World of animals 1 mammals small carnivores

Page 1

Raccoons, Weasels, Otters, Skunks


Boston Public Library

REFERENCE Boston,

MA

02116

i


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WORLD

OF

ANIMALS

Raccoons, Weasels, Otters, Skunks...

PAT MORRIS, AMY-JANE BEER

GROLIER


Published 2003 by Grolier, Danbury, CT 06816

A

1

division of Scholastic Library Publishing

This edition published exclusively for the school

and

library

market

Planned and produced by

Andromeda Oxford members of

Various

Limited

11-13 The Vineyard, Abingdon, Oxon 0X14 3PX

the

weasel family: European

badger

(1);

pine marten

(2);

www.andromeda.co.uk

European polecat (3);

wolverine

(4).

©

Copyright

Andromeda Oxford

No part may be reproduced, stored

All rights reserved.

Limited

2003

of this publication in

a retrieval system, t

or transmitted in any form or by any electronic, mechanical,

means

photocopying, recording,

or otherwise, without the permission of the

Congress Catalogmg-in-Publication Data

Library of

copyright holder.

Morris, Pat. / [Pat

Morris,

Amy-Jane

Beer, Erica Bower],

Contents

v

1

.

Art Editor and Designer:

Small carnivores

-

v.

2.

-- v. 3.

Large carnivores

mammals

Sea

-- v.

4.

-- v. 6. Ruminant (horned) herbivores -- v. 7. Rodents -- v. 9. Insectivores and bats -- v. 10. Marsupials. and lagomorphs v 8 Rodents 2 ISBN 0-7172-5742-8 (set alk. paper) - ISBN 0-7172-5743-6 (v.1 alk. paper) - ISBN 0-7172-5744-4 (v.2 alk. paper) - ISBN 0-7172-5745-2 (v.3 alk. paper) - ISBN 0-7172-5746-0 (v.4 alk. paper) - ISBN 0-7172-5747-9 (v.5 alk. paper) -- ISBN 0-7172-5748-7 (v.6 alk. paper) -- ISBN 0-7172-5749-5 (v.7 alk. paper) -- ISBN 0-7172-5750-9 (v.8 alk. paper) - ISBN 0-7172-5751-7 (v.9 alk. paper) - ISBN

Primates

-- v,

Large herbivores

5

Angela Davies, Penny Mathias

Editors:

cm. - (World of animals)

p.

Graham Bateman

Project Director:

Mammals

Steve

Marian

Editorial Assistants:

Dreier, Rita

-

1

Demetriou

:

.

McCurdy

Tim Williams

Cartographic Editor:

Picture

Manager:

Claire Turner

:

Picture Researcher:

Vickie Walters

:

0-7172-5752-5 1

:

alk

Mammals-Juvemle

Clive Sparling

Researchers:

Dr. Erica

Bower, Rachael Brooks,

Rachael Murton, Eleanor

paper)

literature

[1.

Mammals

]

I.

Beer,

Amy-Jane.

II.

Bower,

Thomas

Erica.

Origination: Unifoto International, South Africa

World of animals (Danbury, Conn.)

IV

Title

Ill

(v .10

Production:

<

OL706 2 M675 2003 599-dc2

Printed in China

RfF

Q

About This Volume

.

1'

2002073860

Set ISBN 0-7172-5742-8

lU'v

MlclS

200 3 volume

n this

we

introduce you to the basic features of

all

mammals.

We

then go on to cover several carnivore

I groups, from weasels to otters to skunks and mongooses, which are mostly distinguished by their small carnivores tend to be

average domestic

cat, yet

Despite their small

and rather

alert, active,

no

less

dangerous to

size, several

o’

.

in

' i

•on'

]

and

,

al

tigers (see

Volume

vorc', inhabit life

m

2),

They are natural-born

like insects,

kill

numbers

live

of

them

smaller than the

which are too small

to bother with.

for the larger

and more dramatic carnivores, such

However, several species have a varied

diet,

ranging from

every continent (except Antarctica) and most major habitats, although only one

the sea

whi e others

many

animals larger than themselves. But being small also offers

Many

are adapted to tree climbing,

derground, emerging to hunt rodents and rabbits at night. group',,

killers,

their prey.

species regularly

opportunities to exploit prey animals r,

fierce.

alone

Some

size. All

where they pursue prey

Some

small carnivores

or lurk

live in

in

fish to fruit.

— the sea otter

ambush. Others

highly organized social

species are quite numerous, but others have been reduced to perilously low

as a result of hunting for their lustrous

and valuable

furs.

live


Contents How to

Use This Set

4

American Badger

Find the Animal

6

WHAT IS A MAMMAL?

8

SMALL CARNIVORES

European Badger panda

18

lives in

temperate

Striped

THE RACCOON FAMILY

20

Common

22

Raccoon

The red or lesser

Honey Badger

Ringtailed Coati

28

Red Randa

30

THE WEASEL FAMILY

32

Least Weasel

36

Skunk

THE CIVET AND GENET FAMILY

88

Common Genet

92

Common

94

Palm Civet

Fossa

Stoat Polecat

Black-Footed Ferret

THE MONGOOSE FAMILY

98

40

44

Meerkat

100

Dwarf Mongoose

106

46

Indian Gray

American Marten

48

Fisher

50

American Mink

52

Wolverine

56

European Otter

58

Mongoose

108

Banded Mongoose List

110

of Species

112

Glossaiy

114

Further Reading and Websites Set Index

North American River Otter

64

Giant River Otter

68

Short-Clawed Otter

70

Sea Otter

72

117

118

Various

members of

the raccoon

family: a ringtail eating a lizard

(1);

kinkajou

(3).

coati

(2);

forests.


How |

M

Use This Set

to

World of Animals: Mammals

If If describes

is

a

10-volume

mammals from

in detail

all

set that

Article Styles Articles are of three kinds.

corners of

the earth. Each volume brings together those animals that

introductory or review

are most closely related and have similar lifestyles. So

all

groups

and

introduces smaller groups

the meat-eating groups (carnivores) are 2,

and

are

all

in

Volumes

3,

and so on. To help you

find

The

volumes that

animals to be found

in

pages 6 to 7 (Find the Animal).

brief introduction to

each volume

also given

A

article

on page

2

(About This Volume).

filled

statistics

of each animal

introduces large animal

makes up most

like families

articles

(The Raccoon

review the

different groups.

of each volume.

It

The

full

variety of

third type of

concentrates on

describing individual animals typical of the group detail,

Data panel presents basic

One

orders (such as whales and dolphins). Another

Family, for example).

interest you, look at

is

article:

mammals)

the seals, whales, and dolphins (sea

m Volume

1

like

There are two types of

such as the

tiger.

Each

article starts

in

great

with a fact-

data panel to help you gather information

at-a-

glance. Used together, the three article styles enable you to

Image of animal typical

in

become

familiar with specific animals

their evolutionary history

and

in

the context of

biological relationships.

pose Article describes a particular animal

Name and scientific classification of

animal Scienti f: c

name of animal

additional information about each

Sizes given in imperial units followed

Captions to photographs provide

animal's lifestyle

by

Common name metric equivalent

of animal Visual comparison of

average-sized adult

mammal and

6-foot

human

being.

(1.83-m)

Scale in feet (meters)

Habits

$oo 4

vrto*»-iy

sem m

groups of up to 20

eccor&ng to age 4nd vx. often moves

Breeding

M-

produce

'T'rflu'*

.!

years after gestation

Vo*a

< »

•Kjuean.

•.

penod

end

of

1

calf

every 3

14-15 months

used

for

Basic description Diet

Habitat

of animal, cjk)

i’

rt

sees

ge^^'iy

na.»r

sea

xr

its life,

and

in

distribution

m

breeding and lifespan Distribution

i

i

,t«-

be based on figui related species)

Conservation status (see Glossary

Volume

1,

and

pages 9

and

10)

Cross-references

Locator

4

maps showing

to relevant

pages

and other

each animal's

in this

normal range

volumes

Easy-to-read and

comprehensive text


A number

of other features help

you navigate

through the volumes and present you with helpful extra information. At the

references to other

bottom of many pages are crossarticles of interest.

related animais, animals that live

in

They may be to

similar places,

\

animals with similar behavior, predators (or prey), and

much more. Each volume

also contains a

the complete World of Animals:

mentioned scientific

with a

Mammals.

the text are indexed by

animals

common and

A

there are words used

in

will also

help you

you do not

list

fully

if

of useful Further

"List of

the

understand. Each volume ends

Reading and Websites

help you take your research further.

heading

All

names, and many topics are also covered.

Glossary text that

in

Set Index to

Species" you

Finally,

will find

of the animals that are covered

in

that

under the

expanded

listings

each volume.

Detailed

maps

clarify animal's

distribution

Meticulous drawings illustrate a typical selection of

group members

At-a-glance Tables

summarize

boxes cover classification

of groups

Who's

Who

tables

summarize

topics of special

and give

scientific

names

classification of

each major group and

interest

of animals mentioned

in

the text

Introductory article describes major

groups of animals

Graphic full-color

photographs bring Detailed diagrams illustrate text

give scientific

names of animals

mentioned

in

the text


Find the Animal x

yi World of Animals: Mammals

If If

library that describes

Each cluster of volumes familiar

in

group of animals

amphibians,

fish,

and

all

is

the

groups of

— mammals,

but

cover a

will

birds, reptiles

and other

insects

the kingdom Protista that were once regarded as animals,

animals.

living

World of Animals

World of Animals. Also included are those members of

part of a

first

now form

group that includes

part of a

organisms. Kingdom Animalia

and

is

single-cell

all

numerous

divided into

major groups called Phyla, but only one of them

invertebrates.

These groups also represent categories of animals

(Chordata) contains those animals that have a backbone.

recognized by scientists (see The Animal Kingdom below).

Chordates, or vertebrates as they are popularly known, include

The Animal Kingdom The

world

living

is

the animals familiar to us and those most

all

studied by scientists

divided into five kingdoms,

which (kingdom Ammalia)

one

amphibians, and

of

the main subject of the

is

— mammals,

fish. In all,

birds, reptiles,

there are about 38,000

species of vertebrates, while the Phyla that contain

animals without backbones (so-called invertebrates, such mice Volume

squirrels, rats,

chinchillas

7;

cavies, porcupines,

as insects, spiders,

Volume 8

many more.

species, probably Q''je'

.

lagomorpha

rabbits, hares, pikas

Volume

1

-e.

.:

at least

To find which set of

in

the World of Animals

is

relevant to you, see

'Oder Scandentai Volume 9

the chart Main Groups of Animals (page •

million

1

8

volumes /

and so on) include

r

• .

es (Oraer

'sect

.

ora

1

7).

shrews, moles, hedgehogs Volume 9

Colugos, ‘lying lemurs (Order Dermoptera)

Primates Order Primates

Volume

8

Mammals

lemurs, monkeys, apes

Volume

Pangolins (Order Pholidota)

Volume 4

in Particular

World of Animals: Mammals focuses on the

9

most

most

familiar of animals, those

easily

Carnivores (Order Carnivora): raccoons, weasels, otters,

skunks Volume Seals

1;

cats,

and sea

recognized as having fur (although

dogs, bears, hyenas Volume 2

Volume

lions (Order Pinnipedia)

Odd-toed ungulates (Order rhinoceroses, tapirs

may be absent

3

mammals

Perissodactyla): horses,

Volume

in

like

many

this

sea

whales and

5

dolphins),

and that provide

Even-toed ungulates (Order Artiodactyla) pigs, camels

Volume

5;

deer, cattle, sheep, goats

Volume

milk for their young.

6

Mammals

Whales and dolphins (Order Cetacea): Volume 3

are divided into

major groups (carnivores, Bats (Order Chiroptera):

Volume

9

sloths, armadillos

Volume

and

primates, rodents,

Xenarthrans (Order Xenarthra) anteaters, 9

marsupials to

name

just

Elephant shrews (Order Macroscelidea):

Volume

9

The chart shows the major

Volume

Aardvark (Order Tubulidentata

9

groups of mammals

in this set

Hyraxes (Order Hyracoidea) Volume 8

arranged

in

evolutionary

Dugongs, manatees (Order Sirema)

Volume

relationship (see

3

Elephants (Order Proboscidea)

Marsupials

Volume

5

opposums,

volume

in

appears

is

page

10).

The

which each group indicated.

You can

find individual entries

by

kangaroos, koala

Volume 10 Monotremes (Order

looking at the contents page for each

volume or by

Monotremata) platypus, echidnas Volume 10

6

consulting the set index.


SINGLE-

ANIMALS

CELLED

Kingdom Animalia

Kingdom

LIFE

Protista

The Main Groups of Animals alive today.

Vertebrates/

Volumes that cover each major

Invertebrates Numerous Phyla

Chordates Phylum Chordata

group are indicated below.

Insects, spiders.

Mammals Class

Mammalia

Amphibians

Reptiles

Birds Class Aves

Class

Class Reptilia

Amphibia

Single-Celled

mollusks, spinyskinned animals,

Fish Several classes

Life

worms

major groups are shown on the chart on

a few). All the

page

6.

few

To help you find particular animals, a

familiar

in

Naming Mammals

this possible.

names

To be able to discuss animals,

Most people regard

animal and lions as another.

are

needed

tigers as

All tigers

one kind of

more

look

for the

They breed together and produce young

or less

it

at different

was one

species.

formal system of classification makes

Domestic cats are

similar to lions

and

tigers,

example, they do not

genus

(Felis),

but

roar).

Felis,

They are put

a different

Panthera, and other catlike animals

grouped together as the family

mammals

in

Felidae.

The

flesh-

eating

animals corresponds closely to the zoologists' distinction

together with a few plant-eaters that are obviously

species. All tigers

lions to another.

The

lion

belong to one species and

species has different

different languages (for example, in

Swahili),

common mountain

and often a

Lowe

single species

names

all

in

German, Simba

in

may have

several

names. For example, the North American lion

is

also

known

as the cougar,

related to

them

(cats,

order Carnivora. These and suckle their Finally,

the

it

all

mammals

in

amphibians,

all

other animals

reptiles,

and some other animals that seem to be in

the

the class Mammalia.

are included, with (fish,

in

the other animals that

young are grouped

panther, and catamount.

Zoologists find

dogs, hyenas, weasels, and so on),

(such as pandas), are grouped

that have backbones

puma,

larger

the catlike animals or

all

themselves. This popular distinction between kinds of

between

(part)

but not as similar as those species are to each other (for

are

like

A

the mammals.

named

make statements about

often necessary to

groups of animals: for example, all

alike.

is

It

the chart.

different kinds.

described and

times without the zoologists realizing

ones, such as sheep, goats, cats, and dogs, have been

included

may have been

species

Volume 21

Volumes 21-30

Volumes 31-40

Volumes 41-43

Volumes 44-50

Volumes 11-20

Volumes 1-10

and

birds)

related to them,

the Phylum Chordata.

convenient to have internationally

recognized names for species and use a standardized

system of two-word Latinized names. The

lion

is

Panthera leo and the tiger Panthera

The

first

Panthera,

is

the

similar species),

name

of the

genus

which includes the

second word, leo or

tigris ,

within the genus. Scientific

the world. The scientific

tigris.

(a

group of

lion

called

word,

closely

and the

tiger.

The

indicates the particular species

names

name

is

are recognized

all

over

used whatever the

language, even where the alphabet

is

different, as in

Rank

Scientific

Phylum

Chordata

Class

Mammalia

Order

Carnivora

Family

Felidae

Genus

Panthera

Species

leo

name

Common name Animals with a backbone All

mammals

Flesh-eaters/carnivores .

Big cats

Chinese or Russian. The convention allows for precision

and helps avoid most confusion. However,

common one

for

scientific

it

is

Lion

also

one species to apparently have more than name. That can be because

All cats

a particular

The kingdom Animalia families, genera,

is

subdivided into phylum,

and species. Above

is

classes, orders,

the classification of the

lion.

7


WHAT IS A MAMMAL? 1

M eating

ammals from

otters

that success

tigers, flying bats

all

become

example,

first

nourish the

them apart from the

much more

new group

of animals;

ancient origin. Reptiles,

appeared millions of years

earlier,

and

date back more than 400 million years.

fish

They include the production of active

enabling

and the secretion of milk to

newborn young.

mammals

Hair

reptiles, since

they depend on sunshine to

keep warm. The

and

first

mammals were

insignificant creatures.

today's solenodons

The Origin of Mammals

began later

million years

mammals

animal world.

evidence of limbs

ago

a

group of small

to develop the special features that

enable

Fossil

some

to

reptiles

would

dominate the

remains provide clear

of the changes. The

became more

upright, supporting

the body from below rather than

sprawling out to the side as

and

crocodiles.

The

skull

powerful jaws, and

needed

in lizards

developed more

spare� bones that were no longer

as part of the lower

jaw instead became bones

the ear, greatly enhancing hearing teeth of reptiles gradually

mammals

to exploit a

fur also developed,

to colonize cold places

Looking

ability.

became more

much wider range

in

The simple

diverse, enabling

of foods than

small

like

and gymnures, they

fed on insects caught at night.

About 200

and

nocturnal. Such feats are difficult for

are a comparatively

others have a

progressively.

offspring instead of eggs

the animal kingdom.

Mammals

fossils

are other important changes that must have taken place

the combination of special that sets

can cope with. Brain capacity enlarged too,

complex and sociable behavior. Less apparent from

animal groups. The secret of

mammals

reptiles

probably associated with increased intelligence and more

and

they have diversified to

In fact,

lies in

characteristics in

tor

ago. They

include humans, seals, shrews, mice, grass-

the most successful of

many

furry creatures that evolved

reptiles millions of years

cows and carnivorous

swimming

rest of

warm,

are

and become


WHAT

MAMMAL?

A

IS

Body Temperature Mammals Reptiles

are often described as

and even some

fish

"warm-blooded."

are also

warm

inside,

but

they cannot maintain a high blood temperature without

basking

in

the sun or constantly moving around to

generate heat from their muscles. Mammals, on the other hand, are "homeotherrmic," meaning that their inner

temperature

is

usually held at

not only high but kept constant.

sun.

so

mammals

Homeothermy

high altitudes and

Adaptability, opportunism,

and

life.

The warmth

generated

is

are independent of heat from the

also provides protection

temperatures, making

©

is

about 95 to 100°F (35 to 38°C)

throughout the animal's internally,

It

in

it

possible for

polar regions.

from low

mammals

to

Mammals can

live at

also

be

intelligence, plus the

capacity for intricate social relationships, are traits that typify

mammalian

success. All are

found

in

the red fox.

©

Fossils

from the early Eocene (49 million years ago) were found

at Lake Messel, Germany. The reconstruction

community

at the

the

dawn of

first

known

the age of

bat

(1);

on the

left

shows a

mammals: Archaeonycteris,

Messelobunodon, an ancestral artiodactyl

(2);

Propalaeotherium, an ancestral horse

(3);

Lepticidium, an insectivore

(4);

Paroodectes, a miacid (5);

Eurotamandua, an anteater

(6);

Pholidocercus,

a

hedgehog

(7).

9


SMALL CARNIVORES

ana that probably enabled them to escape

a:: .e a* nigh

competition with the dominant reptiles of the day.

Body heat *

is

easily lost, especially

from small animals

nat have a relatively large surface area

their mass.

in

So there are benefits to being

evolutionary lines produced very large

culminating

in

comparison, to big; several

mammals,

the blue whale, the largest animal that has

ever lived Elephants and various extinct giants also benefit from size, not least because

few predators can

.ouch them once they are full-grown. Large

mammals

are

also efficient in energy terms, needing less food per

pound

body weight than small ones. But they require

of

a

greater total quantity of food and will soon eat everything

within reach unless they

move

on.

A

body can be

big

a

handicap because moving around becomes progressively

more J

r

jj

difficult

c

their

with larger

size. Big

animals also put greater

bones and bodies, especially when moving

Rodents (Order Rodentia)

Lagomorphs (Order Lagomorpha) Tree shrews (Order Scandentia)

Shrews, moles, and hedgehogs (Order Insedivora)

Colugos (Order Dermoptera) Primates (Order Primates)

Pangolins (Order Pholidota)

Carnivores (Order Carnivora) Seals

and sea

lions (Order Pinnipedia)

Odd-toed ungulates (Order Perissodactyla) Even-toed ungulates (Order Artiodactyla)

Whales and dolphins (Order Cetacea Bats (Order Chiroptera)

Xenarthrans (Order Xenarthra)

Elephant shrews (Order Macroscelidea)

Aardvark (Order Tubulidentata)

Mammals must expend energy -

Hyraxes (Order Hyracoidea)

Dugongs and manatees

warm

or cool themselves. Elephants lose

body heat by flapping

(Order S.rema)

—

Elephants (Order Proboscidea)

fl

their ears so that

cool air flows over the blood vessels,

and by splashing

METATHERIA INf RACLASS

to

in water.

Marsupials (Supercohort) •

The chart shows the evolution of

SUBCLASS PROTOTHER'A Monotremes

mammals between

the Jurassic era (205

(Order Monotremata)

million years ago)

144

10

million years

ago

65

55

34

24

5

1.8

and the

Pleistocene

era (nearly 6 million years ago).


The

T

Diversity of

Mammals mammals

here are at least 4,500 species of

about 8,000 species of

More than

half of a

sma

s.zec or

and over 20,000

birds

known mammal

and so

sma' est

mamma

on), while (Kitti's

one

in

five

is

They also

arger and both are

The

m

a r gest

ion

single

t

mamma

°~es as

much. The naked

may

travel

a r -us! Migrations

exceot

when

the/

me

Hundred

flesh, fruit, blood, fish,

bamboo,

e rat stays

insects, nectar,

a

in

16,000 km on

Some mammals ^eet

ts

grass,

over 10,000 miles

ive solitary

to mate; others

zed socia systems

knows

ndividua

open oceans. They eat

but whales and

Is

numbering thousands. Yet others tightly orga:~

a

p ace.

have highly complex behavior.

wh

only

MAMMALIA

—2

subclasses,

Order

Order: Carnivora carnivores: 240 species in 92 genera and 8 families Order: Pinnipedia seals and sea lions: 33 species in 21 genera and 3 families Order Cetacea whales and dolphins: 85 species in 41 genera and 14 families Order Sirenia- dugongs and manatees: 4 species in 2 genera and 2 families Order Pnmates strepsirhines, tarsiers, monkeys, and apes: 260 species in 64 genera and 1

Order

— shrews: T8 species genera and family Order Dermoptera—colugos: 2 species

Order

Scandentia

tree

in

6

1

—elephants; 3 species

Order Proboscidea genera and

1

Hyracoidea T

hyraxes:

sts the

1

~a

a

species

aardvark:

Artiodactyla

s.

3 genera

species

1

species

ungulates:

— Order Chiroptera— genera and 8 Order Xenarthra— 13 genera and 4 Order — Insectivora

1

family

1

423 species

insectivores:

in

67

bats: over

1

900

species

in

174

families

xenarthrans: 29 or 30 species

in

Pholidota

and

T

families

pangolins: 7 species

in

1

genus

family

INFRACLASS: Metatheria (Marsupials) Order Didelphimorphia Amencan opossums: 63

— Order — genera and Order —monito genus and Order Dasyuromorphia—

5 genera and 1 family Paucituberculata shrew opossums: 5

species

in 1

in

family

1

1

1

del

monte:

1

family

marsupials:

64 species

1

in

1

8 genera and 3

—marsupial moles: 2 —

1

96

Order

in

1

Diprotodontia

—possums,

wallabies,

kangaroos, wombats, and koala: 125 species in

431 genera and 28 families lagomorphs: 81 species 12 genera and 2 families

in

—elephant shrews:

4 genera and

genus and 1 family Order Peramelemorphia bandicoots: 21 species in 8 genera and 2 families

species

Order Lagomorpha

families

genera and 6 families

species

ungulates:

rodents:

are

Order Notoryctemorphia

in T

families

in

in

families

families

in

Mammals

Australasian carnivorous

in

family

Penssodactyla

world's ra;or grouos of

species

Macroscelidea

species

family

Tubulidentata

This chart

2

family

— 7 and Order — genus and Order —odd-toed 6 genera and 3 Order —even-toed 82 genera and 10 Order Rodentia— over 1,990 Order

in

139

in 3 Microbiotheria

family

T

communally.

live

species

in T

genus and

mammal.

territory

orders,

species

families

-

27

SUBCLASS; Theria (Livebearers) INFRACLASS: Eutheria (Placentals)

— —

*

the world's smallest

is

of creatures.

— — —

*

Some defend

hog-nosed bat

the world's most diverse and adaptable group

truly

CLASS:

Many

social

systems, while others

ch each

Some mammals have

Kitti’s

with sophisticated scent-marking and recognition

es

ve permanently n

~

and gum. They

shout, sing, and stay silent

ives

form color

sgr

and swim. They

equator and from deserts to the

is

fe,

burrow for :ne whole of

Tir seals

weighs

fly,

land.

beetles.

sma ertnan many-

(the b ue whale

dig,

on

run

range from the poles to the

hog-nosed bat weighs only a

twentieth of an ounce 1.5 g) The Etruscan shrew scarce)/

Mammals

faster than anything else

bat The

a

many mice breed

almost continuously.

mice,

rats,

at a time, often at intervals

of over a year, but

fish).

species are rat-

e r Qne-third are rodents

squirrels,

one young

(but

38 genera and

1

in

0 familes

SUBCLASS: Prototheria Order: Monotremata

(Egg-layers)

—monotremes:

3 species

in

3 genera and 2 families

11


rapidly c

tney stumble on uneven ground.

if

Consequently, there has to be a compromise, and land

mammals cannot be much

w tnout

exceeding the

support

larger than

limits of

what

an elephant can

their bodies

.

the water things are different. Buoyancy

in

removes most of the ejects of

gravity,

enabling the great

whales to be much larger than land creatures. But they turn

managing

a

body pumping blood through enormous lengths

gigantic of

imited by the problems of

become

in

vems and

oxygen to tons of

arteries, distributing

muscles, and conducting nerve impulses along nerves that

may be over 60

m)

feet (18

long.

Body heat can be conserved by using

mammals

done by having dense

is

it

sometimes with Fur

protection

found

m

layers of fat

layers of fine fur,

under the skin for extra

mammals and cannot be

a feature of

is

insulation. In

any other group of animals. Long hairs protea

from ram and snow, while woolly underfur keeps the

warm Dense

animal

and to stand out

arctic b zzards

mammals

also enables

in

to survive

the open

all its life.

Fur

yet be quite

exploit the possibilities of living

nooks and crannies away from predators and sheltered

•rom the elements Small a

in

warm and

to keep

Tmy mammals can

sma

muskox

fur allows the

owing

treetops

and monkeys to leap among the

squirrels In

size also permits greater aaivity,

addition,

it

ensures the efficient use of tiny

things for food, such as ants

and small seeds

Body warmth has many advantages over coldbloodedness Nerves work activity,

increasingly

reactions quickly,

Warmth

making

Digestion

is

also allows muscles to

more

rapid,

efficiency

and constant

body warmth

comfortable, aspects of

warm

it

is

at

faa

body,

mammals

and even to

is

In

many

of the major

conjunction with their

also have a highly efficient blood

higher pressures than

them

in all

works

faster

other animals except

enough food and oxygen

to function

fully.

Reproduction

Mammal

offspring are born as aaive babies

and so avoid

the perilous process of laying eggs and incubating

fly.

not just about feeling

Their blood circulation

the tissues to allow

contraa more

temperatures are kept high

essential to

birds, constantly delivering

12

if

mammalian success

circulatory system

and

in

greater

and digestive enzymes can

work with greater So,

much

complex behavior, and more rapid

possible to run fast

:t

also

enabling

faster,

to

in

the

same

place for

weeks on end. Newborn young

fed on milk secreted by the mother's

struaure^ that are not present

in

mammary

manage

glands,

to raise a

litter

in seals

fatty,

but milk produced within the

hours after birth also has other benefits

immune

proteins.

and

of babies that

weigh more than the mother that feeds them. Milk nourishing and

are

any other group of

animals Milk allows rapid growth, especially whales. Shrews

them

in

is

first

the form of

They provide valuable protection for the

young animals against

infection by germs.


This idyllic scene of

Hanuman

langurs grooming

hides a sinister side of their society.

they

may

kill

and young playing

When new males

take over a group,

Infanticide

^he

the infants of their predecessors.

killing

of

young by members of

their

own

species (infanticide)

is

one of the most

examples of aggression

in

the animal kingdom.

look after them. Parental care, sometimes involving the

been recorded

00 mammal

male too (although he never produces

possible benefits are varied

I

Feeding the babies on milk requires the mother to

among mammals and another young to

what

is

learn

big benefit.

their parents, enabling

It

is

normal

allows the

them

to

know

dangerous, where to go, and what to eat without

them having results).

from

milk),

to find out the hard

Parental care

especially in primates,

sometimes

way

(perhaps with fata'

lasts for

many

and contributes to the

In

in

over

1

arresting

species.

and depend on

The

species.

chimps, which eat their victims, such behavior

simply be to obtain food. rival's

In lions a

offspring brings the bereaved

male that

mother

breeding condition so he can mate with

has

It

may

kills

a

into

her.

years,

fact that a

13


ao' wn**

5* e-e'r-* of r»e

;

-

as

'

acT

**f

r-. r

.

^ ~f~

-o -ra

-

s oas-: aony ec. ooe^f s -oc *>r to

-¥c. 'ements or -o'? saeca

:. a'

ne vse

ma» oe ea*-e2

mamma

5

rather

*.

and c r as nner : most

eam

ge-es anc

'

";

r

e a: .e *

s

me.

s:a'r:*.pec arp to a dtoere-T

0*

a>

k.,

Va—ma r

.o-"2 a e r

.a

fam

success^

Diversity

nxxe

0*

Many

ole

a-2 not necessan

soec.es also

.

2^3

..

than they

coope-ate to

s

Cunous

.

—e

m ght

1

m

»e

?s

^*oe re

s*

cod

ng

fewer

gh

to ac- e.e a h

groups

soc;a

babies

'a se

,

a result, the

in

more

young

edgehogs.

r

g-ve-n

jp ore or

exa-p e wha es

tn

the case o* whales

m

very .\a'm

o°a rn

or.as

•V'e

'

and the

n that they p'odjce eggs

to babes, but they' st

At r mi.

‘ood

s

1

nourtsh

seasonally scarce,

and other htoernators abandon

bats,

enoug' ene'gy to so K be^nat ng s

burrows Ech

too

t

a'*

a h»gn body temperature, and

inciuc

ng four

feet,

—-as bee- rnodifeo to a*tow the

.e toes

of e gr

ar- d

*'

ers

,

eff

52 r:.'-'

n

of carr ioies

In

s-<c~ 25 *. -.sis

zr ~e~.es r

typ-ca

ro'-e

rena n

the rones

a

-c

*cvi

cent c ggens. runners,

The teeth are also speoa tzed to

wto mar)

.

2 ifferent k nos of *002 *'o~

aa^es fsh, orass. blood, and nseas

'"eat to nuts

Agam„ there are exceptions associated with speda *estyes Ant-eating speoes such as the aardvark or gnaint a'-teater.

have 'eo^ceo teeth or none at a

try "ood ^eeds no d*ev. ng

s nce the;'

1

ii

Instead, a tong, sticky

tongue s 'eo- 'eo to gather the nsects

The spec .a character-stcs of to

.era

mamma

w oe vanety o* habitats more

the f surroundings a

r

enable them

temperature, for example* than any

Mammals

other animals

is

independently of

:

a re found everywhere, from the

poar 'eg >ons to oese-ts mountamtops. and

Some mace protongeo O

miles

m^y

fly

many

on annual

rr

jungles.

ves nto the oeec sea. while

es each night or walk humQ'eds of

migrations.

Mammals can

surv.rve

successfully n a wider diversity of hapnats than

any other

One mammal Homo sa&enst has even

arim.a group

advantageous behavior

The base mammal oody p an

deveopment

fused to fry— the

others

mes the animals do not nave

ma ntam

:ones

b ucoe-'

n Mole 'ats protect themselves from the

nomeothe"^ * At such

f

among

e nearly hairless as an adaptation *'e

.mar and centra

cooe e~ oent

groups. but no other a" mails have

,%a. c*

scaptoc

t'r^re' '-ex b r.

oesc'oeo above a e found

a*-

mhkh the

Wrists in

rxfqoe'xfent

or their own.

r

'ats a

nstead of gn ng c

sw mmers

'ror—a

provided by aroe amounts of fat

msuiat cm

th

r

appropriate

.

one As

.

spec a advantages For

r

par eular

.

sma

some mammals -eve

.

platypus a'e except

14

;s

and Exceptions

anc -a«:eo mo*e

v.

patterns into

-3-2 te-as to oe

es a r e oe-e'a

nonmamma

ail

each

cobras*. 'ept es

irfe.

c* ** e *eat-'es

*nem

the»r

"2 re

c a and ca- oe ac jsiea to

a-

kxxed are' and tend

.-.e

ate

.vtnch .nd

by

'52*5

r.e ~-e 32 .a-*.age

.

*ran *0 .oung, and somet Ties or

to tne»

in

oe**a. <c'a

'

tec benavxx, on :ne ot-er

cera»n

s./C“

so rs o* core tons a-d circumstances

r-ee: a

Some

Denav 0 r c

5 *.2*5

23 '* mnerted

o* :ne

e 5 "tc oe-a. o'

ir'-r'

arc soeoes.

a^o oat

2" C '2'OC'* Dr C*

*-3

hts s

anoec on toe moon, something that no other an imal has eve' dchevea'

Some mammals have become abundant

and

w oesp'eac A few

that

we

treat

them

are so successfu

and numerous

as pests because they

compete with


WHAT

humans

for food or destroy

in

MAMMAL?

what we have produced and sheep have

ourselves. Horses, dogs, cattle,

played a major role

A

IS

helping

humans

our

in

for

also

own

social

evolution and successful conquest of the planet.

Domesticated mammals,

in

exchange for regular food, with useful services, such

shelter, or healthcare, provide us

as transportation

and muscle power, or products,

like

meat, milk, and leather. Such items would cost us a

more

effort to obtain

(like rats)

Some mammals

by other means.

endanger

are significant carriers of diseases that

many

people, but

assist

medical research to combat disease

some primates and

lot

laboratory mice

among humans.

Challenging Present, Uncertain Future Today

we

live in

the age of mammals. This group

dominates the world,

now

effectively replacing the reptiles

and

amphibians that were once the major land animals. Those that remain are reduced to relatively

comparatively restricted range of the

last

place

5

A

lion killing a buffalo.

Even when agape, the jaw can deliver

in

1

to 2 million years

the variety of

of the world.

mammal

used to cut the

decline

Carnivores, as in this wolf, typically have 44 teeth, consisting of three

incisors

last

{ 1),

one canine

(2),

upper premolar and

to shear

through

flesh.

four premolars

first

(3),

and three molars

lower molar have sharp

Jaw power

(right)

is

tips

(4).

crucial for the capture

The masseter muscle force

when

needed

to cut

through bone.

(b)

car

vore

v0

--

and

power

different parts

extinct,

critically

and many

small numbers.

The

—our own. Today we humans

it,

all

species that share the world with us.

Mammals have

suffered heavily from persecution by

people, particularly species considered to be dangerous like

the tiger or wolf. Large

they need a

lot

mammals

are at risk because

of space and eat a great deal of food

every day. The

provides the

and grind

in

manipulate the environment on a global scale and, with

and high cusps

tearing up of prey. The massive temporalis muscle (a) delivers the to suffocate or crunch

The

However, over

mainly due to the overwhelming success of just

one mammal species

S

lifestyles.

species

Many have become

others have been reduced to is

species living a

enormous changes have taken

suffocating or bone-splitting pressure. The meat-shearing teeth are then flesh.

few

same

land

cannot support

flesh

the jaws are almost closed.

elephants, for example,

and

also

produce crops

meat-shearing edges

for people. Expansion of

farmland has reduced the

numbers and forced them back built to

of wild animals everywhere

into less suitable habitats. Fences,

keep mammals out of crops, force them to

live in

areas that cannot support such high numbers. Wild

mammals

that steal from crops planted

on the land they

4

15


SMALL CARNIVORES

used to occupy are shot or snared. As numbers of

humans nave

ncreased, wild

same

aside At the

numbers, replacing

mammals have been

of domestic

and goats, have

especially cattle, sheep, in

mammals have been pushed

numbers

time,

world to conserve the

Some

hunted

IUCN has over 1,000

Switzerland, the

Perhaps the best-known

for their skins, ivory,

among

and ecologically sustainable. Based

fair

is

staff

in

and the help

of 10,000 volunteer experts from 181 countries.

wild

production of the Red

or other valuable products. Catastrophic population

declines have resulted, particularly

and natural

seeks to ensure that the use of natural

It

resources

greatly increased

their wild relatives.

specially

mammals,

systems.

diversity of nature

published

the larger

Lists

role of the

IUCN

the

is

of Threatened Species.

First

1966, the books were designed to be easily

in

species because they take a long time to reach breeding

updated, with details of each species appearing on a

age or often produce few young. They cannot

different

compensate

now

nave

numbers

easily for the large

killed,

and many

reached the verge of extinction. About one-

quarter of

threatened

known mammals

all

some way. That

in

information

came

to light. So

far,

the Red

as

Lists

include information on over 18,000 types of animal, of

which over

are significantly

is

new

page that could be removed and replaced

1

1,000 are threatened with extinction.

Gathering such a vast amount of information

roughly twice the

proportion of threatened birds.

but

task,

The Red

it

is

huge

a

provides an invaluable conservation resource.

Lists

are

now

available

in

CD-ROM

format and

Categories of Threat to Animals

on the World Wide Web. Governments throughout the

Two

world use them

types of classification are used throughout this set

regarding conservation status: IUCN categories and CITES

Appendices. They appear, where relevant,

and

the data

in

panel for each species.

when

assessing conservation priorities

in

policymaking.

In

the data panel for each species

section

on

status.

Some

in this set

The IUCN

cases an

With so much conservation is

activity in different countries

some way

important to have a worldwide overview,

coordinating what goes on planet That

is

in

it

of

different parts of the

known

nfluence, encourage,

and

to

throughout the

assist societies

It

in

not relevant.

such

risk. In

Where an animal

IUCN

risk, its

the latest Red

rating

is

given. All

Lists.

to the status of rare species, they

confer any legal protection. That laws,

and

is

do not

done through

national

internationally by CITES.

IUCN CATEGORIES EX

Extinct,

when

there

is

no reasonable doubt

that the last

DD

individual of a species has died

EW CR

Extinct in the Wild,

when

known

only to survive in captivity or as a naturalized population well outside the past range. Critically

a species

is

Endangered, when a species

extremely high

immediate

risk

is

facing an

of extinction in the wild in the

future.

EN Endangered, when

a species faces a very high

risk

of

extinction in the wild in the near future.

VU

the wild

LR

16

when a species faces a high the medium-term future.

Vulnerable,

Lower

m

when

risk

of extinction in

a species has been evaluated and does not satisfy the cntena for CR, EN, or VU Risk,

is

should be pointed out that while IUCN categories

draw attention

is

is

to be officially at

provided

Conservation of Nature (IUCN), also referred to as the

listing

a

categories of threat have been taken from information

the role of the International Union for the

World Conservation Union The mission of the IUCN

IUCN

is

many

animals, for example,

rodents and bats, are abundant and not at

there

NE

Data Deficient, when there

is

about a species to assess the

risk

Not Evaluated, IUCN criteria.

not enough information of extinction.

species that have not been assessed by the

Note: The Lower Risk (LR) category is further divided into three subcategories Conservation Dependent (cd): species that are the focus of continuing species-specific or habitat-specific conservation programs the cessation of which would result in the species qualifying for one of the threatened categories within a period of five years; Near Threatened (nt): species that do not qualify for Conservation

qualifying for VU; qualify for the

Dependent, but which are close to

and Least Concern

two previous

(Ic);

categories

species that

do not


l«.

arts

CITIES

Carver tier on rrematicna

CITIES s Bine

as :re

Ws&r ngtor Convention

meeting

-atons nave agreed Tradi ig

"

C C

\Aasring:cn

i

an,

•,

:c

~C'e ’"e r r

:

<

"own -

a~e' a

I lists-

Currenth TEE

the C

~ais and their ccc\. cats

been a '•"ace

-"as

species,,

nameiy, those that are heading toward extinction wtfl

be harmed by continued

trade. Such)

permit. Permits

are required to cover the whole transaction exporter

and

speaes are

ther native countries and can only

be imparted or exported wtfi a speoai

egJatiors.

TIES

APPOdCES

the most threatened traded

usuaily protected in

'9~5‘

i

Appendix

(also

was acred

since t

V A V WAL"

A

IS

'

Tirade

Endangered Sceces of \A d Eauna arc -ers

HAT

—both

and importer must prove that there is a scientific justification for moving the animal

compelling

m the

-actor

le of some of die *vcddte rarest sceces

dec:

arc CITES aims to

C ~ES eate^ed

contirallthe

Appendices

sts .called

ite r nat:ona

cv.

orSbem

from one country to another. This includes

animals between zoos for breeding purposes. Permits

:a v

sceces

:.nat

are

are only issued

foe id

trace

.'n'e ' a1

"ccaing

ve cr dead

ai

s*cnddder An

when it can be proved that the animal

was legally acquired and that the

made An nals

are assigned ta will;

Appendix

transferring

,

spec.

s* is ‘asrrers

remaining population

mot be harmed by the toss

^ens

and so cr

Appendix

indudes species that are not currently

threatened with ©rttoctiony but that could easily

ce ccrnffisoted tv customs at ntematicaa borders

,v

seacorts or airports Appendix

sceces can ce traded

'temaconali', ouitory under strict controls. V,

trade s

on©" ,aiuacte

am mate CITES

to ~

t:ie

*

*'s;<

,

*viitih

exaroe

far

shew-

rerroers

-

a finest

the

no:

r

'

MS

gnest

co

rave ‘c

Sene

sell;

try

the rare species pretending they were a similar

common one. animus,

with:

Permits are required to ©apart such

requirements similar to those for

Appendix speaes.

coo gpore

UCM

li

rare

Appendix

speaes are those that are at

risk

or

categcnes

protected in at least

one Gauntry. Other nations may

C~ES entree cr

be allowed to trade

m the animate m their products*

ercia

-urner seta's of

the rabe C ~E£

oskc

—mav

smpis because they nave not eai

traders

eev, fines or nensenment.

—eve" these

~ar> cats

are

-era eccrc<~'> and; that

Mevec^eess

arid cecole

nes

sceces

:.ti©-

resemble threatened species* since criminate could

d fe

dihreue questions accut the relative ~cortance cf

•a,ses

become so if trade is not carefully controlled. Some common: animals are listed here if they closed

Acce"d ces

a

_e ~aoa

but they will probably need to prove that they come is

re catchy es

from safe populations


SMALL CARNIVORES M

-

:i

:

i

T

.z a

I

zsze mezazz'S z"

Bs.z"

.•

rn Ca"

ma

s

mose n^cec

as

Hc/.e\e r >ze

jr

;

j a

e

nan me

e'

rc >,zes 50"~e z~ z^e most

_

'

Z:-

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a:,

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30

3 'eez _

_ z

eae


Who among the Small Carnivores? Family: Procyonidae — raccoons: 7 genera, 19 species, including

Who's

ringtailed coati

(

Family: Mustelidae

Nasua nasua ); red panda Ailurus fulgens) otters, and badgers: 26 genera, 66 (

—weasels,

species, including least weasel

nimble. The smallest of as

little

weigh

all,

as 5.5 inches (14

just

the least weasel, can measure

cm) from nose to

under an ounce (28

such small size might seem

weasels and their relatives lack

make up

for

in

speed,

stature they

and

agility,

(.

ferocity.

more than

They

unusual

in

own

brasiliensisY,

them

to reach prey

burrows, and crevices that would

common

genet ( Genetta genettaY fossa ( Cryptcprocta ferox)

Paradoxurus hermaphroditusY,

— mongooses:

17 genera, 35 species,

banded mongoose

Mungos mungo)

foil

in

nests,

are semiaquatic,

larger hunters.

waterproof

Small carnivores can be solitary

meerkats and

(like

coatis). Smell

is

the stoat) or social

important

in

communication, and most species use scent to mark

most notably the

information to other

members

small carnivores climb well, entire

life in

trees.

is

vision.

Most

and some spend almost

Most can swim, and

a

and

webbed

whose feet,

a powerful, rudderlike

otter

spends most of

even

come ashore

tail.

the water and

its life in

their

few

female.

Compared with other

spend a long time with

and learning the

special

dense,

The sea

may not

to give birth.

majority of species rearing offspring

of their species. The sense

also important in hunting, as

fur,

otters,

Reproductive strategies vary widely, but

ranges and territories and to convey personal

is

(

is

Lifestyle

of smell

civet

including meerkat ( Suricata suricattaY,

adaptations to the water include

home

lutris)

civets and genets: 20 genera, 35 species,

any group of animals. They can also squeeze

into tiny spaces, enabling

(like

sea otter ( Enhydra

common palm

Family: Herpestidae

readily

bodyweight, which

stoat

nivalis)',

including

(.

tackle prey several times their

Pteroneura

Family: Viverridae

disadvantage, but what

in

Mustela

(M. erminea ); wolverine ( Gulo guloY, giant river otter

and

For a predatory animal

g).

like a

tail tip

(

skills

is

in

the

the job of the

small animals, the

young

their mother, gaining strength

they

will

need

in

adult

Play

life.

is

an important part of growing up, and games often involve parents as well as offspring.

© A group of banded mongooses vantage point on a termite

find a

mound

in

the Masai Mara Nature Reserve,

southern Kenya.

j

:


SMALL CARNIVORES

The Raccoon Family accoons are medium-sized, long-bodied

R

with a long l

m

the family Procyomdae,

Raccoons have

its

appearance and ecology. Only the kmkajou has a uniform

as a

body color The

of

facial

have distinctive coats with various

markings and ringed (banded)

on the soles of

partly or entirely

a

Is

most

cats,

except

in ringtails

have semiretractile claws on

tails.

Raccoon?

Camdae.

Their origin

is

reflected

name Procyomdae, which comes from pro.

in

the family

the Greek words

meaning "before," and kuon, meaning "dog."

Recognizable

fossils

of raccoon ancestors have been

found dating back 20

million years

—to

a time

when

Europe and North America were one continent. As the continents separated, the raccoon family unequally, with the procyonids remaining

World

(the Americas)

weighing

ringtail,

The raccoons are descended from ancestors of the dog family,

and the subfamily

ancestors of the red panda,

in

was in

their feet, like bears.

thumb. Generally, the claws do not

The procyonids range

What

footed and walk

flat

The

red panda, however, has an extra digit, which functions

display a remarkable diversity in their

rest

toes on each foot, with the third

five

toe being the longest. They are

Although there are only 19

tail

different species

members

mammals

split

the

New

Ailurinae,

the Old World (the Eastern

little

retract like those

and red pandas, which

their forepaws.

in size

from the slender

more than

.8

1

pounds

(0.8 kg), to

the stockier raccoon, which can reach 33 pounds (15 kg).

Despite being classified as carnivores, most of the

raccoon family eat surprisingly the bulk of their

diet,

meat.

little

makes up

Fruit

although they often supplement

with a variety of insects and small animals. The kmkajou tops up

its

fruit diet

pa. idas feed mainly

with the occasional insect, while red

on bamboo shoots,

and acorns. Raccoons

lichen,

and worms as well as

snails,

will

eat

fruit, roots,

fish, crayfish,

berries, nuts,

and the olingo preys on small

and

fruit;

and mammals.

birds

The most carnivorous of the procyonids are the I

Hemisphere, excluding Australasia).

ringtail

and cacomistle. They have well-developed,

doglike carnassial teeth to catch and

Family Procyomdae: 3 subfamilies, 7 genera, 19 species

large as rabbits. their external

RACCOONS AND COATIS

The

slice

diet of procyonids

is

up prey as reflected by

appearance. For example, the kinkajou

4 genera, 12 species

has a shorter than average muzzle with a long tongue Procyon

Nasua

7 species, including

common

raccoon

(P.

lotor);

crab-eating

raccoon (P cancrivorus)

for obtaining nectar

2 species, nngtailed coati (N nasua), white-nosed coati

flowers.

(N narica)

Nasuella

1

species,

coati

(A/

olivacea)

flexible

Bassariscus

?.

species, ringtail

KINKAJOU AND OLINGOS Potos

(kmkajou)

RED PANDA Ailurus

1

red

genus.

panda

astutus), cacomistle

7 genera,

sumichrasti) (B

snouts to

.

probe

for insects.

6 species

species, including

alleni

1

(

(

B

(no

B gabbii (no

common

common name)

species

A

fulgens)

-

20

SEE ALSO

mon

from

contrast, the

species (P flavus)

1

Baaaricyon (olmgo) 5 name), B

In

coatis have long,

mountain

it

1:22, Coati, Ringtailed 1:28, Panda, Red 1:30


wl 1

*

r/

J>

,!//f

i ri :

•>'

u

,

Jiii /

If'

kM

TP

y oJm

i

if*

wjBSl

IMifmtwm

:

nP Hi ThSt

f

i.

f'.B

7

©A

*.

*

pair of ringtails. The species

was

w it# Ik?-/

originally reared as a

prospectors' camps in the early American West, hence

name

r

its

mouser

in

alternative

of "miner's cat."

Lifestyle Procyonids can generally no

live

for

more than seven

until their

so

their first spring.

and provide

all

in

They give

birth in

occupy

a diversity of habitats ranging

through most of

forests,

are

while coatis prefer more

found

in

American

is

cliffs

wooded

and dry

regions. Olingos

tropical rain forests, while the

adaptable raccoons thrive

panda

rocky

in

in all

of these habitats.

The red

confined to Asia, favoring remote, high-altitude

forests. All

procyonids are nocturnal, except the coatis,

which are mainly active during the

day.

g)

litters

and are poorly of three or four, but

pandas produce only one or two young

at a time,

and kinkajous usually only one.

North, Central, and South America. The ringtails and

cacomistles are found mainly

dens or nests

the parental care. At birth the young

developed. Most species have red

but

the wild. While males

weigh only about 5 ounces (142

The procyomds (excluding the red panda)

in captivity,

second year to breed, females usually do

wait in

10 to 15 years

Recent studies of the raccoon family reveal that their social structure

as the coati

move around

the

seem

like

ringtail,

the Procyomdae

in

to be

is

in

the wild

complex. Species such

large social groups. Others,

more

solitary.

While some of

—such as the common raccoon — are

thriving, others are classified

by the IUCN as Endangered

The red panda, olingo, and cacomistle are among those in

decline as a result of destruction of forest habitats.

21


CARNIVORES

Sr.'A__

Common Common name

Raccoon

Cor"non raccoon Scientific

Procyon

lotor

Raccoons are one of the most familiar North American animals. Their adaptability has allowed

Family procyon cae

Order Cam vora Lengtt-

reacted.'

tenet"

8-12

1

8-27

in

Procyon lotor

name

43-68 cm);

them

tail

to

succeed

a wide range of habitats while

in

,

ir*

20-30

err

;

he ght at

shoulder about 10-12 n 25-30

25%

about *

1— 3 *

Key features

but sometimes up to 33

lb

bandit* face mask, accentuated by

raccoon's \~eujge\ce, atertness, and

gray cars above and below, black eyes; short,

curiosity

rcureed

settlers

and

bushy

ears;

black

r

with alternate

ta

ngs fusua /

body

5);

brown

hairs long

were

.

females

may

so'itary,

although related

ve close to one another

r

speed of learning

somewhere between and the domestic

Breeding

Four to 6 young born around February to

Ap after gestation period of 63 days. Weaned at 7 weeks, fema’es usua iy sexually r

;

mature by the

Voice

May

/e over

16m

the wild

r f ^st 1

m

7 years

captivity,

up to

Chitters, purrs, hisses, barks, growls, snarls,

Fruit,

and seeds;

crayfish, clams, sna.

Almost anywhere

m

Distribution Southern Canada,

s,

and earthworms; crops

North America, including

can become quite a handful.

Masked Bandits Raccoons are unmistakable animals, with their

stout

little

tail.

across the

They have

bodies, typically weighing

and 18 pounds

weights are

and 8

(5

known

to

kg),

between

although

change with season and

U

S

,

and Central

common

of raccoon family; continues to

range and increase

in

numbers

than

southern ones. The heaviest raccoon recorded

reached 62.4 pounds (28.3

its

mask

distribution; northern animals are larger

Population abundant Most

expand

Young accoons make

ntrigumg pets, a though as they mature, they

1 1

America

member

cat.

monkey

r

also fish,

urban areas

Status

that of the rhesus

eyes and their bushy, banded

such as corn and stored grain

Habitat

thought to be

is

characteristic black “bandit"

bernes. nuts,

in captivity,

spring, males by 2 years.

and squeaks Diet

Native Arre r car

in

Raccoons are often kept

*o!kiore.

and the Nocturnal mainly

a source of fascination to early

a^d are celebrated

and gray Habits

kg).

Enormous numbers of raccoons have been trapped or shot for their to

make

kept

jackets

in captivity

and

farming, the to France,

of Russia

in

Raccoons are also

gams

to be

made from

Germam ana

the 1930s and 1940s wild,

Switzerland, Austria,

As a fur

raccoon was introduced

Netherlands,

escaped into the

Some have

which are used

to supply the fur trade.

common

me

skins,

hats.

result of the financial

parts

Many

and raccoons spread to

and the Czech Republic.

also turned

up

in

Poland, Hungary,

Denmark, and Slovakia The European raccoons are

now sometimes

considered a nuisance.

The raccoon's coat

is

made up

of hair The short, fine underfur

SEE ALSO

extremely

popular with people.

-i Biaoc

make them

their appealing looks

Vale

arger than female

5-8 rg

fb

err

R ngtailed 1:28, Panda.

is

of

two types

uniformly

Red 1:30, Old World Monkey Family. The 4:40

XT

:


RACCOONS COMMON RACCOON

gray or brownish and provides the animals with

warmth and some

protection from the wet.

Growing from among stiffer

guard

hairs,

short coat are longer,

its

which are tipped with black

The density of the guard

or white.

hairs alters

the overall appearance of the coat, often giving it

shaggy

a fuzzy or

look.

Raccoons molt

in

the

early spring, with hair loss beginning at the

head and proceeding along the back. fur

New

grows throughout the summer to

provide extra

warmth

for the winter.

Many raccoons have amounts

of yellow

variable their coats,

in

and some albinos have been reported. Apart from size the

sexes are similar

in

appearance,

and juveniles resemble

adults.

Raccoons are excellent climbers, aided by sharp claws

and the

ability to rotate

the

hind foot through 180 degrees (thereby turning

Such

ability

it

backward).

makes them one

of

Š The raccoon's "bandit" eye mask, brown-and-black ringed

tail,

round ears are trademark

and

small,

characteristics of

this highly distinctive species.

Raccoon Currency

T

he raccoon's fur has always been the main reason for hunting

and trapping the animal. During the 17th century bans were

imposed to prevent too many raccoon

skins being exported

from the

United States. At one time the skins were used as currency; and

when

the frontiersmen of Tennessee set up the state of

Franklin, local officials received

each

year.

payments of "coonskins"

Although they are hard wearing, raccoon skins

are not especially valuable nowadays,

and trade

no longer

However, a movie

a threat to population size.

about Davy Crockett, king of the wild a

sudden fashion

frontier,

for coonskin caps like the

in

created

one worn

$

in

the movie, costing the

lives

of

many

them

raccoons!

is


iccoons omen use dens

i

*

-:

.-

r

nr-r and speno The day

'

wvhem mar

almost a

•.

exceo*

©ve

dens mav

r»f

The

Trees.

& about 9 to

wmc^

in

m Tan

Each den

the

its

Alaonpuam worn amkun, wn.ch

o:

only

washmp

ven

rts

oea:

occuped

refers to the

mor

name

scientific

-leamnp To wash

roughly translates as "ne

who

a reterence to the frequent grooming that

accoons. characteristically mouipe

tki

is

The German name waschbar.

in

raccoons nadir of washing

its

tood

taken from the Latin wore /ai*re

~he perception that raccoons wash themselves wrm then hands actually comes Timm observations

when

win®.;

roreoaws are a orominent teature and are name oe common name raccoon is derived from

skiltrui

scratcnes wrrh ms. hands

mom

is

accoon-

reflected

to

they wit

pm-prs protection

rencton anr the .veathe

.

tits:

brushy nests oic buildings

nr mies anc haystacks

.a s

ash nc Sears

«\

above the ground ~nev also use

bumnws

r

itwr

r* v

hollow

in

e— ance hr.* that

.-

? ~

fee*

-

hang:

.a"'

air desceno nee funks near

':t

.

the

'

Tacco°hs carchmp and Teedinp on aauaTic prey. ~ne\ danoie and splash, in the ware in an

tot

1

ongp w/np

over

u.c

a

twrtt\

mothe has sne

will

instinctive

afco

movmc

her

1

to catch,

aooearance of washing instinctive

a nr drfftcutr business

manne

off

(ever

tamny

fish, pivinq

their

food

It

is

every

the same

Pehavior disniaved by captive raccoons

when

there

the notion of the

no

is

"

water', that

washing bears

has encouraged

Water ^o\nnc h»e raccoor % torenaw.' nave e wel--aevetonpr sense o* rover

nrnughout ther range

acmons

anr air caaabK of miiaae mamoviaTior.

are found arm os:

B

a.-

skilttu as t

monkret

a'

everywhere that ware; s avaiiab* They are mosr

mangroves hood niam v-

'-

.."

.

>e*'

^

pi

o

Garmons

»

are less

nev also Tenr tr avoir

true

r

oeoends on

ai

cr The northern

southern a-vn

.

ana da thev

win place-

i

^ tartMT

r

an on

fleser;

to

live

m

areas Thev

me

parts,

iea lonp

fr

of then Tanpe they

nrrt

to huddle tooethe’

Raccoons

m

acmmy

active

21

SCI

rate

The» body temperature stays

ALSO

am

warm whe-

r

tvnically active

fust

is

a neai

morn sunset to in

teedmg

before midnight Raccoons

vOasta marshes may be se*r.

all

they snuggle

a single den

aimouoh there

living

bv

Teedmg durmo

the day whpr ther tood source oi crustaceans is

exposed

opfinnunsts. able to

whatever tood

nedme

long and

ur close Up to 25 raccoons have Peer known

hear

m

do

am sieer Their

tall,

e^^ energy keening

the northern imitec States and

wimp

summer and

buildings

ano moMusks

Thei

resents they have Puih up

1

sunrise

^

surviva

denning togethe ovei wmTe: since they use

m att«ides above t t>HD

fcacroons do not hibernate and

no

oi

Ther body weight Raccoons are ohten found

ivem and springs Raccoons

anpK round

nenod ther

Irrtle

harsh winters raccoons may lase up to naif of

united Stares and like

inactive

the- fa:

ove- the previous

pe ooen

Vherr they have spread out onto

metabolic rate remans

moemsrars. Since they consume

tood during ther

common

ia r

thei

high As a result They use more energy than

ub«ai: woodlands especially where pine

-

common m

hanpuy r suburban areas within

~ P-^'de

3n°C. are

95**F

anr abandoned tarmiands and can

-ec

o.iitt

•.

and mesh anr

marshes ~ney are also

are' .

Tnrests

is

at lovy tide

make a meal from

available

does

take advantage of so

above

The secret ot ther success

Am*

I.

an

.

.

*v ?:3f

Raccoons

tt

ts

then

many kinds

at

ability

to

fond that

is

tr Tact

handime

its

roe raccoor

tooc


RACCOONS COMMON RACCOON

In

most areas plants provide the main food

eaten by raccoons, espec

ally,

will also

born a few weeks

eat

earthworms and nsects and sometimes stored gram. Corn

is

a particular favorite

taken jst before

it is

noe

and

jsua

is

a.^c ready *or

'za r ds.

fy

".ran

Where trey

near turt e nesting beaches, they ;

a^d

steal

the buried eggs. They wi

other vertebrates such as gopne shrews, rabb

forage beside lakes, or

and

ts,

m

s,

aiso eat squi

rr els,

fish,

such

and other aquatic

spring, although

animals. They also take

not breed unt

strong swimmers.

At

birth

raccoons weigh about 2 to 3

ounces ^60 to 75 inches

na

r,

HO

cm,

g-

and measure about 4

in length.

They are covered n

although the mask and

tai

rings are

represented or y by dark-pigmented skin. Afte r

Their iegs

/ely

become

their eyes,

and making ch strong

tter

enough

when

ng noises.

for walking first

molt

they shed the

mfant coat, and the adult fur begins to grow.

Raccoons become sexua

and are

a factor that limits

the spread of raccoons farther north.

occurs at seven weeks,

Breeding

being

and May. The

by the fourth to sixth week. Their

horses.

crayfish, clams, snails,

readily to water

may be

squirming act

marshy areas

where they feed on

to survive the winter

from the corpse. Raccoons occasion a y

mamma s

litters

born, the less chance

about three weeks they open

and even

between

the young have to fatten up for winter. Inability

/e

dig jo

itter is

„sus y a r eady dead, so the r accoons just feed

as deer, cows,

rivers,

r

:

nk; but suc^ an mals are

scavenge the rema ns of arger

C Raccoons often

w

usually

later in April

ater n the year a

consumption. Raccoons also eat small birds and

somet mes snakes and

is

February and March, with most

fleshy fruit,

and seeds. They

berries, nuts,

the breeding season

some

their

y

matu re

particularly

n trie

r

*irst

males do

secona season. Mat ng can

be from Decemoe- through August, occur' ng ater in the season farther south.

Tne pea< of

Weaning takes and the young

place from seven weeks,

start to leave

forage for themse ves. They

the nest and

may

still

be suckled

by their mother for up to four months. By ,jvenies

may we.gh up

fall

to 15 pounds 7 kg

,


City Slickers matings

^phe I

variety of

it

become

Males may

to thrive in a

In fact,

occupy

will

live

alone or

small groups

in

and

a distinct territory ranging in size

from 125 to 12,500 acres (50 to 5,000

has

it

|

I

|

human-dominated

environments.

range, with the smaller males

securing a few matings each.

raccoon's adaptability has

enabled

in his

roam over 1,500

general, raccoons will

very familiar to city

ha). In

a year. Males

may

acres

dwellers. However, these

(600 ha)

masked bandits are notorious

but they disperse during the breeding season,

for raiding

garbage

known

only are they

when

Not

bins.

in

fighting

I

travel together;

and competition between them

increase. Social relations are probably

to carry

]

away whole

established

bins, but the

mmble-fmgered

raiders

The secret of the

to secure the bins, rather than

can

make

common

raccoon's success

that

is

different calls

it

scents. At least 13

have been identified

Sounds are used between

meal of almost any available food,

a

in

raccoons.

individuals in close

proximity to each other. Mothers keep

including the contents of garbage baskets.

through them.

and

postures, vocalizations,

have

also learned to untie ropes used

bite

and signaled through various

in

touch

with their young by purring sounds, while

and snorts express

hissing, short barks,

common

Despite the success of the

but

full

size

is

not reached

year. Families generally share a

and the young raccoons

the second

until

den over

leave their

will

by the spring. Few wild raccoons five years,

but

some

survive

after

20

was

mother

more than

live

up to 16

oldest recorded captive raccoon

raccoon, several related species

winter,

The

years. still

living

Cozumel

Social Organization

by the

listed

as Endangered. The Barbados raccoon

(Procyon gloverellani) extinct

sometime

is little

common Two young raccoons

raccoon {Procyon pygmaeus)

Island

— are

there

years.

—such as the

from southeastern Mexico

IUCN

fear.

to

said to

is

have become However,

after the 1960s.

compromise the

survival of the

raccoon. Predators such as wolves,

bobcats, pumas, great horned owls, and I

by

their nest in a

tree.

hollow

to look for their

food, their

move them ground

to a

start

own

mother

will

den

at

level to prevent

them from

falling.

social organization of

raccoons

However, several females related still

will live in

— usually

males

will also

alligators

solitary.

raccoons actually

inhabit the

One

same

or

The main cause

mate

relatively

for

few are

concern

is

the

can be transmitted to humans, such as

season females mate with between one and

leptospirosis, tularemia,

for

mating

is

competition between males

privileges,

with heavier males

gaining greater access to the females. successful male

is

likely to

(and worst of is

One

be

responsible for over half the

killed.

common

raccoon's susceptibility to certain diseases that

with the resident females. During the breeding

four males. There

few

Common

prey to them.

fall

"coon hunting"), but

more

area and

a small threat, but

raccoons are also hunted for sport (known as

closely

areas that overlap, but they

tend to avoid each other.

may pose

not well

is

known, although adults are generally

As they gain

independence and

The

all)

rabies.

The

the major carrier of rabies

United States and all

and most commonly

in

common in

raccoon

the southeastern

1997 accounted

for half of

reported cases of rabies from wild animals

in

the whole country. Raccoons also often host a

type of parasitic the raccoon

roundworm

itself,

that

it is

species,

common

raccoon

is

in

expanding

in

harmless to in

small children.

so familiar in North America

often the topic of TV cartoons. it is

is

may cause death

but

domestic animals and even

The

that

A

successful

both range and numbers.



SMALL CARNIVORES

Common name

Ringtailed Coati

Ringta>ied coati (coatimundi)

name Nasua

Scientific

Nasua nasua

nasua

Family Procyonidae

and

The ring tailed coati

is

Latin name, nasua,

means "nosy one" and

intelligent

sociable. Its

Order Carnivora

aptly

Size Length head/body:

16-26 tail

in

describes the inquisitive, long-snouted raccoon.

(41-67 cm);

length: 12.5-27

(32-69 cm); height shoulder: up to 12

in

at

At

resemble a slender version of their black-

(30 cm). Male generally

Weight

7-13

lb

flexible snout; long,

banded

common

raccoon. Like

and bold

facial

tail

markings. Their forelegs are

shorter than their hind legs, so coatis always

reddish-brown to black upper

stocky,

cousin, the

raccoons, they have a distinctive banded

(3—6 kg)

Key features Long, tail;

masked

female

larger than

glance coatis (often called coatimundis)

first

in

body, yellowing underneath; coat has coarse,

seem

to walk with their

bottom held high

the

in

long hairs; distinctive white muzzle, chin, air,

and throat

accentuating their tapering

which

tail,

is

longer than their head and body. They have a Habits

Active throughout the day; females form

gangs with

Breeding

males are often

juveniles;

solitary

sensitive Births occur mainly April to June,

gestation period of about 74 days. at

4 months; sexually mature over 17 years

live

in captivity,

after

Weaned May

at 2 years.

9-15

in

is

It

sniffing

and

particularly

well adapted to

out insects and poking

bits of rotten

among

stones

wood.

Coatis are found

woodland

in

They

areas.

require vegetation for cover, since they are

mainly active during the day, although adult

Grunts and chittering used to maintain contact with group, also snarls and squeaks;

Diet

flexible.

is

the

wild

if

and

which

perhaps

2-7 young born

earlier farther south;

Voice

long, flattish snout,

threatened,

will indicate

alarm by barking

Woodland

invertebrates (such as earthworms,

millipedes,

and

snails);

frogs

and

lizards

males can also be active after dark. At night they sleep curled up

in

climbers, using their

tail

rotate their ankles

the trees. They are good to balance,

and can

180 degrees, enabling them

caught with forepaws; adult males tend to prey on large rodents, very fond of

Habitat

to descend trees headfirst.

fruit

Woodlands

Fruit-Loving Carnivores Distribution Colombia south to Argentina and Uruguay

Ringtailed coatis generally travel about 1,600 to

Status

Population abundant Generally

common

and widespread

2,200 yards (1,500 to 2,000 m) each day search of their favorite food

—

fruit.

not available, they forage the forest thrusting their snouts

among

in

When

it

is

floor,

the leaf

litter in

search of invertebrates, such as millipedes,

earthworms, termites,

snails,

and

tarantulas.

Coatis are the most sociable of the raccoon family.

Females and juvenile males gather

bands of up to 20

individuals.

in

However, males

over two years old are normally

solitary,

except

during the breeding season, and are usually

excluded aggressively from bands by the adult females. originally

28

SEE ALSO

r

(

ommon

1:22,

In fact,

they are so isolated that

it

thought there were two kinds of

Mongoose, Dwarf 1:106, Mongoose, Banded 1:110

was

I


animal

—the

coati,

which

lived in

the coatimundi, which led a solitary

place once a year during

life.

The bonds between females are strong,

and

a

young, not

just their

own

group to construct

or those closely

The juveniles are well cared

related to them.

two- to four-week period.

Pregnant females separate from the

band members help take care of the

all

™

Mating takes

groups, and

for,

a tree nest,

may

do not share food, the bands forage together

infants.

so females can watch over and defend the

days and leave the nest at

young. Band members

their

groom each

other,

parasites

climb up to the nest to assist

from each other's fur and gently

The young open

mother and the

A

using their long claws as combs, picking

nursing the

five

rest of

weeks

Numbers peaked

in

coati, its

range

the 1950s

and have since declined. Coatis are now scarce

Bonding Sessions In

in

the breeding season adult males are accepted

into bands, but

behave submissively toward

few days

allowing

young.

them

after the

young

become

to

What sounds

familiar with their

gesture

is

males do not

actually a defense

mechanism.

identify with their

young, they may attempt to

kill

and eat them

when

they

If

at other times of the year

become more

carnivorous.

is

it

in

Mexico.

seems more

threatened by habitat

disruption. Coatis are generally tolerated

because they

are born,

like a friendly

Farther south, the ringtailed coati secure, although

females. Males are also permitted into the band for a

the United States and declining

rarely

Coatis can be practice

is

reduction

show

do any harm

made

to livestock.

into pets,

although the

discouraged to prevent further in

Ringtailed coatis are

climbers

and nest

and sleep

in trees.

They

adopt two techniques for

her group.

white-nosed

Š

good

1

to join

Central America and extended

north to Arizona.

nibbling with their teeth.

in

their eyes after

similar species, the

lives in

give

young. Sometimes another female

birth to their

even after they leave the nest. Although coatis

will

where they

climbing, either

ascending hand-over-

hand or galloping up wide trunks with forefeet

and hind

feet clutching

the bark.

On

they

the ground

move through

forest at a

the

walk or

gallop, holding their tail

upright, except for the slightly

drooping

tip.

wild populations. Captive coatis

high levels of intelligence, often causing

trouble for zookeepers by unscrewing lamps

and hoarding screws and bulbs as playthings.

29


SMALL CARNIVORES

Red Fdnda The red panda

Ailurus fulgens

a puzzling

is

many features of

creature, sharing

Common name Scientific

Family

Red panda

name A

(lesser

raccoons, bears,

panda)

jrus fulgens

famous

more

its

cousin, the giant panda.

Procyonidae (sometimes considered a

member

of the bear family, Ursidae)

T-ere has been

Order

and

much

among

debate

zoologists

Carnivora

about whether both panda species belong Length head/body 20-24 tail

12-20

engtT

m

in

(50-60 cm);

(30-51 cm); height at

shoulder about 10-12

Weight 6-13 b 3-6

in

the bear ramily (Ursidae), the raccoon family, 'Procyonidae), or

(25-30 cm)

Ailuridae.

Key features /ague

darker on

DNA

red

One

m

m

mature

Voice

Norma

Diet

Bamboo

trees

and feeds there n spring

and summer

Weaned at 4 months; sexually 8 months. May live for 7 years in

1

1

8-14

m

the wild

forests,

conifer, often

and

Mountains

in

the trees.

the mixed forests that

slopes of the Himalaya

at altitudes of

between 6,500 and

5,750 feet (2,000 and 4,800 m) above sea It

gets very cold at night, and the pandas

have long, dense fur to help keep them warm.

Thumblike Structure

both deciduous and

on steep slopes

Myanmar

and

and small mammals

Distribution Himalayan 'egions of Nepa

and Yunnan

fruit, berries,

live in

grow on the lower

level.

shoots and leaves;

size,

but

pandas are more arboreal than raccoons,

Red pandas

1

y silent

Temperate

(genetic

appearance are those of a raccoon. However,

spending most o i their time up

days.

flowers; birds, eggs,

Laos.

at night

June) after gestation period of

at

captivity,

belly; tail

young born

to 4

—the

distinctly bearlike,

Arboreal and nocturnal, spends most of day

114-145

.

Like the giant panda, the red

Bhutan, India,

(Burma), and China (Sichuan

additional "thumb."

One

panda has an

of the small bones in

the wrist (called the radial sesamoid) has

Provinces)

become enlarged Status

is

the animal's general shape,

'peaks

Habitat

molecular structure)

banded chestnut and cream, face has cream and white "mask�

sleeping

Breeding

The red panda's

own

raccoor ke animal the size of a

/

;arge domestic cat. bright chestnut-colored fur.

Habits

whether they should be

classified in a family of their

kg)

in

Population unknown, but unlikely to exceed a few thousand, IUCN Endangered, CITES

structure against

to provide a thumblike

which the

five true digits

can

I

grip

and hold food. The pandas are

also able to

hold onto the branches of trees with ease.

Although they are quite average domestic

cat, red

climbers: Their grip

scamper down falling off. Yet

large, bigger

is

than an

pandas are excellent

so strong that they can

tree trunks headfirst without

when

they walk on the ground,

they tend to have a waddling gait caused by their front legs

being angled inward

Nevertheless, they can travel quite fast by

bounding along, but when frightened, they seek safety by climbing trees.

30

SEE ALSO

2:32

Pa-d^ Giant 2:98

will


RACCOONS

sim

PA'.:-

Ree pandas also

lar see.

have a general

stow

y

metabolism which helps conserve enemv Un ve the giant

oa"ca the restreted to

core

a et s not

'

:

bamboo.

eaves flowers

t.

T**ey

ete

so eat the

a.

.v

.

and bark of

roots

other p ants, and "a.e pee" kno.vn to

co"s „'"e ~ung

Occasional

;

a-c eat ,c ous sma a r

and w

also take b ds

the.

.

Âťv

I

catch

indue no

'^a s :

eggs or

'sects,

nestlings.

ma

Meverthe.ess although the, belong *o a

cam vorous group not norma

of mammal's,

hunt an

,

y

pandas do

red'

~"ev behave as

flooc

rial

n

inefficient herbivores iinstead.

Red pandas mate ea

me and

n the

.

the young are bom' about four and a half

months

later

A female gues

ole or nock crev ce to a

birth

sma fam

four cubs. She looks ah:er continuously for about a

n a tree

1

them

week

y

up to

of

moire or less After that she

spends increasing y ong periods away from the nest to feed herself Put returns regularly to suckle and

dean her

babies. After three

months

the young cubs ane ready to leave the den They

go out each

night, staying

mother, earning the I,

r

dose to

their

way around and how

to

find food. They usually disperse before the

breeding season arrives again,

the mother wi

I

if

do

they

drive he' youngsters

not,

away

Warning Displays Pandas have evolved from carnivorous (meat-eating) ancestors. However,

they are mainly vegetarian and eat relatively little

@ Deforestation croc

ss

s~ throughout

the

r.e' efforts are nov,

ce ~g -see fo conserve

~s

,

~ forests,

system'

is still

- di

cere fit the spedes.

basically that

a simple stomach' plant-eating

red penes s range

nounta

animal food. Yet the panda

in

of a carnivore, with

mammals have

Most

very long intestines

maximum

digestive

needed to cope with fibrous plant

food. Therefore

much

what the panda

of the nutritional benefit

eats

is

their inefficient digestion lot

digestive

a short intestine.

order to provide the

efficiency

in

and

s

more than

Red pandas

ive solitary

meet, they engage displays. Actions

in

ves;

and when the, oo

a variety of raccoon

undude arching

their

bacs

n

a

threatening manner, shaking their head, and

snapping their jaws shut. Sometimes they rear

up on then hind

held high

in

the

legs with their front

Red pandas face many forest

w paws

air.

difficult es

n the

r

home. They are sometimes hunted and

frequently get caught in traps intended for

musk

deer. However, the

mam

threat facing red

wasted. Because of

pandas today

pandas have to eat a

habitat as trees are felled for timber

a specialised herbivore of a

we

is

the large-scale loss of their

dear space for crops and

and to

livestock.

31


SMALL CARNIVORES

The Weasel Family Fan

,

Mustelidae: 6 subfamilies. 26 genera, 66 species

WEASELS, MINK. Mustela 16

AND POLECATS

species, including long-tailed weasel

weasel

(M

*

nivalis):

(A/f

stoat

(M

1

species, wolverine (G gulo)

Vormela

1

species,

1

species, zorilla

Poecilictis

Poecilogale Galidis

7

(/.

1

(

V.

banded weasel

weasel

(

L

.

marine otter

(L.

Lutrogale

species,

1

Amblonyx Aonyx

(L

(P.

common

weasel),

mouse burrows. Mustelids

are small

enough

form the

largest family of carnivores

to

live in

and occur

naturally

Australia,

and

smaller islands.

What

Is

a Mustelid?

same

many

patagonicus)

other groups of carnivores, but their exact evolutionary history

river otter (L.

is

somewhat

obscure.

basic ancestors as

Some

zoologists believe

canadensis):

that the skunks are sufficiently different to be classified

felina)

.

sumatrana ); spot-

maculicollis): Eurasian otter

smooth-coated otter

Cape

2 species.

the least

libyca)

(L.

(L.

lutra)

perspicillata)

clawless otter (A. capensis):

their

own

separate family

in

—the Mephitidae.

Most mustelids attack and

their

kill

own

prey on

land, but the various species of otters obtain the majority

species, short-clawed otter (A. cinereus)

1

like

Mustelids evolved from the

3 species, including hairy-nosed otter ( L

necked otter

Some, such as the

grison (G. cuja)

4 species, including North American

Lutra

sea otter, are quite large; others,

weasel (also known as the European

some

little

legs.

throughout the world, except Antarctica,

peregusna)

genera, 13 species

Lontra

T

and short

European polecat

species, African striped weasel (P albinucha)

species, Patagonian

1

vi son):

striatus)

2 species, grison (G. vittata):

Lyncodon

OTTERS

marbled polecat

species, North African

1

frenata): least

mustejids are small, fierce animals with

long, thin bodies

putorius)

Gulo

Ictonyx

(A/f.

erminea), black-footed ferret

(A/f

mgripes ), American mink

A/f

i

8 genera, 24 species

ypically,

of their food (mainly fish) from the water.

Congo

clawless

mustelids, particularly martens

Some

and badgers, are more

otter {A. congicus)

omnivorous, eating a wide variety of Pteronura

1

Enhydra

species, sea otter (E lutris)

SKUNKS

and

nuts,

other vegetable material, as well as animal food. 1

To deal with such a variety of different foods,

3 genera, 10 species

Mephitis 2

species, striped

skunk

mephitis):

(A/f

hooded skunk

mustelids have a very varied dentition (arrangement of

(M macroura) teeth).

Spilogale 3 species, including western spotted skunk

The badgers and wolverine have the most teeth

(S, gracilis)

(38), including

Conepatus 5 (C.

MARTENS

fruit,

species, giant river otter (P brasiliensis)

species, including

mesoleucus): Andes skunk

(C.

grinding solid food and even bones.

chmga)

8

species, including pine

marten

(A/f

More

typically,

the

weasels have only 34 teeth, but they are needle sharp to

2 genera, 9 species

Martes

broad and almost flat-topped molars for

western hog-nosed skunk

marten (M. martes): yellow-throated

stab their prey and tear

it

to pieces.

The honey badger

flavigula)

has the fewest teeth, with just 32. Eira

species, tayra (ÂŁ barbara)

1

Mustelids are characteristically smelly creatures.

BADGERS 6 genera, 10 species Mydaus 2 species, teledu (A/f (M Arctonyx

Melogale 4 Metes

1

Mellivora

Taxidea

stink

badger

hog badger

species, •

specif".,

Skunks are notorious

(A

collaris)

European badger

noney badger

(A/f

(A/f

badger

(A/f

personata)

smells that are so characteristic of mustelids are produced

meles)

capensis)

Mustelids are typically small, fierce animals with long, thin bodies

American badger (T taxus) legs.

Some,

the least weasel,

SF

i

Also

when

weasels and stoats smell strongly. The pungent, choking

and short

32

for the stench they create

defending themselves against attack. Even the small

species, including Indian ferret

,p>-

1

Palawan

marchei) species,

1

javanensis),

1

is

like the stoat (right),

can be

fairly large,

the smallest carnivore on earth.

American 1:48, Wolverine 1:56, Badger, Honey 1:82

but one,



SMALL CARNIVORES

m

:ne ana scent glands underneath the base of the

The scent also to

is

used not only as a deterrent to attackers, but

mark out

territorial

—

like

family includes

likely

markings than any other group of mammals: the various

skunks, for example. as the least weasel

and

several

A

and

become

the marbled polecat, and

zorilla,

may

turn white

in

and even the

sea.

except during the mating season.

solitary

in

to be hostile. By contrast, the European badger

extended family groups known as

lives’

clans. Several speciesjl

of otter, as well as the spotted skunk, are relatively social

couple of northern species, such stoat,

wide range of

exploit a

Encounters between members of the same species are

However, the

more species with black-and-white

types of badgers, the

Between them, the mustelids

Most are

most other

— are some shade of brown.

Lifestyle

habitats, including forests, deserts,

boundaries.

The majority of mustelids

mammals

tail.

winter,

loose family groups.

animals, living

in

nocturnal, but

some

Most mustelids

are

i

o

are active any time of day or night.

Mustelids have five toes on their front and back feet,

paler with increasing age. All

j

mustelids have a keen sense of smell. Their hearing

usually with a sharp claw

is

used to subdue prey or to get a good grip on tree

excellent too, but their visual abilities are probably less

well developed than

many

in

other

mammal

on each. Sharp claws can be

branches. Borrowers, especially the badgers, have stouter

species.

Ij

but blunt claws. The typical slender body of most t

,

>

These mostly southerly species

means

that they are lithe

and

agile creatures,

of mustelid share the same body

able to climb well and squeeze through small gaps.

plan as the stoat and least

However, a long, thin shape also means they are

weasel, but tend to have black, not

generally larger: African striped weasel (1); zorilla (2); little

European polecat weasel

(5);

grison

(4);

Patagonian

marbled polecat

(7J;

body

metabolic

heat.

rate,

mammals

(8).

less

at preventing the

Although they have an unusually high

which compensates still

for heat loss by

have to spend a

lot

of time

hunting to acquire sufficient food to fuel their metabolism. Small weasels their

own weight

in

may need

to

consume

them indulge

in

half

prey every day. Mustelids are

therefore always alert for something to eat.

North African

banded weasel

than shorter-bodied

generating more, they

(3);

black-footed ferret (6);

efficient loss of

brown, as the major coloration, or are

2

mustelids

"surplus"

killing

when

Many

of

prey animals are

abundant, catching more than they need and hiding the bodies

and

away

a larger

become

for later consumption. Badgers are bigger,

body

loses proportionately less heat.

lethargic in winter, saving energy

short supply. They also feed

on

a

They

when food

is

wider variety of food,

including vegetable materials that take

34

SEE ALSO

ack Footed 1:46. Mink,

American 1:52, Badger, American 1:76, Badger, European 1:78

in

'


The mink and various species of otters

cold water and

live in

so keep themselves

dense, insulating

warm

fur.

with

This has

proved attractive to hunters, and species of otter have

many

suffered heavy losses as a result.

numbers of the

Conversely, large

American mink have been reared

on

farms

fur

in

various countries,

making the species

artificially

numerous. Some have escaped

now

and

cause considerable

problems to the not least

mink

native its

in

local wildlife,

Europe, where the is

being replaced by

American cousin. Other species of mustelid

have suffered a severe reduction in

their

numbers. For instance,

the American black-footed ferret reached the brink of total extinction,

captivity

but has since been successfully bred

and restored to the

wild.

At

least

in

16 other

Š Spot-necked

otters playfully chase a terrapin in the clear waters of

Lake Tanganyika, east-central Africa. Such play reinforces social bonds

and helps young

otters perfect their hunting techniques.

species of mustelid are considered to be seriously

threatened. Gamekeepers and farmers have traditionally mustelids

killed

in

defense of their livestock,

Š

Various

members of

with a shell in

its

forepaws

exterminating several species from whole countries. Otters

American mink with

have proved to be highly vulnerable

polecat

out

in

parts of Europe

agricultural

and

due to the

industrial

pollution, dying

tc

effects of acid rain

and

chemicals on their prey.

However, many species of carnivores are protection, both nationally

and

now

given legal

internationally. Moreover,

people are not as dependent on chickens or sheep for their

more

main income as they once were and can afford to be tolerant. Predator persecution

is

no longer

widespread and automatic; and indeed, many species, such as badgers and otters, are very popular.

The European polecat has been domesticated to create the ferrets used

in

the hunting of rabbits and small game.

A

few mustelid species are sometimes even kept as pets.

in its

(6);

wolverine following (7).

(1);

least

a rabbit (3);

winter coat

marten with a bird

a scent trail

the weasel family: Indian smooth-coated otter

(5);

weasel dragging a mouse

European badger

pine

(4);

(2);

European


SMALL CARNIVORES

Common name Scientific

name

weasel)

Mustela

Family

Mustelidae

Order

Carnivora

Size

Least Weasel

Least weasel (European

common

nivalis

Th'e world's

Length

Hemisphere. However,

dense undergrowth most of the

length:

1-5

time.

in

(3-12 cm) Least weasels are solitary, elusive creatures that

Weight

1

.7-3 oz

are hard to see, partly because they

;48-85 g)

fast

Key features Long,

sleek

and short

brown and

in

body with short

tail; flat,

legs

narrow head; fur reddish-

summer, with creamy-white neck turns white

belly;

winter

in

in

northern

and are gone

easily

be confused with

species of weasel

so

However, they are

a flash.

in

more common than people

move

realize.

They can

two other

at least

North America: the long-

in

populations tailed

Habits

Solitary, territorial

animals; fierce predators;

very active both day

Breeding

Up

to 2

and night

all

weasel (Mustela frenata) and the stoat, or

ermine (Mustela erminea).

year round

At a glance

all

three species look similar,

young born each year after gestation period of 34-37 days. Weaned at 3-4 weeks; females sexually

animal. The long-tailed weasel has distinct dark

mature

facial

live

litters

at

of 1-9

4 months, males

up to 10 years

at

in captivity,

8 months. usually

May

under a

but the least weasel

is

by far the smallest

markings, and both the long-tailed weasel

and the stoat have

bushy black-tipped

longer,

year in the wild tails.

Low

trill

a male

screech

to signal a friendly meeting

and a female;

when

between

Habitat

and

The European variant—the European

common

weasel

loud, harsh chirp or

disturbed or ready to attack

Mainly small rodents, especially mice; also rabbits, lemmings, moles, pikas, birds, fish, lizards,

insects

—was once regarded as

separate species, but

same

the

is

now

a

considered to be

species as the least weasel.

World's Smallest Carnivore

Almost anywhere providing suitable cover

Barely longer than a

rat,

and access to rodents, including meadows,

smallest carnivore

the world and the smallest

in

farmlands, prairies, marshes, and woodlands

of Distribution Northern Hemisphere: Canada, Alaska, Siberia, Japan,

northern U.S., northern

all

the mustelids.

Population abundant

One

of the

more

numerous small carnivores

the least weasel

is

the

has a long, sleek body

It

with short legs and a short

and narrow with

Europe, and Russia

Status

tail. Its

large black eyes

head

is

flat

and

prominent, rounded ears. During the

summer

months weasels have

a reddish-brown coat with

a creamy-white patch

on the neck and

early

fall

it

is

winter coat.

belly. In

replaced by a lighter-colored In

some northern weasel

populations, particularly

colder climates, the

in

coat turns completely white

in

winter.

The white

color gives the animals natural camouflage

against the

snow and

helps

them avoid

detection by predators.

Weasels have acute senses of

and hearing, and often stand on to scan their surroundings.

36

SEE ALSO

is

an elusive creature that

it is

hardly ever seen, remaining well hidden in

is

in

(17-25 cm); tail

smallest carnivore, the least weasel

abundant throughout a wide area of the Northern

head/body:

7-10

M us tela nivalis

•t

1:40,

Lemming, Norway 7:9

sight, smell,

their hind legs

They are incredibly


rl

WEASELS

LEAST WEASEL

quick and agile, and often dart around erratically or

bound along with

arched and their

their

backs

held straight out or

tails

pointing slightly upward. They have five clawed

toes on each foot, which they use to grab their prey and to climb trees. Their climbing ability

enables them to reach birds' nests, where they

can consume both eggs and chicks. They are also strong

swimmers. Weasels are powerful

animals for their size and are capable of

running 300 yards (275 m) while carrying a large

mouse

in

their

mouth!

Huge Appetites Weasels are so small and dynamic and have such a fast metabolic rate that

must eat almost

survive, they

weight

in

order to

in

half their

food each day. This means that they

must catch about two mice or one day lot

body

As

just to stay alive.

fat vole per

a result, they

spend

a

of time hunting, although they frequently

take short rests

in

one of

their dens.

Weasels are specialized predators of small rodents, but will also take birds, lizards, insects

whenever the opportunity

long, sleek

into the smallest crevices

and hunting rodents down

its

the weasel's head

body.

If

the rest of

arises. Their

body means that weasels are well

adapted to squeezing

In fact,

and

it

can squeeze

its

body

will

their

is

own

burrows.

the widest part of

its

head

into a hole,

follow without getting

stuck. Access to such tunnels provides weasels

with shelter from predators and also allows

them all

to hunt at any time of the day or night,

year round. They do not hibernate and can

hunt even under deep snow.

Weasels are renowned for being killers.

They catch small

prey,

efficient

which they

kill

with a few swift bites to the back of the neck. If

they encounter their prey head-on

tunnel, they

kill

it

in

a

with a crushing bite to the

windpipe. Weasels also hunt larger prey, which

Š Weasels have extremely acute senses of hearing,

and

smell.

They

will often

sight,

stand on their hind

legs to scan their surroundings.

37


SMALL CARNIVORES

they stalk quietly and then pounce on the victim's

back for a

the base of the

series of precision bites to

Males, which are often

skull.

twice the size of females, are more

hunt larger

likely to

while the females mostly look

prey,

for small rodents.

The weasel's mode of killing

whatever

whenever

can,

it

survival involves

can. Faced

it

with an abundance of mice, the voracious

weasel follows the only pattern will

at

knows and

it

more than

kill

any one time.

it

It

can eat

sometimes

stores surplus food for future

meals its

a side

in

chamber

den. Weasels are extremely

and can

versatile

wherever there shelter

them

live in

They use

log.

Weasels

live in a

variety of habitats,

including thickets

and

woodlands, as long as there

is

a

piles,

mountainous

junk heaps, abandoned

and burrows dug by mice, ground

buildings,

squirrels, or

may

rock

do not normally

They usually make

regions. in

and open

wetland areas, sandy

deserts, or

dens

for

to reproduce

country, but

their

suitable

is

forested, bushy,

a rotten

live

and enough food

successfully.

A weasel by

off

chipmunks.

chambers with grass or

nest

line their

sometimes the

fur

colder climates they

In

and feathers of

prey.

good supply of

suitable prey.

No Time

to Lose

Weasels only have a short reproduce frequently and

life

span, but they

prolifically.

food

If

supplies are high, weasels are able to take

advantage of the favorable conditions, and

diligently for her

female weasels can have up to two

By seven to eight weeks the cubs begin to

year.

The weasels usually breed from

spring to late

about as

litters

few

five

as

litter

size

accompany

early

summer, and the pregnancy

weeks. The

per

lasts

may range from

one or two young to as many as 20,

depending on food average of four to

supplies, although

six is

an

their

38

the

same

deaf.

They only open

eyes after 30 days. The mother cares

SEE ALSO Marten, American

1:48, Mink,

up,

kill

later

mother on foraging

trips

efficiently for themselves.

and

A few

the family group begins to break

and the young

their

rapidly.

mother's

start to disperse

home

away from

range.

Weasels are heavily dependent on rodent

an American one cent coin and are wrinkled,

and

weeks

most common.

Newborn weasels weigh about

pink, naked, blind,

can soon

their

young, which develop

as

populations Often

in

the spring,

populations are low, there in

is

when

rodent

an associated peak

weasel mortality, probably through

starvation.

American 1:52; Lynx 2:40, Coyote 2:58,

Fox,

However, weasels also Red 2:64

fall

victim to


predators, particularly owls

and martens, but

also coyotes, lynx, hawks, cats, foxes, mink,

even stoats. Weasels are also frequently by

traffic as

and

killed

they dash across busy highways.

nesting birds.

In fact,

weasels are often

killed in

a harvest

traps intended for stoats.

Weasels are superbly check populations of

efficient at

many

keeping

hundreds of mice

a year to

crawling

Least weasels are often regarded as vermin by

feed herself and her offspring. Any

gamekeepers and poultry farmers, and have

game

been widely hunted and trapped. They are

weasel's value as a destroyer of pest spec

thought to

kill

young game

birds but are not

birds or poultry

which cause untold

is

far

in

to

outweighed by the

losses to

nest.

is

and the

capable of

down burrows

and squeezing

into

crevices in pursuit

les,

growing crops and

cons dered so serious a threat as stoats, which

stored food. Without predators

can devastate fragile populations of ground-

such losses would be even greater.

;

damage

mouse

the weasel's diet,

animal

weasel

investigates

Rodents form the bulk of

One female

Farmer's Helpers

kill

in

species of rodents

that can be harmful to agriculture. will

A weasel

like

the weasel

39


Stoat The stoat range

Mustela erminea

—or ermine, as

is.

much of its most widespread member of the

the

mustelid family.

name

called in

range includes a wide variety of

Its

habitat from windswept arctic tundra to dense forest.

short-tailed weasel)

Scientific

it is

M usreJa erminea

Family

Muste dae

Order

Cam.vora

The stoat almost certainly evolved as a rodent-

catchmg spec

Leng f ~ head/boc. 7-12.5 length

5-5

1

Weight

in

Key features

nch brown with cream on

brilliant

round Habits

wh*te

m

bei'y;

.vith

tail;

body

may

turn

and long whiskers

and clrnb

well; active

and

solitary

and

rats

S rgle

m

tter of

agile; a territorial,

jsually 4-9>

young born

spring after gestation period of 10

including delayed

months, males at 12 months. 0 years

m

captivity,

May

many fewer

at

live

in

at

Shr

Diet

Carnivorous, includes small especially rodents

and

inside a tunnel.

forests

from

arctic

spine

Its

much

to travel

A

flexible,

is

faster than

its

stoat can streak

sudden leaps and bounds, and change an

instant.

It

direction

can also climb trees and rocks

and swim extremely

well. Stoats

found up to 50 feet

(1

have been

5 m)

in

(800 m) offshore

trees

and over They have

half a mile

crossed even larger expanses of water,

in lakes.

the wild

mammals,

rabbits; also birds

apparently unaided, to colonize small coastal

C

islands in parts of their range. Stoats are light

their explorations at high

enough

speed and not

to run along the surface of fresh snow,

and

but can also

move below

the surface, out of

and sheltered from the wind.

sight of predators Varied,

enough to follow

2-3

eggs. repMes and amphibians

Habitat

The

burrows and to turn

rabbits into

it

slim

s

up to

squeaks when excited

Voice

ii

almost equally

it

months

Weaned

mplantation)

6-8 weeks, females sexuaily mature

1

physical adaptations

through long grass at amazing speeds, take

predator

3-18

and

around

in

Breeding

its

short iegs might suggest.

Mainly nocturnal, terrestnal. but can swim

fierce,

but

smuous body

which adows

wmter; smat head with

ears, large eyes,

st,

and hunting techniques make

animal's

ong-bodied animal

.•."e,

a

effective n pursuit of other vertebrate prey. g)

short legs and longish, black-tipped fur

tail

(4-12 cm)

5-12 oz 42-340

'

17-32 cm);

ir

Stoats carry out

surprisingly

bum

up a

great deal of energy. The

tundra and moorland to

and meadows

Regional Differences

long, thin

body

inefficient

when

is

also

it

Distribution Northern Hemisphere ^Eurasia and North

America from within Arctic

Circle to latitude

There are as of stoat,

Status

Population abundant. v

Common

despread, but trapped for fur

as

29 recognized subspecies

most of them from North America,

where they are often known as

and in

many

some

areas

short-tailed

weasels. They are distinguished as

much by

geography as by any obvious physical characteristics.

As a general

rule,

American

stoats are smaller than those in the Old World,

and throughout the

stoat's

geographical range

males are bigger than females, sometimes twice the size Perhaps the most notable regional difference

is

that in higher latitudes stoats turn

white

m

zones

retain their

winter, while those in

brown

color

more temperate all

year round

Stoats are small, but they sometimes use surprisingly large

40

SEE ALSO

all

1

.

Arctic 2:70, Bear Family,

home

The 2:82

ranges

An

active

male

to conserving heat.

comes


WEASELS


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*3


SMALL CARNIVORES

Mas tela putorius Common name Scientific

Po^cat

name

A

Du tori us Family

Mustelidae

Order

Carnivora

Size

Length head/body:

12-18 tail

Polecats were persecuted relentlessly in the past as a means of protecting gamebirds and chicken houses

Vfusre/a

from

expanding

length:

5-6

in

Weight

female

nobility in

1-3

reputation for being bloodthirsty

lb (0.5-1 .4 kg)

in

were therefore believed to be in

summer, but

winter; bold, dark

mask pattern

on face throughout the year Mainly nocturnal;

solitary;

One

litter

young born about June mature

at

1

forages along the

Voice

Diet

after gestation period 1

in

the wild

May squeak and is

hiss occasionally, but sound not used for social communication

Mainly rabbits and voles; also birds, frogs, occasionally eels caught

whatever

is

in

wet

most abundant and

I,

Later, large

and many thousands were

grass;

killed.

numbers were trapped by

gamekeepers, especially

Weaned at month; sexually year. May live up to 14 years

captivity, 5 in

a threat to

England they were declared vermin by Queen

per year of 3-7 (can be up to 12)

of 42 days.

and

killers

livestock, especially chickens. In 16th-century

Elizabeth

ground; occasionally climbs

Breeding

medieval times. They also had a

with a long,

cat,

sinuous body; almost black pale cream

fur,

which was highly prized by the European

Key features Resembles short-legged

Habits

areas.

Polecats were often trapped for their fine

generally larger than

\

many

in

(30-46 cm);

in

(12-14 cm). Male

1

predation. However, present populations are

the 19th century.

in

Polecats are particularly vulnerable to traps, especially the old leghold traps used to catch rabbits.

The traps were often

or gaps

in

hedges and

polecats dive

in

set in rabbit holes

walls, exactly

and out searching

where

for prey.

Such traps were sometimes deliberately used to catch polecats, but

many more

polecats were

killed as a result of accidental trapping.

easiest

to catch

Habitat

Young

forestry plantations

and woodland, but mainly on farmland where there is plenty of cover in hedges, walls,

and old buildings

Distribution Western Europe from Britain and Spam east to the Black Sea and Baltic countries

Status

Population abundant Previously eradicated from Scotland and most of England, but now

Respite from Persecution Although polecats were once widespread and abundant, numbers decreased the

World War brought

First

from the

activities of

sharply.

However,

welcome

a

gamekeepers

reprieve

in Britain

and elsewhere, and soon polecat populations

began

recolonizing, thanks to reduced persecution

to increase.

survived parts of

was

in

Enough polecats had

remote areas to begin recolonizing

Wales and England, but the species

already extinct in Scotland. Polecats are not

and

much

parasites, but often

affected by diseases

fall

victim to poisoning,

usually as a result of eating poisoned rats.

Modern

many

rat

polecats

poisoning threat

poisons are very powerful, and

is

live

routine practice Another

comes from road

may be

see ALSO Weasel.

traffic.

annihilated as they play

attempt to cross

44

around farmyards where

Least 1:36, Stoat 1:40, Skunk. Striped 1:84

it.

However,

it

Whole in

rat

modern families

the road or

appears that


WEASELS POLECAT

new

forestry plantations provide excellent

polecat habitat, especially

and grass provide

where young

trees

huge populations

shelter for

of voles, a favorite food. Further assistance has

come from

the recovery of rabbits after the

disease myxomatosis devastated populations

the 1950s.

Britain in

The

ferret, a

polecat, has

domesticated form of the

been bred

in captivity

early as the 4th century B.C. initially is

in

now

It

from as

was

raised to assist with hunting, but

a familiar household pet,

especially in the United States.

Ambush Techniques Across Europe polecats seem to be a .

lowland species. However, urban areas and huge,

with

open arable

fields

cover and few prey

little

animals are uninviting habitats. Polecats have acute hearing and

|

good

night vision. They hunt mainly

after dark,

spending about four hours

every night searching for food, mainly

ambushing

rabbits

and

voles. Polecats will also

eat eggs, small birds, frogs, and the occasional eel

found wriggling

usually

wet

in

amble around

grass.

The animals

an unhurried manner,

in

pausing to investigate interesting sounds and smells,

but they can also

bounding

gait

move

rapidly with a

and with the back arched high.

Polecats are playful creatures, but normally live

alone, each occupying a

about 250 acres (100

may

home range

ha), within

of

which they

use five or more different dens. Polecats can produce a

pungent scent

from large scent glands. The scent

mark stones and

is

used to

logs within their territory

and

probably helps the animals recognize each other.

The scent glands

protection:

also provide

Few animals

will try

some

to attack

polecats because of their strong smell.

0 Polecats have luxurious

fur that

cream

mask pattern

year round.

all

dark colored

in

the summer. Their faces have a

winter, but pale

in

is

45


SMALL CARNIVORES

Black-Footed Ferret

Musteia nigripes

The black -footed ferret specialized

in

hunting prairie

dogs and suffered heavily when these animals were eliminated from agricultural areas. The ferrets ha\'e 3lack-footed ferret

been reintroduced

Musteia nignpes

but

Common name Scientific

name

its future is

to the wild from captive-bred stock,

precarious.

Family

Mustelidae

Order

Carnivora

Black-footed ferrets were never

Length head/body: 14-18

lived

Size

in

(36-45 cm); tail length: 5-6 in 2- 5 cm) Male generally about

(

1

1

more than

10%

prairies

2-2 4

lb (0

9—1

3.5 miles (6 km). They used to be

found widely across the American short-grass

bigger than female

Weight

common. They

alone and were often spaced apart by

from Texas to beyond the Canadian

kg)

1

border. Over

90 percent of the

prairie dogs,

but the animals would also

ferrets'

food was

Key features

Slender animal the size of a small cat; pale yellow with short black legs, black mask, and

mice and other small

tail tip

Habits

get by on prairie

underground Breeding

what

Usually nocturnal; spends the day

3ne

tter of

March-Apnl

up to 6

young born

after gestation period of

1

in

(called a town), living as a

residence."

The black-footed for

Voice

Normally

Diet

Mostly small rodents, especially prairie dogs

Habitat

Grasslands

prairie

food and shelter has contributed to

dogs

its

silent

in

the wild. During the 20th

century whole prairie dog towns were plowed over or eliminated using traps, gassing, and

Distribution Formerly widespread

Midwest from Texas

to

in

poisons to

the American

Canadian border,

now

make way

for agriculture

Growing crops

ranching.

survives only as reintroduced populations in

prairie

Montana. South Dakota, and Wyoming Population about 500, mostly I

on

ferret's reliance

precarious situation

CITES

quite a small

in

Disappearing Food Source

1

the wild

Status

single ferret could

could catch

it

dog colony

"predator

A

45

Weaned at month; sexually mature at year May live 12 years in captivity, similar in

days '

(usually 3-4)

prey.

kill

since

dogs was

and

successfully

impractical,

among

the

and farmers found

their tractors often got stuck in collapsed in captivity;

burrows.

1975

Prairie

dog towns were

hazard for horses and their

broken legs and other tripping over dirt In

Kansas

dogs

with

riders,

injuries resulting

mounds and

many from

tunnel entrances.

—a former stronghold of

prairie

—over 98 percent of the towns were

eliminated

As a ferret

was

also a serious

in

fewer than 100 years.

result of the

numbers

also dwindled,

believed to have

of ferrets

and

demise of the

become

their tracks

prairie

dog,

and the species extinct

Sightings

were reported from

time to time, and a small population was '

discovered

46

SEE ALSO

(f

1:44

Dog, Black-Tailed 7:56

in

South Dakota, but

it

had died out


WEASELS BLACK-FOOTED

by 1974. -

Black-footed ferrets

commonly

inhabit the

In

of black-footed ferrets

Wyoming.

was found on

began, but the white-tailed

prairie

(rodents of the squirrel

which they mainly fed were

hit

over part

of the tunnel system as

predator

in residence.

into captivity to

a ranch in

extinction. Their

Detailed studies of the animals

burrows of prairie dogs

family), taking

Meanwhile,

1981 a substantial wild population

few

a

form an insurance against

numbers slowly

enough

by disease and

wild.

and

some back

Wildlife

into the

Over several years a few hundred were

restored to

the wild.

and

for the United States Fish

themselves caught canine distemper and died left in

built up,

total

by the end of 1991 there were over 300,

dogs on

Service to begin releasing

1987 there were none

had been taken

ferrets

suffered catastrophic losses. The ferrets

out. By

FERRET

Montana and South Dakota. They

began to breed, although rather

slowly; but

many

ferret

and the black-footed

died,

not securely reestablished on the

is still

prairies.

While reintroduced wild populations were

numbers

suffering setbacks,

of black-footed

continued to increase. By

ferrets in captivity

1996 there were more than 400 and today the animals are kept zoos and

in captivity, in

many

wildlife conservation centers.

The black-footed and

legal protection,

now

ferret

enjoys

dogs are no

prairie

longer massacred. The current problem challenging conservationists

is

that the

entire population of black-footed ferrets

comes from only

Wyoming inevitably

in

few animals removed from

a

the 1980s. The population has

become

inbred, resulting in limited

breeding success and poor survival of offspring. It

remains to be seen whether the newly

I restored populations ’

this

manage

will

to

overcome

problem and increase to form a viable wild

population. Even

if

they do,

black-footed ferrets

it

unlikely that

is

ever again be

will

widespread, because prairie dogs their existence relies

—on whom

— have long since

disappeared from huge areas of the Midwest.

Widely Dispersed Black-footed ferrets have offspring

in

litters

up to

six

the spring. The young emerge from

underground

in

early July, having

when

until

disperse.

Young females often

fall,

Adult males do not

live

stays

the young males

together

the

grown almost

The family

as large as their mother.

raise

of

stay nearby.

with their family or help

them. Black-footed

ferrets

widely dispersed to avoid too trying to feed off prairie

dogs

need to be

many animals in

any one place.

47


SMALL CARNIVORES

American Marten

SalnPs

A

Martes americana

charismatic small forest predator, the American

marten has suffered heavily from the

and

trappers

Common name

activities

of

also from habitat loss.

Americar

(American pine marten, American sable)

American martens are typically associated with Scientific

name

Martes americana

Family

Mustelidae

Order

Carnivora

Size

northern forests, particularly those dominated

by mature spruce and in

Length head/body: 20-27 length:

7-9

in

in

(50-68 cm);

trees.

They also occur

but the most distinctive

forests,

features of their habitat are the complex

mixture of different species, the different ages

Weight 10-44 oz (280-1,250

If

tail

(18-23 cm)

deciduous

fir

and the presence of glades and

of the trees,

Male

g)

generally at least a third bigger

clearings.

Such habitat generally has abundant

than female

and diverse Key features Slender

cat-sized animal with short legs

and bushy

tail;

fur ranges

from pale brown to

prey.

The martens

using different areas at different times of year,

depending on food

almost black with an orange or yellowish

move around

will

availability.

throat patch

Habits

Breeding

Active throughout the year and also at any

Disappearing Habitats

time of day or night; climbs well

Martens are found across much of North

Up

to 5 (usually 3)

young born once

the spring after gestation period of

Weaned 1

Voice

Diet

in

live

1

America from Alaska through the forested parts of

Canada and

States.

7 years

in captivity,

east into the eastern United

silent,

At one time they were also

common

the wild

sometimes makes chuckling

in

fairly

the southeastern United States.

However, the harvesting of trees and clearing of forests to

Wide

deprived the animals of enormous areas of

variety of small animals; also insects,

and seeds

Deciduous and coniferous forests

also Rocky

Mountains and

lustrous fur of the

Sierra

Nevada

Population abundant Widespread but elusive, rare

make way

for farmland

have since

suitable habitat. Moreover, the dense

Distribution From Alaska eastward to eastern Canada;

Status

in

month.

noises or gives the occasional scream

fruit,

Habitat

15

Normally

a year

6 weeks; sexually mature at

at

5-24 months May

at least

1

and declining

in

some

places

marten made

target for fur trappers,

easy to catch. As a

it

and

a prime

and the animals were

result,

martens were

eliminated from the southern part of their natural range

many

and have become quite

M

rare in

other parts of their range, too.

American martens do not hibernate and are active

throughout the

year.

They hunt mainly on fcv

the ground, although they can also climb well.

They are perfectly capable of swimming, but do not enter the water unless absolutely necessary, preferring to cross streams by

overhanging trees and

way

of

logs.

American martens have a very varied

and more than 100

48

SEE ALSO

1:50; Mink,

different types of

American 1:52. Vole, Southern Red-Backed 7:94

diet,

food have

8:

4

-


WEASELS AMERICAN MARTEN

substances

—

precisely to deter animals such as

martens. However, the American marten

wholly carnivorous and quantities of fruit

in

will also

the

fall

eat large

and even

seeds. The indigestible remains of

its

can be seen as chewed fragments

ripe

varied diet

in its

shiny

open

black feces, which are often deposited on tracks

and prominent

mark out

its

not

is

probably to help

logs,

territory.

Martens are extremely active creatures and

move

can

when

rapidly

they need

They

to.

trot

and bound around, stopping frequently to investigate likely places to find food. their foraging

done

is

early in the

in

the evening, although martens

at

any time of the day or night.

Most

of

day and

late

may be

active

Delayed Implantation Martens are

territorial,

and each one

lives

alone, except during the breeding season.

male may

live

with a chosen female for a

couple of weeks

in

time the animals

will

in

mock

the summer, during which

with several partners

becoming quite result of the loss of to farming

and

rare in places as a its

forest habitat

egg remains

the wall of the uterus and developing

normally. until

mammals such

As a

However, they

will also

result,

food, and the weather

many

fallen log.

They grow

ground

mother. Males

mice

is

pocket gophers, and even

due to the

common and

fact that such prey

easy to catch

forest clearings.

Martens

in

and

eat birds

if

they can catch them and any large-bodied insects,

plenty of

challenging.

in

the family.

den among

a

in

rapidly,

except those that produce evil-tasting

and by

are almost as large as their

will

grow

to at least one-third

bigger than females. The young face few predators and are nimble

is

the long grass of

will also

is

boulders, a hollow tree, or the shelter of a

eat mice and

hares. Their preference for voles

less

is

there

as five babies

The offspring are born

midsummer they

snowshoe

when

average three young, but sometimes

occasionally larger animals, such as chipmunks, squirrels,

the young are not born

the following April,

Litters

as red-backed voles.

fertilized

as

200 days before implanting

for over

there are as

small

in

delayed implantation. The

logging.

been reported. Their main prey consists of

only a month, but births

known

itself in is

lasts

The

a season.

in

midwinter are avoided by a process

dormant

Š The American marten

and indulge

play together

They mate many times, often

fights.

pregnancy

A

likely attackers.

time

when

it

is

enough

However, winter hard to find

deep snow. Nevertheless, and some martens

live

for

is

to escape a difficult

mammal

prey under

survival rates are high,

up to

1

5 years.

49


SMALL CARNIVORES

Fisher The fisher

is

Martes pennanti

a large marten.

an opportunistic

It is

predator and one of the few animals that hunts porcupines. However, trapping for the fur trade has

Common name Scientific

name

eradicated

Mustelidae

Order

Carnivora

Fishers are medium-sized

Length head/body: 18.5-29.5

Weight

12-16.5

length

tail

Male 7.7—1 2

lb (3

(47-75 cm);

in

5—5.5

kg), occasionally

up

(2-2.5 kg)

Key features Cat-sized weasel with short bushy

long,

relatively large,

tail;

head

rounded

is

long body,

legs,

wedge shaped,

ears; coat

Active any time of day

in

Breeding

probably

fishers

litter

do not eat

named by American

mature for

year,

1

10 years

males

settlers

who

fitches or fitch ferrets.

Fishers are

sometimes

because of

their luxuriant fur.

called

American sable

Growls, hissing coughs, or

clicks,

but

which

means they

will

eat any animal they can catch.

few animals

f

young

trees by

that will eat

makes them popular with can

removing

damage

or even

their bark. Fishers

also eat the flesh of large

dead animals, such as

deer and moose.

Porcupine Hunt

When

hunting porcupines, a fisher travels long

normally silent

distances on familiar routes, inspecting most of Diet

Porcupines,

snowshoe

hares,

and small

the porcupine dens along the way.

If

it

rodents; also carrion, birds, eggs, insects, reptiles,

Habitat

Mature

and amphibians; some forest

fruit

and nuts

encounters a porcupine, there

drawn out attack

and swampy woodlands

is

usually a long,

that can last over half an

hour. Porcupines are usually attacked Distribution Northern

U

S

extends south

Status

and Canada, in

ground.

up and

Population unknown, perhaps tens of

thousands Protected

in

some U

S.

states

If

one

is

up

try to force

the porcupine attack.

The

falls

to

keep

its

a tree, the fisher will climb it

to the

end of a branch.

to the ground,

its

face.

When

During such charges

which has no protective

it

it

is

quills,

n,

soft,

its

tries

face

spiny

flailing its its

and the

when

exhausted, the fisher

and attacks the

i

to

can, the

exposes

attacks repeatedly. Eventually,

SEE ALSO

open

The porcupine

porcupine charges backward,

porcupine

is

back toward the fisher and

close to a tree trunk.

tail.

it

If

fisher circles the porcupine,

attempting to bite

50

on the

distribution

mountainous regions

porcupines

a branch

attacking

live

the wild

in captivity, similar in

will kill tree

ground.

Fishers are generalized predators,

kill

May

trees.

end of

spring

sexually

They

branches of

which are also known as

352 days (including

at 2 years.

in the

by knocking them off the

foresters, since porcupines

2-4 months; females

at

at

They were

fish.

and even hunt

noticed their similarity to European polecats,

trees with ease

an uncertain period of delayed implantation).

Weaned

the wild. Contrary to

alive in

porcupines. Their diet

after gestation period of

Voice

name,

seen

meeting during the mating season; mainly

in

Fishers are excellent

some of their prey

rarely

They are among th

per year of 1-5 pups born

0

and are

hours; territorial and mainly solitary, only

One

largest

dark and

is

short bouts of 2-5

ground dwelling, but climbs

mammals and the

climbers

vanes between sexes, seasons, and individuals

Habits

States.

of the martens. They are shy, elusive creatures

their

(30-42 cm)

in

to 19.8 lb (9 kg); female 4 4-5.5 lb

and

from many parts of the United

Marxes pennanti

Family

Size

it

Fisher

face, fisher

the

flips

it

over

unprotected underbelly.

American 1:48, Porcupine, North American 8:20

All

and

them on the


come together

briefly for

breeding, and the males do not stay to help rear the family.

spring

in

other prey bite to the

fisher

the

is

killed

with a quick

back of the neck. The

consumes the porcupine from

liver,

and leaving the

feet,

and

skin, large

bones,

During attacks on

intestines.

porcupines fishers freguently get their body.

in

quills

They do not seem to

cause serious discomfort, but can be lethal

if

they puncture a

vital

organ.

Fishers travel fairly long distances

when ),

hunting and use temporary dens for

when

particularly

logs,

trees or

progress

is

made

brush

piles,

or tree roots,

are rarely used for

more than two

Fishers are territorial and, like

mark

and scent from flank,

in

hollow

ground burrows, or under snow. They

mustelids,

hollow trees or on rocky

Although blind and helpless

months and

after four

at birth,

own

their

kill

prey

are fully independent by

their first winter.

Threats to Survival Threatened by the fur trade and logging, fishers have been eradicated from many parts of their original range, particularly

coastal parts of the United States. Their

persecution to

still

Because they

most severe

will

in

the early furs,

but

silky pelts of

the

eat carrion, fishers are

easy to bait and trap. They are legally in

some

still

states. In others

hunted

they are

listed

as threatened

and given protection.

many

also suffering

from the effects of deforestation.

anal, cheek,

abdominal, neck,

like

is

also

used to communicate during the breeding

find receptive females.

its

females are particularly valuable.

Many

will travel

at

continues today. The black,

ranges with urine

when males

was

mid-1900s owing to the fashion for

home

their

along the western

or three days.

and feet glands. Scent marking

season,

in

they grow guickly. They can

by heavy snowfall. Dens are usually

under

in

eating the heart, lungs, and

belly,

stuck

are born

nursing dens that are usually well

concealed high ledges.

Young

long distances to

Males and females only

to

down

of the old-growth forests

make

their nursing

for timber

if

which they

dens are being cut

and other commercial

Fishers dislike crossing

vulnerable

in

Fishers are

uses.

open areas and so

the forests they inhabit

broken into small patches.

are

become


SMALL CARNIVORES

American

Mink

Mustela vison

A widespread waterside predator in North America, thi. Amt lean mink has also become established i

Common name Scientific

name

m parts of Europe, American mink

successful but

Mustehdae

Order

Carnivora

American mink are widespread and

Length head/body: 12-18.5 tail

Weight

9-4

1

(0

5-9

length

lb (0

dark-brown

in

(30-47 cm);

substantially smaller than otters

(13-23 cm)

9-1 8 kg); female 1-1

darker .8 lb

pencil.

They are

Mainly nocturnal; swims and dives; uses

and smell

lairs

among

tree roots at water's

One

distinctly unpleasant.

Mink are

usually associated with slow-

rivers

and lowland

lakes, preferring

4-6 young born April-May after gestation period of 39-78 days, including a

places

variable period of delayed implantation.

bankside vegetation. Each has a hunting

litter

of

Hisses

when

lakeshore.

about

threatened;

some

high densities, as

Mostly lowland areas beside

on rockpool

and

Distribution .anada. eastern and most of central U introduced to Europe: in Britain. France, Spain, Ireland, Scandinavia,

common

in

upland

places they have established

many

mink behave rather

rivers, lakes,

less

colonies along the coast. Here they can reach

mammals, waterside birds and their eggs, also some invertebrates such as beetles and worms, especially along coasts

ponds, also marshland and along seashores

plenty of overhanging

half a mile of riverbank or

may scream

small

is

Mink are

areas, but in

defiantly in self-defense, but usually silent Fish, frogs,

where there

territory of in

the wild

mile (1.6

km) of

fish,

as three individuals per

coastline. like

Along the seashore

coastal otters, feeding

but also climbing steep grassy

slopes to raid gulls' nests.

cliff

S. Italy,

Lone Rangers

and Iceland

Mink are Status

wet. Their droppings are easily

burrows and

Weaned at 5-6 weeks; sexually mature at 2 years May live for 10 years in captivity, 2-3

Habitat

and much

deposited on rocks and logs at the water's edge

flowing

Diet

when

about the diameter of a

pointed muzzle

for itself

Voice

common

recognized, being black and cylindrical and

short-legged, glossy black or

edge, also rabbit burrows, but does not dig

Breeding

fairly

appearing almost black,

in color,

especially

cat;

proving a

across most of North America. They are

5-0 8 kg)

Key features Resembles

Habits

in

it is

unwelcome invader

Mustela vison

Family

where

:

Population abundant Increasing

in

Europe

active at dusk

and

after dark,

bounding around on land and swimming and diving underwater. They are unsociable

A

creatures, territory.

and each one tends

Male

territories

to live in

do not

its

own

overlap, but

often include parts of the territories of one or

more females.

spring

In

out on long journeys

may

travel

There

52

SEE ALSO

1:58; Ott>

is

new

mate mink

no

of the males set

search of females and

widely across the countryside,

helping colonize finding a

in

some

pair

areas.

still

However, even after

do not

set

up

families.

J

bond, and after mating the

an River 1:64,

Otter, Sea 1:72

l t


WEASELS AMERICAN MINK

animals litter

year, usually in April or

each

average

produce only one

apart. Females

live

size

litter

four to

is

six

May. The

young, although

mink can produce many more. The

captive

two months

babies spend up to

mother's nest, normally

dense tree

in

in

their

burrow or among

a

They are weaned

roots.

about

at

five

and the family disperses soon

to six weeks,

grow

afterward. The babies

rapidly to reach

adult size before the end of the year and are

capable of breeding the following spring.

Females can

few reach

still

breed at seven years, although the wild, and most mink

this

age

in

die within the

first

three years of

life.

Dangers Faced Young mink tend

to disperse

away from

mother's

sometimes

traveling

territory,

than 6 miles (10 km) to

live.

new

established a

mink tend to stay

territory,

the

in

same

patch well, since they are

in

place,

know

often for several years. They need to

home

more

search for a place

in their

Once they have

their

their

constant

danger from gamekeepers, farmers, and trappers.

Thousands are

may drown

killed

each

year.

On

traps set to catch fish.

in

other hand, mink have

little

to fear

Others the

from natural

predators and are well able to defend

themselves

attacked by foxes or cats.

if

Perhaps that

is

why

they are often active

broad daylight, although their main takes place after dark,

more

beetles,

activity

prey animals are

be found.

likely to

Mink eat

when

in

and

fish

birds,

worms, and other

but also take

invertebrates.

Most

of their prey consists of small creatures, but

often mink

will

attack rabbits.

they

In fact,

will

eat almost anything apart from fruit and other plant material.

they spend

less

Mink are such successful hunters than 20 percent of their time

away from the den. The away

are safely tucked

Š Mink are

rest of

the time they

asleep or grooming their

solitary creatures

and only meet

However, after breeding, they do not set up the female raises the

young

to breed.

families,

and

alone.

53


SMALL CARNIVORES

produced, ranging from silver-white to cream, as well as the natural dark chocolate-brown.

The mink industry expanded

and

in Britain

Europe during the 1950s, producing more than a quarter of a million skins per year. But the

mink

is

able to climb and squeeze through

and

small gaps,

many

inevitably

escaped. The

animals often proved adaptable and survived successfully

in

the wild, despite attempts to

Mink spread widely and within

eradicate them.

30 years occupied most also

of Britain.

found over most of

Iceland,

They were

Norway, and

Sweden. Today they continue to spread eastward

established

now

mainland Europe and are

in

also

in Ireland.

Invaders of Europe The American mink's extraordinarily successful invasion has

mammals

added another species to the

of Europe, but

accompanied by serious sleek

and

lustrous

old rabbit

The den may be

burrow among rocks or

brushwood, but edge.

fur.

it

is

a separate

may

entrance underwater. Each mink several

Mmk

dens at different times of the

use year.

do not hibernate, although they become

significantly less active during winter

months.

spread has been

losses of native

animals. The rare European mink ( Mustela lutreola) has

now become

almost extinct, and

Britain

the water vole has disappeared from

lucrative business,

nearly

90 percent of

and

common. Mink

trade

a

was once

a

mink coat was

considered the height

managed

of luxury. However,

offshore

fur has since fallen

in safety,

out of fashion.

Here they have

are

sites

where

was formerly

to reach islands over 2 miles (3

where seabirds had

km)

previously nested

protected by the surrounding water. killed

hundreds of adult birds

and

Mmk

colonies have been wiped out

have a glossy coat that has been highly

it

chicks, as well as eating the eggs. in

Whole

the space of

few breeding seasons, and

some

prized by the fashion industry. Although wild

only a

mmk

places seabirds have been almost eliminated.

are easy to catch, trappers could not

enough

result, special

especially in

skins to

meet demand. As

When mink

a

mink farms were established,

Europe Mink were

from America to European

fur

first

farms

in

Second World War. However,

postwar years

raising

seen as a lucrative animals bred well

mink

in

the

for the fur trade

was

new moneymaker. The in captivity

and could be fed

cheaply on unwanted animal waste, including bits of

get into parks with captive

cause mayhem,

killing

many

birds.

chicken from the expanding broiler fowl

where there eat.

Gamekeepers regard mink

because they take many gamebird chicks and eggs. However, recent studies suggest that

mink numbers may be recolonize

mink.

In

rivers,

falling in Britain as otters

perhaps displacing the smaller

the United States mink

seem

breeding even more valuable colored furs were

better balance with nature

Water 7:98

They also

as vermin

problem, so perhaps one day they

Vole,

they

are high densities of juicy trout to

industry Moreover, through careful selective

SEE ALSO

fly),

create havoc on chicken farms and fish farms

the

1920s, but only small numbers were kept until after the

in

ducks (often pinioned so they cannot

imported

in

good syvimmers and have

Fur Farming

obtain

54

fur farm in Estonia.

Breeding mink for the fur

piles of

always close to the water's

may sometimes have

It

ŠA

an

in

its

in

will

less of a

achieve a

Europe, too.


WEASELS AMERICAN MINK

The In

Freedom

Price of

1998 animal

British fur

was an

fur farming

is

right or

irresponsible act, widely

conservationists

European Mink

wrong,

it

condemned by

and animal welfare groups. The

mink posed

released

broke into

farms and released thousands of

Whether

mink.

rights supporters

a serious threat to

many

its

other species, and such releases were also cruel

bred

many

captivity for

in

generations: Like pet

mice or guinea pigs, they were unaccustomed to

the wild. Large numbers were easily

life in

recaptured, since they had no idea

being caught.

Many were were

roads. Others

how

also run over

to avoid

on the

annoyed landowners and gamekeepers. Some mink even found

way

their

similar in

American cousin, but has

a

white area around

appearance to

upper

lip.

its

European mink prefer to

rather than lakes

Spain, Estonia,

and

coast, but have

into people's houses,

is

live

now found

only

has disappeared from

It

about 80 percent of decline.

in

small areas of France, of the rest of

and the invading

American mink.

in

in

notably scarce

and Romania, having been driven out loss

still

beside fast-flowing water

become

Europe by habitat

made

by dogs or shot by

killed

is

recent times. The species

mink themselves, since they-had been

to the

T

he native European mink (Mustela lutreola)

its

former range and

is

Urgent efforts are being

to establish captive-breeding

colonies from which the species might

one day be restored to the

wild.

having been driven by hunger to enter through the cat flap

in

search of food.

The rare European mink (above) has a white

Fashions have changed, and fur coats are

no longer so much less profitable

in

demand. Mink farming

today than

it

patch around

its

upper

lip.

is

once was, and

animal welfare legislation imposes

many

conditions on the managers

numbers of

of fur farms. Large

businesses have closed leaving fewer

mink

down,

in captivity.

However, populations of wild

American mink to prosper

in

Europe

and are

likely

contii

to remain

permanently established there.

Š An American mink returns to a

henhouse

in

Britain to

feed on hens

killed the

previous night.

Mink have

also

it

been

responsible for losses to

many

native animals.

*

f .


SVA„ C~'\\ Oc ES ,

Wolverine A

a ferce and greedy predator. Ruthless

persecution res resulted in

Soe-'rrfK rvame

?ve--ne 3 ,"Cr. sku-k .

iVcl.t-

’jes

a*£ olinc a^cxnd the

ocw

Family

Arctic

Oder

cota conditions sng

in

As a rcsAt an >ea'

We>gnt 2Z-65

fc

A-Ef

*.

;

Mate

.

ctse' a-

:

ma .vw

a-ge oowe^j paws. coat pare» cr *ace

Mabrts

a: *eas:

'

0*i*

ana

Soi'-a*. creaturc

s*vrt oa'*.

o-g

we:

rrvats

m

>ov\

covering over 30

woe>

tt

One

fctrcr

c* jC to

Fefcnary-MVct)

wW enough *ood

bom esanor pe-'oo

afte^

The summer

a:

S-'O

<m

arc orowfc wn?- annoyed

he spent out cr

w w grate to We shelter of

some: mes youmeynng jp to 50

A though

SO sses

n a day Often

open tund'a or mountainsides Out come the

of

*orc$ts

~

myths may

wnter wohennes vVeanec

v

we'e s only one wolverine n nore wan 200

w oe\ mams

- bates

oe avec replantation

Voter

<e the wolvenne

to *eec sometimes

es

so^arc mnes 500 so. Breeding

sno\%

afsc playful

f

werc

too

ts

much

of it the s*xxt-

chasing ang

dtfftcutty

caotunng prey As a result winter

Wet

m we

spent »'»ne'e

Mountainous

‘o'ests

ts

rcatne shelter of con

normally

*er forests

of,

ocx. arcas ana Tunara

a-srexte:: across

nerwem

Eurcoe

Although wolverines normally prey on sma crcaturcs such as Arctic narcs and

lemmmgs,

Wev must mave We most o4 any food

manage

to

*

no

n the

ripe Pe~*es to eat

and many

young mammals and During leaner

:

mes

they

there are plenty of

fall

experienced

birds that arc easy to k

the

wo verne wl

sometimes attacx animals as arge as oee* and

w»d

sheep, storing surplus food for ate' (The

woher ne$ powerful aws mean

down

animals

much

*

te-

There arc

can drag

larger

than

to t

more than

wohen ne s tendency has gnen nse tc

it

l

ts reputation as

many

foHc tales

tsetf.)

The it

needs

a wanton

about the

cunrung and boodthirsbness of wolverines Rjr

56

SEE ALSO

2:70. Bear

2 --

.

'c

'»

a<-.

7:9C

'-arc

-

C

8 S2

.*

o

rc s

r>e-~re’ o*

fmmffi iv

I

«5 ;

me

Of

e-ges:

me and

It is territorial

oerc-c ts

OeSf

s-a

33Ce3'r '‘aE

aoo/'ist

we same

the sncn% s usualy ess deeo

Waste Not Want Not Ostnbut»on

'ic: w *

m les

they can cope with oeec

eggeo wofvennes nave

Habitat

Arctx: a

number. Wofvennes ~eed

~we* must ,\5ooe'

ana bushy

we

rue cm

—pawicua-ty D'esators

tend tc be

3->a

we

mean ws:

nters

.v

brow*' bur

'.axs, tai thict

war -cams

egs

eases of

Amenca and E„-ope The

North

hjge arcas to provoe them

Ke> features

rare.

g~cw stow>> and orey animals s re scarce

plants

-

becoming quite

it

oe-S'

3 c

3*

ouh

woh enne has a

nararalh' scarce animal the

re pi tatior as

Com'non nanie

Gulo

sex

home

orvn

of


WEASELS AOLVER

trappers

comp

a n that t h e an mafs steal :

from

their traps,

and e naeer henfs^en fear that

wolverines

w

r

attack the

r

Wolverines are senstive creatures and eas y disturbed. The remote areas

animals.

Wolverines mate n the

summe and would r

normally give birth after a few weeks However, they avoid producing young process result,

just

m

known

in

midw

nter by a

as delayed implantation.

time for the long days and

abundant food supp es of the

As

a

about Apr

the babies are not born unt

relatively

Arctic

summer.

The young continue to use the mothers de~ until

May. They stay within her normal

range

until

home

about August. After that they tend

once

accessible.

are shot

sight,

not just

species

seems

likely

Wolverines were at one time found

it

to rvn

snow without

on

sinking

in.

V

much

farther south in Europe than they are today,

even into Germany.

the United States they

In

used to occur across the whole continent as far south as Arizona and

New

Mexico. Today they

are extinct east of Montana, with a

British

5,000 animals.

widespread

still

Columbia

Scandinavia wolverines have

freeze together

now

and

Siberia,

Canada, about

known about numbers

also for their fur. Their long hairs tend not to

Ruspia

in

a stronghold with

s

Little is

few

and Idaho.

in California

in

t^e intense cold of tne Arctic,

feet allow

to suffer further decline.

because of the damage they might cause, but

in

The wolverine's broad

woivennes are regarded as threate°ed, and the

and on

which they

The remaining populations of

Wolverines are

The Human Threat

in

becoming more

lived in safety are

remaining

to disperse, especially the males.

Many wolverines

ME

but

in

northern

become

rare

and

occur mainly n the remote mountains of

so the pelts are prized by local people for

Norway and Sweden.

trimming the hoods of winter coats.

are

left

n the

It

whole of

s

thought tnat only 40

Finland.

57


SMALL CARNIVORES

European Otter European

popular creatures. However, few

otters are

people have ever seen one or are Common

Mustelidae

Order

Carnivora

Size

Length head/body 24-35 tail

The European otter

length: 14-18.5 in

m

shoulder about 6

20%

so, since

in

family

(60-90 cm);

(35-47 cm); height

at

(15 cm). Male about

and has

a member of the weasel

s

body with

a typical long, thin

short legs. However, the similarities

much

Otters are large and spend

end

there.

of their time

bigger than female in

Weight 13-37

[

do

urbanization and pollution.

Family

•HI

likely to

they are becoming increasingly rare as a result of

Scientific

1

Lutra lutra

water. They are probably

more abundant

(6-17 kg

lb

than people think, but they are very elusive

Key features _ong,

tapenng

long,

wth

creatures. Finding signs of otters

all

4

feet are

on

webbed

Usua

ly

bom

in

mud, but the most

soft

important otter signs are spraints (droppings).

may

during the day

2-3 cubs

easier than

seeing the real thing. Their web-footed tracks are distinctive

land, usually nocturnal, but

come out

is

dark-brown fur

L ,es alone; swims and dives well but can be act ve

Breeding

light- or

tail;

and

legs

broad, flattened head, small ears, and

small eyes,

Habits

body with short

slender

Evidence of Otters

after gestation period

Otter droppings are smeared onto rocks and

of 2 months; births at any time of year in

western Europe, more seasonal

and

Weaned

east.

mature

at

captivity,

at

about 3-4

Voice

Occasional

Diet

Slow-swimming

shrill

the north

in

live

up to 15 years

in

the wild

Edges of

rivers

rocky coasts

in

fish, especially eels,

and

but also

in

oil.

50 years

unpleasant to humans.

hairs,

Declining

in

extinct in parts of

most

Europe

areas; in

the

meaning that

Otters are riparian species, river

In

few

a

places, Ireland

and

western Scotland, for example, they may

live

along the coast, feeding on rockpool

and

A

the day. different

live,

in

often

may

among

from grass and twigs, deep

An

fish

need a secure

which to shelter during

single animal

lairs,

otters

use more than 20

tree roots or

ground Sometimes they make a

otter can travel over

in

soft

above

couch

a thicket.

6 miles (10 km)

in

a night

and sometimes uses 50 miles (80 km) of

river in

a year Coastal otters usually have

much

smaller territories, only a mile or so long That

C tter.

N

which are essential

for keeping the animal

banks, lake edges, and other

place close to water

SEE ALSO

otter

underfur and long guard

crabs Wherever they

58

© A European

bits

droppings of most mustelids, they do not smell

waterside habitats.

last

bones, scales, and

possesses short, dense

Population widespread but scarce, IUCN I.

fish

of crayfish or crabs, for example. Unlike the

they frequent

become

patch of thick

Closer inspection reveals characteristic

of North Africa

has

like a

south

Lanka, Malaysia, Sumatra, and parts

Vulnerable. CITES

only about 2 to 3

itself,

areas

Europe east to China and Japan, also

Status

proportion to the otter

fragments of prey

Distribution From Britain throughout most of western

India, Sri

fresher than others. They are small in

look tarry and black, almost

lakes, often in reed beds;

some

some

small groups,

in

inches (6 to 8 cm) long and rather slender. They

whistle, otherwise silent

mussels, shrimps, crabs, and lobsters

Habitat

logs as scent markers, often

with

4 months; sexually

May

year

1

in

encan River 1:64,

Otter, Giant River 1:68

is

well insulated in water.


WEASELS EUROPEAN OTi

perhaps Decause *ood r

f

^ore abundsr:,

is

since

oc*'Doo *3" are easier to catch than :he b'gger sh

found

n rivers. _arge r -ers are noi

^ecessar y best for otters and they ofter nunt in

streams and

here and

Otter Conservation

ft is

tr

outaries There

is

more food

r

eas;e to catch fish n pools and

O

nce a population of a scarce species becomes small and thinly dispersed,

it is

highly vulnerable to extinction

population by releasing captive-bred animals

may

and may be

Boosting the

help save a species

sha low streams than n arge bod es of water

That has been attempted

Tne

Beginning n 1983, captive-bred otters were released into

r

so

i

otter's ears are qurte

very acute. Sight

ofsmei sigrt

but rearing

aiso good,

ar d

ts

the sense

mam

senses used

land otters can run at speeds equivalent

to a fast

waikmg

swtm

pace, but in the water they

faster than

adapted for feet *or

.

kewise very important; Underwater,

ts

and touch are the

On

car.

ts

smai

life *n

humans. Otters are wel

water. They have

swimming, and the long

tai

webbed forms a

rucder behind The the body twists and turns to allow swift

changes of direction and detatled

expl orator of

underwate r nooks and crannies.

systems to bred,

ft rs

known

population

been

gaps

fill

m

in

in Britain

river

the wild otter population. The animals soon

that at least 25 otters

were added to the

local

eastern England The use of captive-bred animals has

criticized

because they have been produced by a limited number

of animals and might be inbred. However,

—even

inbreeding

tried elsewhere.

in

all

rare animals nsk

the wild. Restocking with unrelated animals

may

actually he'p reduce the problem. Further support can be given to

otters by providing suitable lying-up sites in the

thickets or

mounds

of earth

and

form of planted

logs at the water's edge.


SMALL CARNIVORES

The

fur

very thick, allowing the otter to

is

warm and

remain

dry even

water However, as pressure squeezes characteristic

surface.

The

trail

it

air

freezing-cold

in

dives, increasing

out of the

water

leaving a

fur,

of bubbles rising to the

insulating effect of the fur

is

therefore reduced, causing loss of body heat.

That

is

dive

in

is

less

one reason why

shallow water where the water pressure

The

making otters

otters normally only

it

the fur also creates buoyancy,

air in

hard work to stay underwater, so the

soon get out of breath. Dives are

therefore short, and otters rarely stay

submerged

for

more than about 30

40

to

seconds. They also spend less than 20 to 30

minutes get

the water before coming ashore to

in

warm They

shake their fur

dry,

sometimes

running around with an arched body or in

rolling

the grass.

Versatile Hunters Otters are predators of prey caught

in

and feed on

a

wide

the water, including

variety

fish

weighing several pounds, frogs, and crustaceans However, they also catch land

animals such as rabbits and the occasional

They are very

flexible in their

bird.

hunting behavior,

being able to adjust their technique to cope with whatever opportunities arise They

will

nuzzle submerged stones to catch crayfish hiding underneath, grope feel for fish

among

sleeping bird on

its

in

muddy water

to

tree roots, or grab a

Most

nest

of their diet

consists of fish seized in the teeth. Shallow

pools are best, because the

fish

cannot

easily

get away, and the otter can breathe every

seconds

Many

dives are unsuccessful, but

few

young animals. Adults cannot spare much

in

energy for play

effort

and time are better

shallow rockpools the animals might find food

spent searching for food. Adults do not often

every once

meet

in

three dives. Otters need the

equivalent of about three

1

-pound (0.5-kg)

fish

per day to satisfy their hunger. Otters normally

alone or

live

and

also

make

in

a family

a sharp bark, but otherwise they

are not particularly vocal animals. Captive otters play a

lot,

behavior

but play seems to be relatively rare

in

the wild and usually involves only

SEE ALSO

f

in

the wild; and

be a serious

when

fight, especially

they do, there

between

may

rival

males. Fights involve squealing and chasing and

group They occasionally emit a loud whistle

60

—the

(T

iropean 8:70

sometimes

biting, until

up and runs away they

and

may in

indulge

If

in

a

one

of the animals gives

male and female meet,

vigorous romping on land

the water. Mating takes up to half an

hour and happens either on land or while

swimming. Afterward, the two animals separate

The European

lithe,

otter's

elongated body and

webbed

feet are built for

vigorous swimming. The

long

tail

forms an

effective rudder

and

allows for deft

maneuvers when searching for food.


WEASELS EUROPEAN

make

Otter cubs suckle frequently and

grow

chirruping noises. They

weeks

or

weight

—the

more

size at

open. Cubs take

weeks

(0.9 kg)

in

which the eyes begin to food from about seven

solid

of age, but

little

slowly, taking five

pounds

to reach 2

OTTER

it

will

be another three

weeks before they venture outside the den months

Otters are three

play.

first start

to swim,

to

old before they

and the mother may

be

still

catching over three-quarters of the food for

cubs nearly eight months leave

home when

old.

territory of their

than

five years,

finally

they are about a year old.

New-found independence and many young

The cubs

a very tricky time,

is

otters die as they search for a

own. Few

otters survive

and probably only one

more 100

in

reaches 10 years of age.

Chemical Poisoning The main threats to European otters are food shortages and accidents, especially getting run over on the roads.

Some

and drown. Others

fish traps

caught

otters get fall

in

victim to

insecticides used to protect crops. Toxins build

up

in

the animals' bodies,

leaving

them

killing

them

or

infertile.

Across Europe the effects of industrial pollution

and acid

the food

rain are affecting

supplies of fish on which the otters depend.

Otters are

becoming scarce or even

wide areas as

a result.

However,

extinct over

strict

controls have allowed otters to survive

of western Europe from which they

pollution in

may

parts

slowly

recolonize suitable habitat once the threats

have been removed. Repopulation

and go about

their solitary lives.

not help raise the family

needed by the cubs

is

—

all

The male does

the support

water, the long guard

provided by their mother.

Otters can breed at any time of the year

where the climate

is

not too severe and food

is

always available. Although females can have a litter

every year, nearly half of

only raise a family lasts just

born rising

in

in

them probably

alternate years. Gestation

over two months, and the cubs are

a special den, well

above the

flood waters. There are usually

three cubs, but there

may be up

to

level of

two

five.

Fresh out of the

or

hairs in

an

tend to

stick

otter's pelt

together

creating a spiky

appearance.

happening

more

in Britain,

which

now

is

already

probably has

otters with a healthier future than almost

any other country

in

western Europe.


CARNIVORES

Disappearing Otters #

T fa

r

he

survey of otter distribution

first

was

in Britain

carried out in

the mid-1950s. The animals were found to be widespread and

ly

common,

despite decades of hunting for sport. Otters

were

even reported on the fringes of London. However, by the 1960s

some

were reporting reduced

surveys

fewer than 6 percent of

sites

and

had signs of

central England

legal protection

Meanwh

still

otters. Otters

However,

egislation alone

is

is

it

full

fall.

now

extinct, proving that protective

not the answer. Research has identified causes of

decline that are potentially

more dangerous

to otters than hunting.

demise coincided with the spread of the introduced

American mink and a massive increase Habitat destruction

is

riversides, since tree roots

backwaters

in

and boating.

fishing

another threat, especially the clearance of provide otters with both

which to catch

in

was the

the otter

were given

the Dutch otter population increased tenfold following

e,

otter's

1970s

inspected over large parts of southern

from 1978, but numbers continued to

egal protection

The

success. By the late

fish.

introduction of

and sheltered

lairs

However, the main problem for

new

agricultural pesticides in the

1960s Tiny amounts washed into ditches and streams, and were taken up by plankton The plankton was eaten by turn

consumed by

fish,

which were

in

otters.

Long-lived species

are especially vulnerable, since

like otters

adults steadily accumulate the toxic chemicals, but stay alive, giving

no

sign that anything

long before poisoning

population collapses

Many dangerous

sources

New

If

the older animals die, since there are too

going.

it

danger has emerged

—

fish

this

especially eels, thereby

turn results

in

amount

at

minute concentrations. Heavy metal

and cadmium)

fewer otters

It

62

SEE ALSO

M

'

In

of predators like

more problems because

and other food

may account

for fish,

for the

which

in

disappearance of

Europe

European otters are scarce or declining

and pollution

also contaminate

damaging the health

of insects

otters in parts of Central

destruction

time from industrial

through pollution from factories and waste

and are damaging

reduces the

now been phased

have

agricultural chemicals

the otter Pollution from acid rain creates yet it

reduced

the adults cannot breed, the

pollutants (notably lead, mercury, fish,

is

chemicals called PCBs (polychlorinated biphenyls) are

contaminating effluent,

fatal.

is

when

few offspring to keep

out, but another

wrong. However, breeding success

is

some

m

most

areas, mainly as a result of habitat

parts of Europe the species

American 1:52

is

now

extinct.


VI

V

ItV

4

*V?I

Ry vV«u A K j

k

1 a*

yi

f

A

*^.'7;


SMALL CARNIVORES

North American River Otter

Lontra canadensis

North American river otters are playful, intelligent animals that were once common throughout most the United States

Common name name

Lontra canadensis

Family

Mustelidae

North American

Order

Carnivora

all

Size

Length head/body: 26-42

12-20

length:

in

in

(66-107 cm);

(32-46 cm); height

and Canada

until at least

at

were the most widely

1

cylindrical

legs, long,

pointed

and

small ears

body and tail;

short, stocky

small, blunt

with plenty of

brown

do not Lively

and

playful: lives

alone or

in

November-May

sexually

21 years

Voice

Diet

mature

Shrill chirps, soft

14

in

May

and scrub cover and

polluting water with pesticides. Because otters

live

to

are at the top of the food chain, they are

the wild

sensitive to

"chuckles," grunts, coughs,

any pollutants. Another threat to

Mostly

30,000 animals are harvested every

also crayfish, frogs, crabs, birds'

mammals

Coastal and freshwater:

large

rivers,

is

Distribution .anada and mainly northwestern,

U

areas of

but extinct or rare

in

U

S.

actually

year.

Such

be sustainable as

is

healthy. Sadly, this

frequently not the case, and pollution,

and

loss of habitat

about the extinction of otters

S.

Population probably low thousands; CITES

the fur trade. About 20,000 to

numbers may

disturbance,

southeastern, and Great Lakes states of

is

long as the otters' habitat

streams, lakes,

marshes, swamps, and estuaries

Common m some

cities

a threat to otter

otter survival

reservoirs,

Status

away from

and growls; loud screams when frightened fish;

are

industry, or recreation,

habitats by removing tree

5-6 months;

at 2 or 3 years.

in captivity,

eggs, and small

Habitat

at

any

them

When waterways

disturbance.

and towns agriculture poses

after gestation period of

Weaned

in fact,

Yet they are shy animals and

the otters usually leave. Even

approximately 50 days (including delayed implantation).

like

fish.

developed for housing,

1-6 (usually 2 or 3) cubs born from

Litters of

—

small

groups, semiaquatic; active at any time of day

Breeding

and streams

healthy water systems that can provide

head with

eyes; fur light to dark

mammals

distributed wild

the United States. They favor coastal areas,

estuaries, rivers, lakes,

Habits

the 18th century.

Along with the beaver and timber wolf, otters

in

Key features .ong,

in

the major waterways of the United States

than female

Weight

used to be found

river otters

tail

shoulder: 10-12 in (25-30 cm). Male larger

t

in

have brought

many

areas.

II

and Canada,

Favorite

Games

others

Otters are intelligent/ quick, curious,

and

exuberant creatures. They are famous for their playful nature

~r

activity.

games

and

make

will

a

game

The young and even adults of tag, tumbling,

favorite pastime

is

of any

will

belly

play

and wrestling A

sliding

down mud

or

banks into the water, tobogganing on

snow

their

with their front legs folded back. Small

objects such as shells, stones, or sticks are used as toys

64

rivers are

too polluted or urbanized to support them.

(northern liver otter) Scientific

and Canada. Many

North Ar.erÂŤr*r.^;,er otter

SEE ALSO

Ott*

opean

1

:

58 Otter, Sea ,

1

2,

and

for

games

of hide

Beaver, American 7:30

and

seek. In

of

now


WEASELS NORTH AMERICAN

captivity otters

have been shown to have good

memories and can be trained

to perform tricks

their aquatic

Š North American

river

body and

river otters are well

life.

They have

feet with

webbed

a long, streamlined toes.

The

fur

with long, glossy guard

is

dense and

widespread as they once

Long, sensitive whiskers help the otter locate

were and are

now

mostly

prey

in

murky water, and the

hairs.

small eyes are set

confined to Canada and

high on the head and close to the nose, so the

the northwestern,

otter can look

southeastern,

and Great

Lakes states of the

United States.

around above water while

keeping most of

its

swim by

in

side.

m)

They have

a top

6 to 7 miles per hour

and

tail

speed of (1

Waterway

Travelers

Otters can have very large

may

use

many

home

ranges and

miles of waterway. Males

generally travel farther than females. The length

home

of a (8 to

range can vary from 5 to 50 miles

80 km), with the

An

size

depending on

otter living

in

an area with

plenty of food, for example, will not have to

the water. They

twisting their hind quarters

from side to

feet (18

deep. They can stay submerged for over four

habitat quality.

body submerged.

Otters are very graceful

60

adapted to

otters are not as

oily,

at least

minutes, but most dives are shallow and brief

such as retrieving objects.

American

km/h) and can dive to

RIVER OTTER

0 to 11

travel as far as

one that

lives

where prey

scarce. Territories of individual animals

overlap,

and the home range of

crosses that of

a

is

may

male usually

one or more females.

Within their

territories

have many resting

American

sites

river otters

and dens. Rather

digging their

own

dens,

they usually use holes


female together, or even a group of bachelor males. Groups are temporary, have no apparent

and do not cooperate

leader,

share

what

is

hunting or

caught.

Most American

when

in

they are

two

river otters start to

breed

years old. Receptive females

advertise their condition by markings at scent stations,

two

and the powerful smell may

attract

or three males. Although American river

do not form strong

otters

once

pair bonds,

a

male has mated, he may drive away other

who come

males

receptive.

part

made by

other animals, such as beavers. They

otherwise use natural shelters such as hollow

even abandoned

Tj

A

will

rearing the offspring. However,

the family group

a stream in

when

months

boathouses. Nursing dens have an underwater

born blind and helpless,

November and May, but

entrance with a tunnel leading to a nesting

but

chamber

and

lined with leaves, grasses, mosses,

bed

hair to provide a soft

for the

young.

fully furred.

They are

April.

usually in

The pups are born

fully furred,

about seven weeks, bjt

the adults.

six

The young are born between

old.

introduced to water at

rejoin

the young are about

Montana. The young are

tree trunks, driftwood piles, or

is

then leave her and takes no

sometimes fathers have been known to

river otter with

young by

in

He

near the female while she

and look

blind

like

and

March or but

helpless,

miniature versions of

are often reluctant

Scent Marking Otters

other

mark

swimmers and have

their territory using scent. Like

members

dragged

of the weasel family, an otter

has scent glands near

its

anus that produce

a

be

Reluctant

Swimmers

Otter milk

very rich,

quickly

on

is

it.

and the young grow

They open

weeks and begin

their eyes at

their mother.

onto scent posts throughout

to water by the time they are seven

tree stumps,

usually well

They

prominent stones, or

above the water

marks do not get washed levels after rain.

its territory.

off

line

by

logs,

so that scent

water

rising

As well as depositing musk,

otters leave spramts (feces),

which provide

messages

for the inquisitive noses of otters

visit later

Scent posts are usually

places

on the

who

obvious

in

otter's route, including dens,

rolling places, slides,

runways, and haul-outs.

Otters will also scratch up

mounds

of

soil

and

debris or twist tufts of grass together, marking

them with scent deposits

or spramt.

As well as

defining boundaries, scent marking also signals

when

the otter

probably

its

last

passed through,

its

sex,

and

age, helping avoid potential conflict

with other otters

Although generally

spend time

made up

in

solitary, otters will

a small group,

which can be

of a mother and her pups, a male and

SEE ALSO Hare

owshoe 8:74

about four

playing with each other and

strong-smelling musk, which the animal wipes

may be

66

there.

to

The mother introduces the pups

weeks

old:

They are often reluctant swimmers and may

need to be dragged

into the water.

A mother


wil 5 pend a lot of

catching sma!

and releasing them agair so

fish

r

the young otters can develop the s*

ore

B>

ills

or

reared unt the>

w -'o

r

morths but somet mes memoe's together tor a year or

amounts

lots of fist

,

:

j r ne

of a

s

>

’a” \

along wrtr s~,a

frequently

1

rg of 'food

ir

iVher hunting

time diving and chasing stories at

a single

the<,

or

fisr

a

lot o'

oggmg m mud

the botto" of ponds and streams

r

r

seasno e

n the mouth, not the

Otters catch prey

r

5mal 'oop rtems a e eater

-

the water

paws vh e

>arge r ories are tare’ asnore

Otters tend to catch ,arge or

slow-moving *'eas*

fsh concentrated 'to sma

exhausted "spawned-ouT salmon ere

easy to cater

depend

Fo most r

of the year

or other types of fs'

no sever

Acoough

fast-swimming species suer as trout anc pire

commor m

fish

.

but are more

be

visible

cr

me

e> r activity

to dry

*

r

^

so4*

me

webs oe* weer

r

sorrietimes on twisted turns of

n anc vegetation

rvers and .ahes, otters usually

some soo

r

t

re\ to eat other species

that actually corioete with soon fish for food.

w

Otters

sometimes eat water? dw such

and ducks, and rad the

as coots

eggs. “'ey

ma^ take oead o

particularly

ir

active /

streams or shallow pools with nowhere to

a e

piies of c

aiso

'iame giver by angers Driers eat

1

During salmo r spavv r ng times m~e rs

or the hordes of

r

ge a r eas of battened vegeta: on

r

go fo r ease r prey They do not oese've the pad

mportant pan

of the diet, as a e craos along the

otters

o

r

or the bar

rol

show the ost octve marks

So rants may r

,a

the water, tney

t

mea They

spend

for smaller prey Crayfish are ar

escaoe

toes

o*

relatvey easy to find dues to

ter

have a rapid metabolism so they need to eat

fi$h

their footprints

is

it

of othe r pre> The> corisume up to

2 pounds

and

come out

themseves, leaving

g'ass

-..up

and elusve, few peope a r e ucky enough to see

>

them However,

/y/her they

!y

months so

a

O

tters are sr

ns: oe fu

are a: east three

They a re not fu u y odspendem

Oners eal

hunting

ID weeks of age toev star to

eat soup ’oop, although they

wil stay

Signs of Otter Activity

-

time lead ng her pups,

me snoot ng

unt and k

h

r

'

r

nests

ured Di'ct

seasor

heathy

*'or

but

w

An o ~ er ’eeos on

also

birds They stalk

trout caught

me" by swim m ng underwater and grabbing me birds from below Ome s are aiso among the

will

~ew o'eoators m.at

noting

r

Occasional

1

/,

mammals on American sm.a nares.

oners wi

river

Trey wi

and

mar

turtles

piros a nd

land There have been

c~e

mammals up

rosenips

snapping

k

r

s

r

epons

chasing and catch ng

to the size of

a:so ea: berries

snowshoe

sum

blueberries, bu* that

is

as

unusua

G e rie ral‘y,

m

river

a

a rver.

omers

take la'oe o r slowfish rather

than

fast-swimming species, of

such as trout. During

salmon spawning times otters will feast

exhausted

on

fish resting in

shallow pools


SMALL CARNIVORES

Giant River Otter

Pteronura brasiliensis

The giant river otters of South America have been severely reduced

Common name

numbers as a

in

result

of

hunting for their fur They also

Giant river otter

suffer from habitat Scientific

name

fortunately, small populations

Family

Mustelidae

Order

Carnivora

Size

survive in protected areas.

Length head/body: 34-55

13-39

length:

tail

shoulder: 16

in

(86-140 cm);

in

(33-100 cm); height

in

at

(40 cm)

Giant

48-57

Key features Large

lb

(26-34

otter with short, glossy

when

creamy nose and

and

is

female

kg);

(22-26 kg)

that looks black

tapers

lb

brown

webbed

many

Groups

20

as

individuals living together.

usually consist of a

feet; tail

flattened with a flange along

otters are the

most sociable of

otter species.

They stay close together, often

all

the world's

each other or indulging

in play.

water

The adult male and female often share the

Up

each

same den,

to 5

Weaned 2 years.

young born

at

in

a single

litter

65-70

days.

3-4 months; sexually mature

May

live

over 14 years

a habit that

is

not seen

Giant otters are also believed to help each other

by driving shoals of

where they can be caught more

Mainly

fish,

and

whistles; very vocal

but also freshwater crabs and

mammals

Slow-moving

and swamps,

fish into

on slow-moving

usually feed catfish,

rivers, creeks,

the more

at

in captivity,

Loud

yelps, barks,

in

familiar river otters of the Northern Hemisphere.

possibly similar in the wild

occasional

Habitat

along

Lives in family groups, mostly in the

year after gestation period of

Diet

pair,

animals from the previous year as well. Giant

calling loudly to

Voice

mated

with their young born that year and often a few

each edge

Breeding

family groups of six to

fur

wet; often white or

throat;

river otters live in

eight animals, although sometimes there can be as

Weight Male 57-75

Habits

damage, but

Pteronura brasilien sis

shallow water easily.

They

species, such as

which are easy to catch. Some such

fish

can be over 2 feet (60 cm) long.

especially within forested areas

Distribution Once over

much

of tropical South America

south to Argentina;

Status

now

rare

and patchy

Population perhaps fewer than 2,000 the wild, IUCN Endangered; CITES I

left in

Coveted Fur The giant

river otter

is

the largest of the

freshwater otter species, and has a distinctive flattened fur

is

tail.

As with other types of

when

giant otter's fur

is

in

the water. However, the

very distinctive, being short

glossy, like velvet.

sought after

was

in

It

became

the 1960s

particularly

when wearing

selling a single giant otter skin

than they would get

in their

working hard as laborers

for

pay packets for

two

or three

months. The same skin would be worth

SEE ALSO

furs

very fashionable. Native hunters could earn

more money by

68

the

dense and helps protect the animal from

getting cold

and

otter,

ropean 1:58, Otter, North American River 1:64, Otter, Sea 1:72

five


WEASELS GIANT

RIVER OTTER

Giant otters are mainly active during the

daytime and so expose themselves to many

humans with

dangers, particularly from

spears

or guns. Giant otters are also very vocal,

frequently calling to each other using a wide

range of sgueals, barks, and whistles. Their noise draws attention to

them and makes and

easier for hunters to locate

whole family

kill

groups. Another fatal characteristic otters are very curious animals:

around with

their

their

that giant

They often swim

surroundings

to shoot. They will

makes them easy

swim toward

often

is

heads held high out of the

water to keep an eye on habit that

it

and

intruders

potentially

dangerous situations to investigate more closely

— again making

and to

it

easier to shoot

them

others of the group that stay around

kill

what has happened.

to see

A Host

of Threats

As huge areas of the South American forests

were opened up

made

tracks

it

for logging,

much

new

roads and

easier to reach places that

had otherwise been remote

retreats for the

giant otter. Gold mining has been a problem, too: Large

amounts

of cyanide

and other

poisonous pollutants have been released into the

rivers

where the

otters

Dredging for

live.

gold also makes rivers very muddy, which severely reduces the fish populations

the otters. People still

more is

It

the

fish

rivers,

needed by

too, removing

of the otter's food supply.

little

wonder

that giant otter

populations collapsed, and soon the species

seemed Trade

to be heading for early extinction.

was banned

in

1970, but there was

considerable black market

activity,

and skins

were smuggled through countries where law

©

times as

The giant river otter

of South America

is

much

in

the fur trade after processing.

Financial rewards provided a major incentive for

the

and

the 1950s over a

enforcement was

slack.

Nowadays

are not fashionable, so the market richly

rewards hunters for

killing

most sociable of all the

hunters to

otter species—

thousand giant otter skins were exported each

Fortunately, giant otters are

characteristic that has

year from Peru alone. Since otters reproduce

in

sadly helped decimate

slowly, with only a single litter

populations. is

Nowadays

a rare sight.

it

kill

otters,

in

each

year,

no longer

the animals. fairly

numerous

the Pantanal (bordering Brazil and Paraguay)

and

in

parts of Peru.

Some

breeding rates were not enough to keep up

are relatively secure

in

with such high levels of

the giant

enjoys

loss.

still

otter skins

river otter

of the best habitats

national parks, full

and today

legal protection.

69


Short-Clawed Otter -

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-


WEASELS SHORT-CLAWED OTTER

Fun and Games Young short-clawed

to help

otters are social

and

playful

creatures.

They tumble, play chase and tug-of-

war, slide

down muddy

boisterous.

When

and are generally

hills,

otters are not entertaining

themselves, they bask on rocks, hunt for food, Short-clawed otters are very sociable in

and

live

loose family groups

numbering up 12.

to

or

go

otters, they

many other

spend more time on land than

in

Short-clawed otters tend not to dig their

own

dens, since they lack strong claws. Instead,

make use

of the

playing games, such as

they usually

chase and tug-of-war.

of other animals or shelter In

areas

where there

in

is little

The

driving shoals of fish

otters are put

keep any

to

Short-clawed otters often feed rice paddies.

the

are largely nocturnal.

flooded

farmers because they sometimes uproot young rice plants

while probing

the

in

mud

for food.

However, they eat huge numbers of crabs, a pest

in

nee

fields,

so

it is

likely

they

do more good than harm. Short-clawed otters are

officially

otter species. However,

In

in

They can become unpopular with

dense vegetation.

disturbance, short-

on a harness

they catch.

fish

at a lower risk of extinction than

paddies of Malaysia, where there are often

many people around, they

and allowed

work by

abandoned dens

clawed otters are active during the day. rice

at

into their nets.

which are

water, preferring shallows to deep water.

about

The young enjoy

for a leisurely paddle. Unlike

them

range, such as

deforestation

opens up

in

is

in

some

classed as

some other parts of their

parts of Malaysia,

a serious

river valleys for

problem because

it

development.

Development means disturbance and often

Short-clawed otters are sociable and easily

food. Increasing use of pesticides

trained. In Southeast Asia fishermen tram otters

pollutants also

since

many

harms the

less

and other

otters, particularly

of their prey species are very

sensitive to pollution.

entanglement

in

Other threats include

fishing nets

and

fur trapping.

Asian short-clawed otters are part of a

jyyvorldwide captive-breeding program that aims to protect the species from extinction.

program

is

The

focusing on the best ways

of keeping and breeding

so no

them

more need be

taken from the wild.


SMALL CARNIVORES

Sea Otter

En hydro

lutris

*

The sea otter was once widespread along the coasts of the North Pacific, but hunting for skins brought the

species to the brink of extinction.

Common name Scientific

name

Enhydra

protection

lutris

Order

Carnivora

Size

Length head/body: 29.5-35

and some successful

Today sea otters are

length:

shoulder:

1—12.5

1

8-10

in

in

(75-90 cm);

(28-32 cm); height

at

lb

(14-38 kg)

make them

antics

on

otter with blunt-looking

that turns pale

a popular species to watch.

head

in

their fur

One pup born each

year

after gestation period of

mature

May

at 3 years;

live for

in

early

summer

4 months (including

in

the waves. their

a strand of kelp

their chest. Periodically, they feed

by making short dives of about a minute to the

seabed to look

rabs, sea urchins,

for

and

mollusks. They cannot dive deeply, so they have to stay

in relatively

cannot

last

shallow waters. They also

males at 5-6 years, but do until at least

over 20 years

7 years.

in captivity, similar

the wild

make

long without food and are unable to

long journeys out to sea

crossing large areas of

Voice

Normally

Diet

Crabs, shellfish, sea urchins,

silent

fish,

and other

marine animals

Habitat

—often anchored by

months; females sexually

not breed successfully

m

over and over

rolling

draped across

up to 8 months delayed implantation). at 5

and

They also spend long periods dozing on

and calm waters;

kelp beds

dives to feed from seabed

Weaned

back at the water's surface, grooming

their

backs on back

their time floating quietly

cream with age; feet

completely webbed; hind feet form flippers Floats

observed, especially

along the California coast, and their playful

They spend most of

Key features Dark-brown

easily

reintroductions.

(20-25 cm)

in

Weight 30-85

Breeding

now

substantially recovered, thanks to strict international

Mustelidae

Habits

has

Sea otter

Family

tail

It

Kelp beds and rocky seashores

if it

entails

deep water. However,

northern Pacific from California to Kamchatka

The sea otter

they do sometimes undertake long journeys

possesses the densest

along the coast, sticking close to shore where

coat of any

they can continue to feed along the way.

help keep

and

Generally, sea otters are solitary animals,

Distribution Formerly along coasts across eastern and

(£)

males are

territorial,

competing

probably to avoid

for limited

chilly

warm

in

to

the

seas of the North

Pacific.

food resources.

it

mammal

There are

estimated to be over half

and northern Japan, exterminated over most of

its

range,

now

California, Alaska,

Status

a million hairs per square

reintroduced to coasts of

Oregon, and Washington

Unwitting Conservationists

inch of

Population about 150,000 and growing;

Sea otters are intelligent animals and have

IUCN Endangered; CITES

learned to bite open old drink cans that have

II.

Given

protection in 1911 and probably

full

now

legal

secure

sunk to the bottom of the ocean and provide a hide.

A-

lair in

which a small octopus may

They also eat large numbers of sea

urchins, helping control their

numbers down eat

now

growing

is

kelp.

numbers. Keeping

important because the urchins If

there are too

many

the kelp beds are unable to flourish and protect the coast from the Pacific tides

result

full

urchins,

do not

force of the

Beach erosion and flooding may

Hence sea

otters are very important

ecologically for maintaining a healthy coastline.

72

SEE ALSO

Ottfr. Giant River 1:68,

Whale, Killer/Orca 3:62

fur.


WEASELS

SEA OTTER

Densest Fur on Earth Sea otters are probably the smallest

blooded animals that spend water.

all

warm-

their time in the

The coastal seas of the North

Pacific are

very cold: Even far south off California the sea is

and

cool

quickly.

will chill

The sea

mammal's body

a

needs very

otter therefore

effective insulation to prevent loss of Its

protection

is

are estimated to

twice as

be more than

as

found on the

The fur has long, shiny guard keep the water

half a million

at

otter's

body

larger fur seals.

hairs that help

bay and prevent the underfur

from becoming matted and losing value.

heat.

mammal. There

square inch on the sea

many

body

provided by a thick coat of the

densest fur possessed by any

hairs per

quite

Below the guard

hairs

is

a

its

insulation

dense mass of

extremely fine underfur that traps a layer of

air

against the skin and acts as insulation to

prevent heat

loss.

The sea otter

is

fabulous fur to enable

without becoming

much

of

its

dependent on

totally it

to

chilled.

live in

It

the cold seas

therefore spends

time grooming and caring for

precious coat. That

is

also

its

why

its

the animal

cannot dive deeply, since the increased water

Š Sea to the

otters

make

dives

seabed to look

for

prey such as crabs, sea urchins,

and

mollusks.

However, they easily get out of breath, so cannot dive deeply,

and must

stay in shallower waters

along the coast.

73


SMALL CARNIVORES

pressure at depth squeezes the

causing

fur,

to lose

it

The trapped

air

buoyant, so

also

insulating properties.

makes the

into the

it

breath.

from staying

to dive repeatedly until

When

more, the hunter speared

not just valuable to the

200 years

it

was

also

one of

human

A

for winter wear.

last

was

miles of coastline. Sea otters have

plenty of incentive to hunt the otters.

one had been

for the early exploration of the

occasionally

them smash open hard

North

During the 18th century Russian

withstand heavy

Prey

is

navigators expanded their trade

placed on

the otter's stomach

Pacific.

colonized Alaska and

and

what

is

in

now

Columbia, as well as the Aleutian

crushed with the stone.

the

British

and Americans joined

skins

and

rapidly

year

British

Islands. Later

Skin Trade were

killed

between 1750 and 1850, with 17,000

skins

in

shipment made

in

were bartered with the Chinese

in

fine porcelain,

Expeditions

for

which could then be taken to

Europe and sold for immense

would stock up with axes and

made

America, then

back to the North

sail

in

natural

losses.

They do not breed

and produce only one youngster per

—often not even every

not breed

until

year.

Females do

they are over three years old

first

have a breeding

five years old

baby.

Male sea

mature and may not

territory established until they

Slow breeding meant that sea otter populations could not cope with heavy exploitation,

and the animals soon disappeared

over wide areas. By the early 20th century the

profits.

other useful tools

few

are 10 years old.

1803. Pelts

exchange

every

until

them) and are not adapted to

kill

otters take even longer to

a single

otter.

whales, bears, and bald eagles

(killer

and sometimes can be over

in.

million sea otters

and

along hundreds of

killed

before producing their

Over three-quarters of a

skinned. The

into the sea,

an entire year's wages for a seaman, so there

main reasons

shells.

was

they could be hunted systematically

carry a flat stone to help

back. Sea otters

it

did not seek safety by dispersing out at sea,

sea

predators

its

where

Since the otters only lived along the coast and

use,

Hunting sea otters and fur seals was one of the

on

to dive any

the hunter paddled on to find the next

could be sold for the equivalent of

otter's skin

of

and dragged the

it

body was then thrown back

is

into the kayak,

it

was out

it

was too exhausted

it

depths. The otter's fur

making warm coats

swimming

was forced

body

the world's most prized furs for

shellfish while

hunted from canoes

easily

and kayaks. An animal would be chased so

soon gets

It

were

otters

underwater long enough to reach greater

animal: For over

sea otter feeds on

The

otter rather

a small seal or whale.

more

trade the tools with local hunters for yet sea otter pelts.

water than would be

out of breath, which prevents

A

from the

has to expend more energy

it

swimming down needed by

its

vital air

sea otter had

Europe and North Pacific

and

become exceedingly

been reduced to perhaps fewer than thousand animals

was on the

a

the whole North

in

having

rare,

Pacific.

brink of total extinction. Yet

It

its

range crossed several international borders so

Tool User

giving

it

legal protection in

one country would

not necessarily help: The animals might be

he sea otter

is

one of the few animals, apart from apes,

learned to use tools.

armpit and uses

It

to help

it

often carries a

its

paws

in

to

crushing

it

calm water, with lift

its

the stone and

against

its

stone tucked into

smash open the hard

mollusks, and sea urchins on which floating

flat

it

feeds.

prey lying on

hit

that has

shells of the crabs,

The its

its

otter

chest.

lies

on

its

back,

The animal uses

the prey hard and repeatedly,

chest until the juicy insides are exposed and can

killed illegally in

smuggled out

country, but then

to be sold

was needed was an

international

any marine animal.

In

The 4:12; Chimpanzee 4:28

What

agreement

1911,

in

the

first

such

international agreement, the Russians, British

(on behalf of Canada)

total protection for the sea otter

throughout the North

Family,

else.

to

However, that had never been done before for

agreed to

SEE ALSO Ape

somewhere

give the animal legal protection everywhere.

Americans, and

be eaten.

one

Pacific.


Repopulation Success Slowly, sea otters

have regained

and today there are about about

half the

In

number

1

their

numbers,

50,000 o f them,

that probably existed

300 years ago. Gradually, they are spreading back to

many

parts of the coast

where they

have been extinct for more than a century.

was once thought

that sea otters had been

eradicated from the California coast, but a

were spotted increased to

in

It

1938, and numbers have

lately

more than 2,000. Fishermen now

say there are too

many

that the otters eat too

of

them

many

—complaining

mollusks, crabs,

and other valuable sea creatures. Population growth leveled out small decline

in

the late 1970s, and a

may even have taken

place since

brown

surplus animals have been transported to

pale cream on the head

Washington

State,

Oregon, and Alaska to

repopulate areas along those coasts. Overall the sea otter seems

now

expanding future

new

in

to have a secure

the North

Pacific.

dangers, notably from

are almost as dangerous.

comes from TBT found

in

(tri

butyl

tin),

adult.

Sea otters

and

for over 20 years

However,

now a

oil spills

near the coast. Detergents used to clean up spills

coloration turns

with age, as with this

fully

may

live

and are

protected

species.

oil

Another threat a

substance

the special paint used to prevent

barnacles and seaweeds growing on the hulls of boats.

marine

The substance also

life,

including

some

kills

other forms of

of the

main foods

of the ÂŁea otter. Nevertheless, the sea otter's

one

of the best examples of

1998, perhaps indicating that the habitat

comeback

cannot sustain any more, and sea otters are

successful international cooperation to secure

back at their natural population

the conservation of a rare animal.

size.

otter's dark-

recolonization of the sea otter's former haunts,

there are

few

Š The sea

an attempt to speed up the

is

75


SMALL CARNIVORES

American Badger I

Common name Scientific

name

Family

Order Size

The American badger catches most of its food by digging. It can tunnel with amazing speed and will turn over vast

Mustelidae

AMERICAN BADGERS HAVE A BODY

Carnivora

With

tail

quickly.

in

at shoulder:

move through

lb

(8-12

female 13-18

kg);

lb

(6-8 kg)

soil

webbed

surprisingly

membrane)

third

to protect their

eyes from dust. Loose skin on their back and shoulders gives

Key features

partially

There are stories of badgers digging

eyelid (a nictating

Male 18-26.5

for digging.

through asphalt roads. The animals have a

(20-25 cm)

in

bllilt

powerful claws and

in

4-6

length:

their

toes they can

Length head/body: 16.5-28

8-10

in its nightly hunts.

Taxidea taxus

(10-15 cm); height

Weight

amounts of soil

American badger

(42-72 cm);

I

,v\

them

mobility

in tight

tunnels.

body with short legs, long curved foreclaws, and shovel-like hind claws; gray to Flattened

yellowish-brown fur with cream

belly; sides

face white; dark patches behind ears

of

and on

cheeks; white stripe from forehead to nose

Habits

Breeding

Forages at night; does not hibernate; solitary, except for breeding pairs and family groups Litters of

1-5 young born

March or

late

April after gestation period of 7

(including 5.5

Weaned at

for

months delayed

early

implantation).

6 weeks; female sexually mature 12 months, male at 14 months. May live in captivity,

12-14

Dens are the cente of badger usually simple tunnels with

excavated

When

in

badgers are

in

Diet

Normally

silent,

Mainly burrowing

mammals such

reptiles, insects,

Habitat

as pocket

and occasionally some plants

to help keep

more complex. The rejoin the

meadows, and cold

and covered

extensive, nursing

US, and Mexico

still

and

rear the

side tunnels

uncommon

and chambers,

tunnel are dug as

dens have larger

Mounds

contain fur because they are dug

when

the badger

is

material.

Since they are so

in soil.

unknown, perhaps low

thousands Increasing, but

warm. The

main thoroughfare,

mam

outside than normal dens. Population,

birth

sometimes containing grassy nesting Shallow pockets off the

desert areas

Status

soil

are additional side tunnels

latrines

Distribution Parts of Canada,

up outside.

allowing the badgers to pass each other. There

squirrels; also birds,

Treeless regions, prairies,

piled

the wild

but occasional yelps

gophers and ground

is

Soil

sometimes block the

dens used by females to give family are

They are

the den, especially during

will

entrance with loose

branch off and Voice

in

cold weather, they

life.

one entrance.

making the den

months

at

26 years

Multipurpose Dens

in

piles of soil

often spring

molting.

American badgers are uniquely adapted

for

catching underground prey. They dig into

burrows to catch pocket gophers, ground squirrels,

they

food

and many smaller rodents. However,

will also is

take advantage of whatever other

available

and

will

eat

many

small

creatures, including snakes, toads, frogs, birds,

76

SEE ALSO

‘

ur °P p an 1:78,

insects

and

worms.

In

their grubs,

wasps, bees, and

addition they will occasionally eat

Badger Honey 1:82, Wolf, Gray 2:54. Coyote 2:58


WEASELS AMERICAN BADGER

plant material, too, particularly

when

in

the

fall,

they take sunflower seeds, corn, and

coyotes follow badgers while they are hunting, catching rodents that the badger flushes from

Š An American badger

other grains. They also eat carrion and are

burrows. Coyotes help the badgers find

emerges from

known

burrowing and hunting areas, sometimes

its

den.

Its

powerful shoulders and strong claws

make

it

exceptionally proficient

to store food

in

old dens.

Badgers sometimes develop close

new

appearing to encourage them with "chase

me"

and sharing the proceeds of

their

associations with coyotes, tolerating their

play behavior,

presence and even playing, with them. The

joint

hunting

efforts.

at digging.

Plowing the Land Badgers are an important part of their habitat

because they act as miniplows,

literally

the land. Their digging loosens the creates patches

where

soil

shaping

and

different types of plants

can grow, increasing the diversity of prairie species. Their holes are

often used as ready-made

dens by other mammals and as nesting sites by birds such

as burrowing owls.

Badgers have few natural enemies, since they are such ferocious fighters.

Once they

are over a year old

past their vulnerable stage,

probably the greatest threat. run over or die

in

humans

and

are

Many badgers

are

traps set for fur-bearing

animals. Others are poisoned by bait put

down

to control wolves and coyotes. Badger hair has

been used to make shaving brushes, but on the

whole the animals are not hunted

for their fur.

Tolerated Farmers generally tolerate badgers because they eat large

numbers

venomous can

of rodents

and

will also

kill

snakes. However, badger burrows

damage

crops and are sometimes

hazardous to livestock and machinery. Badgers are one of the few larger

mammals

that are actually increasing their

range

the United States. Because they

in

in

treeless habitats, they benefit

logging and other

open up the

human

land.

live

from

activities that


SMALL CARNIVORES

European Badger

Meles meles

Common name

Badgers are

European badger

large, distinctive

Family

Mustelidae

creatures that live in groups of closely related animals called

Order

Carnivora

clans.

name

Scientific

Size

Metes metes

Length head/body: 27.5-31

(70-80 cm);

tail

t

in

at shoulder:

(30 cm)

Weight 18-26

Badgers are great diggers and excavate an

lb

(8-12 kg)

Key features Dog-sized animal with coarse

on

Habits

hair,

grizzled gray

face

belly,

stripes

members of the mustelid family.

in

length: 5-7 in

(12-19 cm); height 12

is

extensive burrow system called a

and long, on back and black

labyrinth of perhaps

white with prominent black

Normally

100 yards (90 m) of

tunnels and galleries. Main setts are the

running backward through the eyes

inactive for long periods in winter, but

sett.

there are about 10 entrances leading to a

short legs

headquarters of the social group, which consists

Nocturnal, occupies a clan territory; often

Breeding

They are the most social

of closely related individuals collectively referred

does

not hibernate

to as a clan. Often they have small "outlier"

Usually 1-4 cubs born around February after gestation period of 10-12 months (including

setts

8-10 months delayed at

implantation).

Weaned

about 4 months; sexually mature at 2 May live to be 16 years in captivity, 10

the wild

visited for a

few days

weeks

or

during the late summer.

Badgers have special adaptations to help

years. in

nearby that a e not occupied continuously

and are only

with digging, notably powerful limbs and big

Voice

Occasional yelps and loud whickering noises

paws, each armed with long, curved claws on

Diet

Almost anything edible found

all

including

acorns,

Habitat

ground

at

worms, small mammals,

roots, fruit,

and beetles

soils

shoulders and forelimbs are specially modified

on slopes

for

earth. Digging

in

Status

to China

Population probably at least Generally scarce, even extinct

but increasingly

common

in

and

done with the broad forepaws,

is

burrowing

most of Europe from

and Spam eastward

is

and loose earth belly

Distribution Widespread

The bones and muscles of the

to provide leverage for shifting stones

Woodlands, farmlands, even suburban areas where there is access to food; prefers welldrained

five toes.

level,

scooped backward under the

and kicked away with the hind

feet.

Britain

and Japan

Creature Comforts 1

.5 million. in

some

Underground

it

is

areas,

parts of Britain

damp

as well. To

generally cool,

make

the sett

and

it

may be

more

comfortable, badgers import large amounts of

bedding -

in

the form of dry grass, bracken, or

'

-

other suitable material. They rake up the grass using their front paws, tuck a bundle between

the forearms and chin, and shuffle backward to the

sett.

Badgers are scrupulously clean animals.

They use a special their

.

I

toilet

area outside the sett so

underground nests do not become fouled.

Nevertheless, the bedding gets soggy after a

few months, and badgers

usually clean

it

out at

the end of winter, leaving a large heap outside.

78

51

1

ALSO

1

i

}


WEASELS EUROPEAN BADGER

The badger's

fur

coarse and sparse, but

is

protects the skin from

damage and does

get clogged with

A

long claws, and

spend

dirt:

falls

away. Badgers

when

a lot of time scratching

they go

especially to get

above ground each evening, the

not

quick scratch with the

the mess

all

it

out of their coat. However, the sparse

soil

much

wiry hairs do not provide

insulation, so

instead of having a layer of fluffy underfur close

keep them warm, badgers

to the skin to

accumulate large amounts of under the

insulating layer

an

fat as

skin. Fat

also useful

is

drawn on over

as a food reserve that can be

winter. Badgers are not true hibernators, but

they do remain inactive underground during long periods of inclement weather, relying on their fat reserves to tide

them over

until

conditions improve.

and

Social Badgers to the

Solitary groups, but they are not social

live in

same extent

as lions, for example.

do not help each other travel in herds or packs.

spends

practically

ground alone.

It

all

of

raise

young or hunt and

Each adult badger its

active periods

is

therefore

developed than that of many monkeys, but

more in

above

hunts alone, travels alone, and

feeds alone. Badger social behavior less

They

highly organized than

large groups

once

which

in seals,

live

a year only for the

purposes of breeding.

A it

is

group

special benefit of living in a

easier to

defend

territory.

is

that

Instead of each

badger defending

own

its

individual

whole

territory, a

group of related animals (the clan)

defends a communal patch of land, which

may be

over 250 acres

(100 ha)

Š With

five sharp,

curved claws on each

European badgers are well adapted for a for food

in

and digging underground

extent.

paw life

(inset),

of rooting

tunnels.

79


I

_

_

Badger Mobility

B

adgers amble around with a

sniff

footed

gait, like bears.

and search

rolling flat-

They slow down to

carefully for food.

When

alarmed, a badger can sprint at 20 miles per

hour (32 km/h), but not for long distances. can also climb, using

its

It

claws and powerful

forelimbs to grip rough stonework or the bark of logs. Sloping trees are often explored, and

badger (3

may be found more than 10

hairs

m) above the ground.

w A cub

Resident badgers tolerate each other, while intruders are chased off.

each other by

is

useful for a

others fur social

to see in the dark, especially in the thick scrub

consists of

and bushes often frequented by badgers. Scent also makes recognition possible at a distance,

territory to

warn

off

Living in a social

badgers

around the

Work

Light

each would have to dig

12 inches (30

its

cm)

in

measure about

diameter. Even a small sett

needs a ton or more of earth to be dug out. a shared sett each

and might never dig much can be handed

it

left

The whole

all.

sett

the generations,

little

work

do except

to

result,

is

common among

bond the members

SEE ALSO Rabb

and

play.

social animals.

of a social

8

each

may be

six

animals, but sometimes

over 20. Each clan usually has a

than the females and consequently

move around more

within the group's area.

Territorial activity

greatest

and

is

is

found, particularly

Breeding sett,

in

the time that most bite

is

among

the early spring

wounds

males.

normally confined to the main

and only one family of cubs

will

produced, with most of the females not breeding.

In

are

some

be

in

the clan

carnivores, nonbreeding

females often act as helpers and

assist in

bringing up the dominant female's family. But

the badger

home

in

only the mother suckles the

young and takes them out

to forage. Badgers

to use helpers,

nonbreeders contribute

little

and the to the welfare of

the breeding group. Males sometimes play with

more time and energy

for feeding, breeding,

lick

part of the system for creating

about

do not seem

clean and perhaps dig occasional

Play

helps

at

the advantages of a complex

extensions As a are

as other animals

down through

burrow system with keep

In

badger can simply use the

same passages and entrance

all

If

and may

dominant male. He and the other males are more active in scent marking their communal

— hard work for an animal of such

a size, since the tunnels have to

Providing

there

all

ar other

bonds. Normally, a badger social group

territory

group has other advantages.

lived alone,

own burrow

left

badger trespassers.

Many Paws Make

80

groom one

also

nocturnal animal because visual signals are hard

and scent marks and feces are

in a British

enables them to recognize each other. Badgers

The animals recognize

their scent. Smell

forages for food

the cubs, but

do not help feed them. Most cubs

born within the group are fathered by the It

group and

males

in

manage

that

same group However, some boars

to sneak into neighboring social groups

woodland.

feet


WEASELS EUROPEAN BADGER

The badger

and mate successfully with females there. While

most badgers stay with

some

older males

their clan

may go

to

live

all

their

life,

elsewhere,

which helps prevent inbreeding. However, such individuals take a great risk.

Newcomers

often

get savagely attacked by resident badgers sustain serious bite

infected

and

wounds, which may get

and lead to a lingering death.

is

well

adapted for

the jaws have such a strong hinge that they

cannot be dislocated.

A

willingness to eat

vegetable material leads the badger into

unpopular

activities,

such as raiding cereal

crops. Besides eating the grain, they also

sett,

spring, but the

Badgers eat worms, small mammals, beetles,

delayed for

will relish a nestful of

rabbits or birds' eggs. But

members a

baby

although they are

of the order Carnivora, badgers have

broad diet and eagerly dig up a wasps' nest

or root out bluebell bulbs, wild

and even garden bulbs

in

as blackberries, bilberries, a favorite in

also

the

fall.

crunched up

in

arum corms,

pregnancy

difficult to harvest.

in

starts.

first

silky

white

year, usually

fur.

The

facial

The cubs spend about

weeks underground and until

Š As

before normal

they have pink skin covered

stripes are barely visible.

weaned

is

The delay ensures that the

with a thin layer of

and

large quantities.

many weeks

February. At

eight

Beech mast and acorns are

when

development of the embryos

cubs are born at the best time of

the spring. Fruit such fallen apples are

roll in

they are two years old. They mate during the

have a good scratch, and go off to feed.

and they

it

Badgers normally reach breeding age

At dusk the badgers emerge from their

for

grinding up tough nuts and gritty worms, and

Meat and Vegetable Diet

frogs,

varied

The molar teeth are broad and knobby

diet.

the ripening crop, making

and

its

will

not be

fully

they are about 16 weeks old.

carriers

tuberculosis,

European

badgers have come into conflict with farmers,

particularly in Britain. To

combat the

disease, culls

have taken place Britain

and

although their

20 pounds

effectiveness

by the end of their

first year.

in

Ireland,

They grow quickly and can weigh as much as (9 kg)

of bovine

is

in

doubt.


SMALL CARNIVORES

Honey Badger The honey badger

animals Honey badger

Mustelidae

Order

Carnivora

Size

Length head/body: 24-30 length:

The strikingly marked honey badger

8-12

formidable animal. in

(20-30 cm); height

in

Key features

at

animal with short

tail

and

brown elsewhere; head muzzle and small eyes to

Habits

Breeding

on back, black

small with short

not particularly large

it

is

without doubt one

animals, always ready to pick a fight with living creature.

Honey badgers

have been known to attack horses, wild

cattle,

°ot (3.4-m) long pythons, deadly cobras,

-f

1

Nocturnal, excellent digger; usually solitary;

whole packs of domestic dogs, and any other

extremely aggressive

animal they perceive as potential food or a

One

to 4 (usually 2)

young born

at

any time

possible threat. Attacks are often launched

of year after gestation period of 6 months.

quite suddenly from a

Weaning and sexual maturity unknown. May live up to 26 years in captivity, perhaps 10 in

vegetation. The

the wild

Voice

Harsh growls

Diet

Mammals, insects,

up

give

burrow or patch of dense

honey badger

have done nothing to provoke reptiles,

poisonous snakes,

and worms;

Anywhere with

quite readily

will

a perfectly secure location in order to

attack an unsuspecting victim that appears to

when angry

it.

birds,

also carrion, eggs, fruit,

Fearless Attacker

and honey Habitat

a

is

a typically shambling, badgerlike

almost any other

suitable sites for

forests, grassy plains,

dens-

and rocky slopes

The honey badger has

large, strong teeth

and

long claws. While they certainly serve as

Distribution Africa and southwestern Asia

effective

Status

Population unknown, probably a few

explain the extraordinary self-assurance with

thousand Widespread but declining

which the animal throws

weapons, they do not

apparent fearlessness justification,

extremely

is

itself

in

themselves

into

combat.

Its

not without

however, since the honey badger

is

difficult to

kill. It is protected all over with an immensely tough, rubbery skin that is

thick

enough not

to be pierced by

most

including the needle-sharp fangs of

snakes.

means firmly

and

Its

skin

is

that even

by the neck,

bite

its

Despite

SEE ALSO

venomous

the honey badger it

is

which

grabbed

can easily wriggle around

attacker. its

potential ferocity, the

badger generally prefers to mind

82

teeth,

also very loose fitting, if

It

will attack

its

1:76. Badger, European 1:78. Jackal. Black-Backed 2:62

honey

own

not entirely

very difficult to

is

of the world's most relentlessly aggressive

short, sturdy legs with extremely long, strong

claws; fur usually grizzled gray

and has

appearance. However,

15 5-28 5 lb (7-13 kg)

Solidly built

It is

(60-77 cm);

shoulder: 10-12 in (25-30 cm)

Weight

animal

sora capensis

Family

t

its fearlessness is

(ratel)

unjustified, since the

name Mel

tail

a natural aggressor

is

larger than itself with remarkable

ferocity However,

Common name Scientific

much

Mellivora capensis

kill.


WEASELS HONEY BADGER

business and areas

is

seen

rarely

where the species

common.

It

its

den only

its

in

relatively

is still

emerges from

and spends much of

the open, even

in

at night

time digging or sniffing

around for small food items.

Sweet Tooth As

its

common name

badger has

a

suggests, the honey

sweet tooth and

fond of honey.

It

tracks

down

is

particularly

bees' nests

often by teaming up with an unlikely partner, a bird called the African

honey guide. Unlike most

small birds, which stay well

away from

honey badger, the honey guide out the badger, attracting special

Honey guides

call.

its

hungry

a

actively seeks

attention with a

like

honey too, and

they especially love the nutrient-rich

wax

honeycombs made by bees

to store

honey and

grubs. However, the bird

not strong enough

is

to break into a bees' nest on

its

own. Having

discovered a hive, the honey guide

calls for

help.

The honey badger follows the

hive,

breaks

it

open with

most of the honey.

Its

its

bird to the

and devours

claws,

thick skin protects

it

from

the stings of angry bees. Meanwhile, the honey

guide gets

its

reward, feeding on the remaining

broken honeycombs, which are scattered and accessible

now

widely

owing to the badger's

destruction xaf the nest. Other creatures also benefit from the

foraging

abilities.

honey badger's strength and The black-backed

example, pounces on rodents and

jackal, for

reptiles

flushed out by the badger. The badger gains no

advantage from association with the

jackal.

The breeding biology of honey badgers

is

not well known. Animals are sometimes seen traveling

in pairs,

but usually they are

The young are born

in

spring

in

solitary.

the north of the

species' range, but breeding appears to

seasonal to

six

in

much

be

of Africa. Gestation lasts

months, and the young are cared for

less

up in

a

secure den lined with dry grass.

B Although honey badgers find most of their food on the ground, they are

known

to scale tall camelthorn acacias in

search of raptor bird chicks.

83


r SMALL CARNIVORES

Striped

Skunk

Mephitis mephitis

Skunks are one of the few mammals

When

defense.

to

use chemical

threatened, they squirt a repulsive-

smelling liquid at their attacker Scientific

name

ephitis mephitis

Family

Mustelidae

Order

Carnivora

Size

The striped skunk

Central,

Length head/body: 12.5-18 tail

7-10

length:

shoulder 4

in

in

Breeding

lb

(1.5-6 kg)

striped skunks serve the

After a

first

Mainly active at night and at dusk and dawn;

that sees another TKunk will

generally solitary; squirts foul-smelling liquid

dramatic coloring

may swim

necessary

if

62-66 days

mature

captivity,

at

fewer than 3

in

the wild

vegetables,

wooded

Distribution Southern Canada,

U

S

ravines;

,

tail

around the

and

may be

a

tipped with white, and

ears.

The

on the back vary

stripes

length and width, so that an animal

almost entirely black or white or variations in between. There

town gardens

may appear

many

no difference

is

in

in

patterning between the sexes, and the

and northern

markings do not change with the seasons.

Mexico Status

body with

there are often white spots on the legs and

Forest or field edges, patches of brush, rocky

outcrops, and

The

tail.

garbage Habitat

a hasty

variable, but usually

white stripes that extend from the hood to the

Mainly insects; also small rodents, rabbits, fruit,

make

probably

will

white blaze on the nose, a white hood, and

growls, grunts, and snarls; also churring and short squeals; occasional screech or hiss

eggs, carrion,

that the

associated with a

consist of a predominantly dark

Low

birds,

remember

(including

Weaned at 6-8 weeks; year May live 8-10 years

1

are a warning signal.

The markings are

retreat.

as the

unpleasant encounter any animal

repugnant smell and

Three to 9 young born May-June after delayed implantation).

Diet

wasp— they

white stripes on back; white patch and stripe long, bushy tail

threatened;

markings of

same purpose

on head;

gestation period of

Voice

striking black-and-white

is

Population abundant

Foul-Smelling Spray

A well-known stink.

bad

fact

about skunks

Even their Latin

smell.

The smell

is

name their

is

that they

Mephitis means

one

effective

defense mechanism, since skunks are not

fast

runners, vicious fighters, or clever at hiding.

When

a skunk

is

threatened,

warning display by stamping

warning

body

84

see

also

i

t

Family,

its

is

into a

raising

forefeet

it

will first

its tail

give a

and repeatedly

on the ground.

If

the

not heeded, the animal curves

"C" shape so

The 1:88; Fox, Red 2:64

5 A striped skunk on a log

on a

m

family of their

in a

own: the Mephitidae.

stripes

sexually

North,

from other mammals that

they are frequently classified

Cat-sized animal, with small head tapering

when

live in

the world. Skunks are

in

tail

to a bulbous nose; black coat with forked

Habits

skunks

of the

and southern America and do not

sufficiently different

The Key features

common

most

All

occur anywhere else

(32-45 cm);

(17-25 cm); height at (10 cm). Male larger, but in

female has longer

Weight 3-13

the

is

10 species of skunk.

that

it

its

can point

its

in a field

rests

of

flowers. The white

forked stripe that runs

from

its

head

to its

haunches distinguishes it

from other species.


5T =

WEASELS

anus at ire target

.while

PEC

S*

JKH

keeping a dose e.e on

adversary Theskjn*. then squ hs a ye o.vish

its

ng

ffou;l-sme

norm

sera,,

an us

either side o* the

a*

;>e

.tie :

.he

_

comes th'auc"

mus<

"-e se'=

pouch es or

""uscl; 5'

.2

.

a n.pple

that car oe a~g:ec to mpiove the s<ur< a

can

im. It

hit a target at a distance

to 6.5 feet 1""

times

«era

up

S*xrks are able to spra.

and the

quick successor

in

off

3

car ether oe a tre .apor or a more

shower

uiiected

spra nq ihemse

.

.

scent .vih tre

'

and do not

es.

dh

.vr

m

1

r to ire

sirne

e (0.5

e.es,

temp:

it

sulfur

~

2

tr

^

reread

:

r.

-

'

r

xr

.

e .5 ^

gets

-

-

_

e

5 :

The posturing

"-a sprays r

:

ffor

.

r

-

-

t

.

is

t

to

.

^

r

on the Menu

Insects Striped

_

"

it

taxes ^p to -tS hours

ng before the an

T. r

m

iff

and even

causes extreme p it

t

1

<m downwind; and

undress

display

bad eggs,

-_"r a"s ca" s^e

the skunk to rep ace the must

anc

scatta' the

ffrorr.

also sulfur oasec

is

up to

<e the

""as aw oo

peop e oe e.e

as man_,

la

The mus< contains .cat e

compounds.

r

crop els ’"le

off

s<

omn

j"<s are

.ores The_. are

opportun Stic ffeece r s. eat rg almost ar /thing that appears .ague

o' the

r

.

ec

made up

diet s

About ~0 Perce":

ble.

nsects,

off

such as

grasshoppers crickets oeeTes, pees, ard

wasps.

vr a

Ore

off

their favorite

foods

ch they dig up from the so also feed

on sp

s"a

cers,

is,

s grubs,

Striped skunks

earth worms,

clams, crayfish, frogs, saamaroe’s, snakes,

b

rds eggs,

sma mammals,

nuts, roots, gra ns,

carrior

.

berries.

and garbage

,

Most s

ffooo

ts

ocated by sound or sme

nee the s<un<s distance

T aociuon, the an, ma

.ision

s are

is

poor n

too slow to chase

fast-mow mg prey, nstead. the. hunt cats, lying in

victim

..a

a act or stai<ing the r

"a.'

Cu.m pee. es ano

grasshoppers by sprung ng on

them with the ' forepa as

85


Urban Warriors

Females prepare maternity dens

The young are born rban areas provide everything that a

U

skunk needs.

that there

Human mess means

always plenty of garbage to

is

feed on, and where there

is

garbage,

Skunks

They can

and

dig

will also

there are several animals

if

the color patterns are

birth

patches on the

residence, the smell can be overpowering.

gets too close

is

in

likely to

towns.

Any human

be sprayed.

that

dark

Males do not help rear the

skin.

and may even attack or

The three

kill

them, so

kittens are fast-growing: After

six

to eight

old.

about

weeks they can assume the defensive

hunting Skunks are unwelcome

visible as

posture and squirt scent. They are

in

are

females defend the maternity dens aggressively.

up lawns for grubs.

burrows under buildings;

live in

mid-May and

March.

wrinkled, blind, and almost hairless. Even at

kittens

there are usually rats and mice as well.

in

in

at

weeks and follow the mother on

trips

When

weaned

when

they are about two months

out with their mother, they keep

close behind her

in

a single-file

trail.

By August

they have reached adult size and are able to

fend for themselves. They

will

be able to breed

the following spring.

They

will

use their long front claws to dig for

ground

as mice, rats, moles,

beehives and

will

eat the inhabitants without

appearing to be affected by the

known

to

consume the bee

They are

stings.

and probably

larvae

the honey, too. Skunks have a special trick for dealing with poisonous or hairy caterpillars.

Before eating them, they

ground with successfully in

their

the prey on the

roll

forepaws, an

removes the

activity that

irritant hairs

or toxins

the skin. Striped skunks have also been seen

breaking eggs by rolling them between the hind legs with their

forepaws

until

the egg strikes a

stone and cracks open.

A

Skunk's

Way

of Life

Striped skunks are not social animals. They

come together

to breed

in

the rest of the year alone or of mothers

home

and

their

spend

spring, but in

made up

groups

young. Females occupy a

range of 0.8 to

1

square miles

.5

(2 to

sq. km). Their territory will overlap that of

4

many

other females, but they tend to avoid contact

with each other. The males travel farther

much

—about 8 square miles (20

cover the

home

ranges of

many

86

—and

females, as

well as overlapping with other males.

mate with many females

km)

sq.

The males

living in their

SEE ALSO Raccoon, Common

range.

1:22

use dens above ground, In

although they do not hibernate, they

and

skunks also break into

squirrels. Striped

summer skunks

selecting hollow logs or rock piles.

grubs or to tear apart the nests of small

mammals, such

In

winter, rest for


WEASELS

Š

Two young

SKUNK

STRIPED

and

striped skunks take in the sounds

smells near their birthplace a hollow log den. ,

sometimes spend the winter

in

the

same

burrow as opossums, woodchucks, or occupy

rabbits, but

different

cottontail

chambers.

Death by Natural Causes In

skunks may

captivity

the wild

for

live

10 years, but

in

more than 90 percent never reach the

age of three. Natural causes of death include starvation disease.

long periods

made

in

underground burrows, often

by woodchucks or badgers.

dig their

own

Some skunks

burrows, but they are not very

long or deep. During winter striped skunks

Š A striped skunk digs for

worms and

favorite

food

grubs,

its

in a largely

insectivorous diet.

den alone or together animals. in

in

will

groups of up to 20

Communal denning

is

range.

It

seems that skunks

usual dislike of

company

will

overcome

to take

Skunks are preyed on by great horned

owls and some other birds of prey

—

all

Mammalian predators only

will

on the verge of

if

enough

to deter

A common

all

hunt them too, but

The smell

starvation.

advantage of

is

but the hungriest hunter.

cause of death

skunks

in

disease. Striped skunks are susceptible to their

which

of

have a poorly developed sense of smell.

more common

the colder, more northerly parts of their

harsh winters, predation, and

in

and

diseases, including leptospirosis

is

many

rabies,

which may be passed to humans. Rabid skunks are often very active

present

in

and aggressive. The

their saliva, so animals

only catch the disease

Humans

if

virus

is

and humans

they are bitten.

many skunk

are responsible for

deaths, including trapping for the fur trade.

Skunks are also

killed

attracted to roads

Skunks

by vehicles, since they are

when

at

searching for carrion.

Your

Service Ithough skunks usually have a bad

b

%. reputation, they probably

good than harm. They

eat a

of agricultural pests, such as

do more

huge number

armyworms,

cutworms, Colorado potato beetles, grasshoppers, beetle grubs, and sguash bugs. They are also

and

rats.

money

in

good

at catching

Skunks save farmers

mice

a lot of

terms of the amount of

pesticides that

would be needed

if

the

hungry skunks were not there.

87


small carnivores

The

Civet

and Genet

Family ivets

C

and genets are

small- to

medium-sized

animals with a catlike appearance. The family

and

Viverridae (civets, genets,

linsangs)

and the

Herpestidae (mongooses) are similar to an ancestral

Family Viverridae: 6 subfamilies, 20 genera, 35 species

carnivorous in

mammalian group, the Miacoidea,

the Eocene period, about 56 to 35 million years ago.

Miacoidea

fossils

genus,

'

Nandinia African palm

PALM CIVETS

civet

What

Is

1

Paradoxurus

3 species,

palm

in

species, small-toothed

palm

common palm

civet (A. trivirgata)

civet

(P.

civet

(P.

many ways,

Civets

hermaphroditus)] golden

palm

civet (P zeylonensis), Jerdon's

species,

1

and linsangs are the

Arctictis

1

masked palm

civet

and

short legs

there are also a

their relatives

number

species, Sulawesi

palm

BANDED PALM Hemigalus Diplogale

1

civet (M.

have long bodies and

compared with

cats.

palm

civet

They also have a longer,

vegetable food, so are

Cynogale 2

species, otter civet (C. bennettii); Lowe's otter civet

civet (C.

owstoni)

A

2 species, spotted linsang (P pardicolorY

,;

Viverricula

3 species. Oriental or

civet

banded

civet's

or stripes.

linsang

Malayan

(V.

indica)

civet

and

Eupleres

(V zibetha), large spotted

generally sandy gray with dark spots

is

tail is

usually

marked with bands or

civet

FOSSA

1

1

Some

masked palm

civet

is

uniform

is

a

species,

The

color.

color except for

in

(V megaspila its

face.

Some

and genet species are

civet

1

species, aquatic genet

species, including

armed with

common

genet (G genettaY. large-spotted

When

in

danger, they are able

The

to spray a stream of vile yellow fluid at an attacker. liquid

2 genera, 2 species

species, falanouc (f

stink glands.

(0 piscivora)

is

nauseating and repellent, and

may

temporarily

blind or cause discomfort to the attacker, so giving the

goudotii)

intended victim a chance to escape. Fossa

powerful claws.

species, African civet (C civetta)

CIVETS 1

shorter, less

such as the binturong, have a coat of a single

(V tangalungaY, large Indian

genet (G hgrina

MALAGASY

coat

The

markings on 1

Oenetta 10

prey, large

different color than the rest of the body.

species, small Indian civet

1

Osbomictis

Cats have

7 genera, 19 species

species, African linsang (P richardsoni)

Civettictis

less specialized killers.

genets have more slender, weaker jaws, comparatively

(P linsang )

Viverra

on hunting

canine teeth, and large, powerful claws. Civets and

small canine teeth,

TRUE CIVETS, LINSANGS, AND GENETS

Prionodon

cats are

lowei

(C.

1

and

viverrids

jaws for gripping struggling

short, strong

1

Poiana

between

of the differences

animal prey for food, while civets and genets take more

(D hosei

Owston's banded

their feline relatives.

linked to their lifestyles. Cats rely totally

Chrotogale

species.

relatively

musschenbroekii

CIVETS 4 genera. 5 species species, banded palm civet (H. derbyanus)

species, Hose's

1

of differences.

larvata)

(P.

species, binturong (A. binturong)

1

closest relatives of

jerdoni)

Many Macrogalidia

herpestids.

a Civet?

more pointed head and face than Paguma

and

viverrids

the cat family. However, although they are similar to cats

binotata)

5 genera, 7 species

Arctogalidia

the skeleton

similarities to

species

1

(A/,

show many

and tooth structure of modern

Civets, genets,

AFRICAN PALM CIVET

that lived

It

may be

that

some

species, fanaloka (F fossa)

genus,

1

of the bold coat markings

found

warning to predators,

the distinctive patterns of a

in civets

serve as a

species

Cryptoprocta fossa

(C

ferox

like

skunk, discouraging them from making an attack.

In

other species the markings help break up the outline of the animal, protecting

88

see also

i

net,

Common

1:92; Civet.

Common

it

by camouflage.

It

is

important

Palm 1:94; Fossa 1:96; Mongoose Family, The 1:98


©

>4

binturong forages for

fruit in a tree.

dwelling species with a strong while picking fruit with

its

tail,

which

The binturong it

is

a tree-

uses to hold onto branches

and compact, and they only walk on

dwelling species generally have broad feet with bare soles, a factor that

paws.

their toes. Tree-

improves their grip

when

climbing.

They use the whole of the foot when walking, for civets

and genets to have such defenses because,

bear,

and use

all

like a

the toes to get a better grip. Feet also

although they are hunters themselves, they are also

vary depending on the type of food taken by a species.

targets for larger predators.

Linsangs, like cats,

Different species of viverrid have different lifestyles

and types of

prey.

Hence there

within the family. Although

some

— such as the true

all

civets

is

a range of

species are

body forms

good

climbers,

rarely climb trees to find

food. Such species tend to have feet with well-padded soles that are covered with hairs. Their

paws

are small

themselves and

live

almost entirely on animals they

like cats

— have

fully retractile

kill

claws

with protective sheaths. Species that eat a greater variety of food have feet that are less specialized for prey taking.

The form of the teeth

also varies

between

species.

The

meat-eating linsang's molar teeth are sharp and bladelike, similar to those of cats. True civets eat

both meat and

89


AR\ VORES

small

of the

So .~w

owt

fom//y

ihown

*eedng

-acten st>< activities *f~can

on a r*st1<ng

(II.

in

bonded pa-m cnet eatrng •V*v

Ma ayan

3 i,;ord (21 Oneoiji or

common pa m cnrf

scenting the air (41

b>nturong foraging for

fruit

grasping a brand) ivirt

its

prehensile

tail

crvef fJl

white

(51

vegetable matter, and have

broad-crow ned molar teeth that are suitable both for cutting

and

grinding food The paim civets

take mostly vegetable food. Their

molars are even broader, making

them

chewing

less suitable for

meat but better

for grinding

up

vegetable matter

Un ke

which have rough

cats,

tongues that they use to clean, ovets

tongues because

dogs

and genets have smooth

ve dogs This

in

bones

lick

the

is

same way

as

— they crush and swallow

:ne bones of their prey and so not need to

do

the bones clean

lick

The form ot the sku

;1

also varies

betyveen speoes Genets have catlike white the skulls of civets are more heavily

skulls built

Palm

civets

have comparatively sturdy

The stand of Madagascar

skulls

is

home

to three unusua

speoes that are found nowhere ese me udma the

Where

Civets

TH e argest

and Genets

number

in

T

rue civet fossils have been

Madras, southern

m

the rocks of the Siwalik

p

>v

d

Hills

members

throughout most of Afnca.

India,

southwestern Europe, the Near

90

ns*,

m

g evidence that they have

preh .tone times Today

FV"

i

pantherl se fossa Palm civets

of vivernd species are found in India

•md the Malay Peninsula d covered n a cave

Live

(

u

(

,

and

also

northern India,

ved

in this

v

Asian although there

s

one African speoes The

true cnets a^e mostly confined to As a

Lmsangs are found

n both Atnca and As'3 Genets are found only but

A*nca

one speoes occurs n southwestern Europe Most

region since

:n

and genets

species of civets

live in

forested

of the family are found

areas Thev are found from ram forests to woodlands, n

and Southeast Asia to

woody

scrub and occasionally in savanna Only

spec

of civet

East,

and the Arabian

ere are no ovets or genets

SEE ALSO

India,

genera

and banded palm ovets are

h* 2:10. Dog

m

the Americas

Earn, h

The 2:S0

sma

'Os

manage

Indian civet

speoes

w

ill

live

to

and the

ve

m

drv

open country

common palm

on mountains up

two

civet

to a height of

—the

Some


THE CIVET

Civet Oil

C

source of civet (or musk)

oil,

a scent

making perfume. The word "civet"

used

actually

most

that issues from the glands of

Indian civets, are the ones used

Genet

industry.

animals,

is

and

The

oils.

The

a clear,

fluid, collected

and small

oil

and

civet

oil in

dates back at least to the 10th century

was imported from is

it

Solomon. Civet

Africa by King

believed to have medicinal properties

and has been

and Asian countries Civets

of

animals.

but East African

export large quantities.

and genets use

for scent marking.

means

still

oil,

their

The process

the most important

communication between these

It

is

used to define

territories

generally In

The true

Africa genets occur

and are

particularly

are almost

all

grow

them

are ground dwelling

habitats except desert

in all

living in

or hollow tree.

Some

in

some

Viverridae. There are three species, secretive animals

found only

even build themselves nests up

in

females

civet

and genet

otter civets.

They have smaller

body form,

in

still

tails

good

come together

briefly

a rock crevice, burrow,

may

and

during the breeding

season; but once mating has taken place, they separate,

and the female

raises the

exception to the rule lives in pairs.

is

young on her own. An

the Madagascan fanaloka, which

3

/

The fossa has a catlike head and

retractile

claws for capturing prey

shown here (2)

and the

(1).

Also

are the mongooselike falanouc foxlike

are small,

civet,

Madagascan fanaloka

than other

such as

(3).

civets.

their toes are less

at climbing.

a thick

alone. Males

all

ears, a blunter muzzle, a

the branches.

live

and

of the

forested areas.

in

Lifestyle

Most species of

fully retractile claws.

members

of the rarest

more compact body, and shorter

species, such as the linsangs,

the trees. Apart

have dark rows of spots

all

coat for the winter. Most species forage and hunt for

food at night and spend the day

and

viverrids.

carnivorous and have

webbed, and they are

cold places they

and

civets are a highly varied

adapted to

While resembling otters

In

the

almost entirely

There are also some aquatic species of

during the breeding season to attract females.

7,000 feet (2,100 m).

civet takes

in

or stripes along their bodies. Like the linsangs, the genets

solitary

and by males

Banded

the trees and are generally

in

more doglike than other

Linsangs are

pungent secretions is

time

from the aquatic genet, they

The development of synthetic substitutes

has reduced the need for civet

less

group, but the majority of

oil

used as a cure for sweating and certain skin conditions.

spend

civets

invertebrate prey.

when

paws. The diet of palm civets largely

mammals. Owston's banded

perfume

b.c.,

its

uses to hold onto branches while

trees at night for invertebrates, small lizards,

has an offensive smell to

The use of

diluted.

it

a

(capable

tail

more carnivorous, foraging on the ground and

from captive

humans, but becomes a pleasant scent when purified

The binturong,

consists of fruit, but they also take animal prey.

palm

yellowish-brown mixture of fats

original

which

of grasping),

picking fruit with

the commercial

in

climbers that use their

species of palm civet, even has a prehensile

scent has a subtle, pleasing smell, but true civet species, such as the African, large Indian,

skillful

semiretractile claws for extra grip.

liquid

viverrids.

both tree- and ground-dwelling species. Palm civets are mainly tree dwelling and are

in

comes

from the Arabic word zabad for the scented

perfume

and genets are good climbers, but there are

All civets

and genets are probably best known as the

ivets

AND GENET FAMILY


SMALL CAR,M VQK£S

Common Genet

GzieUC

common genet is

The

Ce"r;;j

the only species ofgenetfound

outside Africa Individuals ,vere kept as pets tv the amici'. creeks and Romans before the i

domestic cat

became popular Family

»e"-ade

.

Order

4

U-rg— '•*» bocv 6-22 r A 3-5 5 or 3 e^gm T6-2C n

S<ze

-

-

-50 err

L -

-8

n

(1

8-20 cm ’

ces

st" -

r

ta

uo<

sam

Africa,

me

sects farming c rc

arc white tip or enger far dovvr spire -rigs

burrg

Act .e

l 're's of ’-4

after

May

/e

up

to

w*t

may orgra

ca.

tip

G ore. tar

on

t

of Spain with Morocco n

may

aiso

20 years

have oeer*

intentionally

“traduced to Euroce by humans n the

me common

Ages,

to

islands

were

genets

definite'/

ip

common

Voce

the Saearc

on-

ta*en there ov people

-genets are s&i

found

iim

Europe,

Vx-ra y ”ev<

occurred as ran north as Be-gium and Germany

'.'cst £>.

Kab«tat

silent. but w-ll hiss, growl, purr and "a/ also cough, whine, or scream

,

'oderts. especially mice also rabbits, izards. fruit, bernes.

semto. -voociand. ^

SCL

*t

_

300

-

Creatures of the Night

and nsects

and rocky

areas,

Common

jp

n

rr

Distnbutxxi E.-ope France and

me berar

Peninsula

and Afnca (except for the Sahara Desert and Congo River Basing •

es* ne.

exua^or abundant

r-e-

UCN

genets are ncctuma

dawn,

dusK.

and most of the n

rest at

some

point

and

w

dunng

Vulnerable

(

ght. They usually

the hours

mer

rock crevices or burrows

Stand subspecies, Widespread and

being active a:

n

occasionally be inactive

morning They spend

Alternative

1

/

they

same

shelter

the eahy

davs tucked away in

dug by other animais.

may use fee

hoi

ows

square

m

es 5 sg

femates tend to be

'ange

may

or

-use

the

day after day for long per-ods

Va-e genets have home ranges 2

darkness

off

sometimes large bracches They often

km n -

much

size

smaller.,

overlap the ranges

off

off

up to

The ranges of

and a male's several

'em a<es Latnnes and scen-t-mark-ng ytes are used by more than one ndiv<dua soiitary-

pa»rs.

92

y

but omy in the southwest. At one time they

b*

Status

A

th

’Oday

Wxe

most a regions excect

once connected

qestator

Sectemoer Weanea at 6 montns. sexually /ears.

that

the southern

per-od of 7 ; aays 5 reedng occurs, yearCure, but mainly Apn to Matt and August

'•ture at 2

urn a-

reached southern Europe oy -crossing the

and bridge

I'iu-

.curg oern

found

others,

umps. and svvms well

cJ rpps.

Breeding

~eerrgs with

is

s.ongo River Basin. The speces

me mgbt arc twi light perccs

soitary out tcierates

t

Sanara Desert and dense forests of the

'•ave

Habits

cutsde Afnca, the common

ge^et s also the most w-despreac genet m

m

iVith

3arK crest

-

species a* ge^et

5-2 kg

*ey features 5ancv c.v

3T mais With

pointed muzzles. As /ve as being the or y

« *

ne-grt at shoudfer

Weight 3-5 b

GE*iE~5 ARE LONG, TWIN CATLIKE

SEE ALSO

*

1:56

Genets ar?

animated but they sometimes

carticuany

dunng

travel- «i

t^e breeding season


"

C

.

E

5

i'»:

SESE'S CC i

'• :

1

1


I*

SMALL CARNIVORES

Common Common name Common

palm

Civet

civet (toddy cat)

Scientific

name

Puradoxurus hermaphroditus

Paradoxurus

hermaphroditus

The

Viverridae

Order

Carnivora

Size

Length head/body 16.5-27.5

common palm

eating

fruit

civet is also known as the toddy cat because of its habit of raiding

Family

tail

F&lm

which ferments

(42-70 cm)

in

16-24

length

plantations in search' of the sweet sap of palm to

make a

mildly alcoholic "toddy/’

in

(41-60 cm)

I

Weight 6-10

The

lb

(2.7-4 5 kg)

down

and base streak

of

to dark-brown coat with black

back, spots on shoulders, sides,

tail,

face has

on the forehead,

mask

tail tip

of spots

and a

may be white

time

Nocturnal, expert climber, spends

much

2-5 young born mainly between October and December after gestation period Litters of

months Weaned at 6 months; sexually mature at 11-12 months May live up to 25 years in captivity, about 10

in

civet. Instead,

fallen fruit

Latin

it

civets

most spends

It

less

most other species of palm

forages on the ground for

and small animals

only active at

is

It

and spends the day curled up

in

the

name, Paradoxurus hermaphroditus,

scientists is

Diet

Small vertebrates such as mice and lizards,

and

difficult

often confused fruit,

A hermaphrodite

to sex.

The reason why the two sexes are

silent

also insects,

in

the

common palm

can make a male animal look areas,

may

shelter in

Palm

thatched roofs and pipes Distribution Kashmir. Indian Peninsula, and

female Sri

islands of Hainan (China). Sumatra. Sulawesi, Simeulue. Enggano, Kangean Islands, Java

Palawan

(Philippines),

and many other nearby

is

like a

female

civets are solitary animals,

will

only tolerate a male

in

and a

her tree

Lanka to

southeastern China and Malay Peninsula, also

(Indonesia),

civet

that the scent gland, located near the genitals,

and seeds

and wooded

a puzzle to

is

an animal that has both male and female

genitalia

Normally

Forests

of the

reflects the fact that the species

the wild

Voice

Status

one

branches of a tree or the hollow of a trunk The

of 3

Habitat

is

time

in trees

Breeding

trees than

in

night

Habits

palm civet

widespread species of palm

Key features Sandy gray stripes

common

Borneo,

during the breeding season Mating takes place in

June Dominant male palm civets have larqe

territories that

number

overlap the living space of a

of female

islands

palm

civets,

but they

will

generally allow smaller, immature, or elderly

Population abundant. IUCN Vulnerable (subspecies P h hgmcobr A common

males to be present However,

animal

male matures, he

is

a threat to the

male's right to the females usually determines

keep the

territory

when

which

in

young

dominant

A

fight

two males

of the

Dominant males

ranges, covering the

a

the area

whole area

will

patrol then

at least

once

every five to 10 days

Sowing the Seeds Although they take some animal civets eat a

m

*

Java the least

vines

94

SEE ALSO Genet. Common

1:92, Fossa

V96

wide

variety of fruit

common palm

35 different

When

fruit

trees, is

prey,

palm

For example, in

civet eats

the

trees,

fruit of at

palms, shrubs, and

eaten, the hard seeds are


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SMALL CARNIVORES

Fossa

Ca -p to p rocta feim

The fossa resembles a short-legged, dwarfpuma a powerful predator and has a fierce reputation in native Madagascar.

a testament Common name Scientific

name

:

Order

Carr .ora

m-v

The fossa's

Zrycroprocta fern*

Vivemdae

was often

the Fe idae.

t

:

Ler;— nead/body 24-31 in (60-80 cm, engm 24-30 in 60-75 cm height at

14-35

sroc der

Key features Redash-brown

to

tail

occasionally up to

is

daws but ur toes

short, thick,

head with rounaed

weobed and have

Mar

and

active

mostly

Liters of

2-4 young

of 3 >

in

trees

and

is

20 years n

at

flat-footed

family, reflecting simi anties

The genera;

ke bears.

l

paced

s

in

the

mongoose

with that group

view, however,

is

is

that the

bom

after gestation

at

4-6 months; live

probably fewer

a cat Hence

a civet trot lives like

fam

y

Vivemdae Perhaps

result of long isolation

4 years May

captivity,

typical

member

s

it

of the

a skilled

predator; solitary except

months Weaned

mature

body than

its

—which wax on them

in

dunng the breeding season

se» -ai

ike true cats

—fossas wax

too.

puma, but nas

cat family. Fossas have cat! ke

Sometimes the fossa

ears, feet

the

in

the cat farm y

n

classified as a rather special ,es

c moer ana powerful

penod

dassifieo

that

short, retractile claws

noct^ma but can be

y

day* gn:

Breeding

distinctiveness

its

on the

is

a

island of

Madagascar, allowing the species to evolve into

something different from

its

ancestors on the

to at least

African mainland. in

Despite being a

the ferocious predator,

wild

Top Island Predator

Voice

Generally

Diet

Lemurs, small mammals, and

sil lent

rept'les. frogs, snakes,

Habitat

Ram

skills.

rid uding

cats,

meant

n relation to

members of the

•ossa

Habits

with

s.nilar to a sma:

is

neac

a longer

dark-brown coat,

occasionally biack. *ur ca: <e

.

cm)

Weight '5-26 b 7-’ 2 <g 44 lb (20 kg

5

s.mila,=;ties

ts strong predatory nature,

past t

are

diet

cssa

Family

Size

hunting

to its formidable

its

Lemurs provide much of its

:

It is

forest

and

‘ossa

Fossas are the top predators

Madagascar and

in

is

~cw

Traatarec

re

a

species,

birds, also

insects

and wooded savanna

have no natural enemies apart from people

largely

They are also Madagascar's largest land

clearance. 4 single arirai

mamma

agile in trees than

occupies an anaa at 4

a trunk headfirst and

sauare rules (IQ to 20

1

more

Fossas are

due

:c forest

a*

Distribution Madagascar

They can run

cats

Status

Population probably only a few hundred,

IUCN Endangered. CITES

jump about 6

down

feet (2

m) between

They

trees.

SO-

need

to

be excellent climbers

in

order to catch

the lemurs (the primitive primates of the

Madagascan

forests) that

half their diet

It is

feature regularly

in

very unusual for primates to

the diet of any animal, since

much

better

climbers than most predator species The fact '

that lemurs are often part of a fossa's diet

shows what an

agile

Fossas are as at they are steal

SEE ALSO

,

zestr-ctcr

1:

2:42

l*

m

and

skillful

home on

hunter

it is.

the ground as

the trees and have been

known

to

chickens and sheep, and take small wild

4

*

-

-,"J

of its habitat s ea'j'rq forest areas toe

make up more than

they are generally smarter and

96

km arc

II

4

—l

small to succor?

sec-re zccuacors.

1


4*

-

pigs Tr-e native cere

e of 'Madagascar consider

thief,

^not an sight.

mgnt

in

visit

mating About a week,

the fossa to be a dangerous and ferocious chicken

allow several mates to

later a

and known offenders are o~er

arrive at the tree to

When

and the resident female

local

the forests, they

cecc e spe r d :he

w

I

often <eep a

her one at a time for

new female wr

mate with the males there will

leave

hire

going throughout the hours of darkless

Unusual Puberty

because they are frightened of being attacked

Dunng puberty

by a fossa while they are sleeping.

strange stage of sexual development: The

Although fossas

camps

for food, they

occasional!',' "aid

do so very

human often

discreetly,

the female fossa goes through a

becomes

clitoris

large

male

similar to the

reputation as a dangerous animal

greatly

colored underparts

at the

orange

exaggerated, and

it

will usually flee

first

male

sght of a human. The mating behavior of fossas

is

different

from that of other vivemds, which usually mate with the one or

two animals

temtones overlapping will

spend

their

several days high

own: A female

m

a tree,

fossa

dunng

which time a number of males group together beneath

it

and

call

up to the female She

will

liquid that

It is

not

penis. Also, the

is

-i

creamy-

become covered

by a bright

known why females go through

side effect of the

and

it

may

simply be a

hormonal changes that occur

trees vim e

emu's. Fcss85 s/xare characteristics

rats but are i trees.

C3 fch

is

now

officially listed

homeland through greatest threat to

demo

as

Endangered by the IUCN. The destruction of

its

wr—

more ag e

the, are acve fo

and

<

lemurs,

despite those an

puberty.

The fossa

fossa leaps

between

usuai lv only released by the

this ''masculine" period,

dunng

that have

small

supporting bone and a coven ng of spines

not even waking guard dogs. The fossas is

and develops a

nte<

-v

5

gent end

capable creatures.

forest clearance poses the

its

survival

97


SMALL CARNIVORES

The Mongoose Family _

M either

ongooses are extremeK

ear v carnivores, the Miacoidea active

and aa e

terrestrial

What

form to the

similar in

that

sma

live

qroups or alone With the exception of recent

'

product ons mongooses are

rest-

ha

cted to the Old World

Taste: n Hemisphere, excluding Australasia).

Some

nclud ng the West Indies,

and the Hawaiian

Fiji,

attempts to control loca vermm, such as

rats

Herpestidae 2 subtami es

rounded

and snakes.

1

1

ounces <312

1

pounds

same

nvm

species

Bdeogjle

specie

5

mongoose

paudnosus)

mongoose

Kludi "g bushy-tai ed

mongoose

lacCsonS

vA

cm) and weighs

es

-

inches

002 cm) and weighs

family

is

closely related to the civet

They are sometimes

family.

classified as the

all

Mongooses

family, the \ iverndae

and genets (B crassrcauda);

in

differ

Cynktis

mcluc ng Alex.5 nder

mongoose (C

s

rounded ears that

rarely stick

out above the profile of the

speces dwsrl mongoose

Herpesfes

5

\

speces

,

l»6eo«tTS

s;>\ e-

t it»er.

They also

their toes.

differ in their

parvula)

remain on the ground and can be active during either the

mongoose (H ichneumon)]

day or night depending on the species

e ng Egyptian

e’

no

nocturnal, and solitary Mongooses, however, tend to

social

Mf

speces.

or

behavior Civets and genets are generally tree dwelling,

ata)

long-nosed mongoose iH naso)

fthynchogak

little

alexandri),

owes mongoose {D d’bowsi.ii)

NWogale

civets

that they possess nonretractile claws,

webbing between

soecies yellow "icngoose (C oentc

Dokygjtr

from

>acksoni)

(5

mongoose (C ansorgei1

Angola's

just

largest species, the white-tailed

head, four or five toes on each foot, and Crossarchus

total

OfT.i. 70

:

species

AtiUx

Their legs

<5 kg).

The mongoose and genet

The

g).

body

of about

long, nonretractile claws.

” inches <43

1

tail

dwarf mongoose, has a

species, the

mongoose measures 40

17 genera, 35 species

MONGOOSES AND MEERKATS

AFRICAN AND ASIAN

a tapering, bushy

and the paws have

length of about

Islands in

and

ears,

to three-quarters the length of the

are short,

1

\

*

are long, thin creatures with a pointed face,

The smallest

been released n other areas of the world,

species have

Mongoose?

a

Mongooses

They are

mammals

Is

nso goose (R melleri)

and

live in

Some

species are

large family groups.

1

’s

an mongoose

(L

Mongoose

fur

long, coarse,

is

and generally brindled

xuhni)

or grizzled (where the individual hairs are colored

Mungoj

Ga >P an mongoose mongoose iM mungo) species

.

XVieumu

l

ParacynKtn Surxitj

speces spe<

.oivts

re-tailed

\vt

v.eoes

es Selojs mongoose

(/

r

sgi a ned

back,

{P selous^

mongoose (G species

..sevies

mongoose (G

legs, feet,

elegans.)

mongoose (G

and

tail

or

the rest of the body.

tasciata), giant-striped

species, such as the

stripes that run

No mongooses have

tail tip

Some

may be

their

spots, but

a different color from

species have different color

forms depending on where they

live.

For

example the

gnanddierf)

wow striped mongoose (M

brown mongoose

(S

slender

decemlmeaTa)

m

concolor)

mongoose

soil

mss Meerkai

1:100,

southern Africa is

Mongoose Dwarl

less

1:106.

it

is

red,

found Such variations

color of the area

making the animals hr

in

areas a black form

match the

SEE ALS

usually gray to yellowish brown, but

is

the Kalahari Desert

some

98

A few

and two Madagascan species have

the length of their body

6 speces

op need

along their length)

meerkat and banded mongoose, have bands across

albtcauda)

"leerLal iS suncaTta)

Galfdxtn

MungotKTn

banded differently

mongoose

MALAGASY MONGOOSES G+hdt*

garnbianus)]

and

in

usually

and may be important

obvious to predators.

Mongoose, Banded 1:110

in


THE

Some mongoose

mongoose— Kenyan

1

„

nngtailed

subspecies, sniffing the air in a

mongoose

(5):

Selous'

white- tailed

(1);

(4);

mongoose

mongoose with an egg

mongoose

dwarf mongoose adult

(3);

feeding a beetle to a juvenile

mongoose

FAN' LY

species: bushy-tailed

~ongoose posture

typical

MONGOOSE

narrow-striped

Egyptian

(6);

marsh mongoose

(7);

Mongooses have

(8).

phenomenal

a

sense of smell, and they mark their territories

with scent. Social species

mark each other help recognize

same way

the

in

members

to

of the group.

Scents are produced both by cheek and

communicate the

anal glands. Scents will

individual identity of the marker, as well as its

sex

and breeding condition. A

mongoose's eyesight

also generally good;

is

and unlike most mammals, which only have partial color vision,

it is

able to see color.

Although mongooses are carnivores, they

will also

fruit.

eat vegetable matter, such as

The canine and

carnassiai teeth (those

adapted

for shearing flesh) are relatively small for a carnivorous

animal, probably

owing to the number of invertebrates

and small animals unnecessary. The

and

is

generally

in

the

diet,

mongoose

which make carnassials

skull

more robust than

has a long

facial section

determine where the animals

a genet skull.

where the

Where Mongooses The mongoose family solitary species.

The

mounds

Live is

split

between group-living and

social species also

smaller ones, such as the dwarf

They use safety

in

numbers

tend to be the

mongoose and meerkat.

soil is

good

mounds. Most species are

are semiaquatic or tree dwelling.

the day, while solitary species are usuai'y nocturnal, using

and

the cover of night as a defense.

chewing wild melons and digging up

general, solitary species tend to inhabit

group-living species,

den

sites rather

species. In

wooded and

forested areas, while social species prefer

open

plains. For

than habitat

Mongooses

but are

They may also eat eggs

small insects to full-size cobras.

between

terrestrial,

opportunist feeders and take a wide variety of prey from

as protection against

habitat varies greatly

The

may depend on the

predators. Group-living species tend to be active during

Mongoose

African grasslands,

alternative to burrows.

presence of mongooses here

some

On

hard and unsuitable for digging, termite

provide a

availability of

live.

may

fruit.

Species living

communal

in

dry areas can obtain water by

species, such as the

meerkat, forage

in

The

smaller,

dwarf mongoose and

groups and tend to feed mainly on

invertebrates, lizards,

are usually solitary

roots.

and small rodents. Larger species

and feed mainly on

larger prey.

99


SMAJ.L CARNIVORES

Suricata suricatta

A. meerkat sentry

a familiar sight on the southern

is

African savanna, demonstrating the selfless cooperative behavior of this plucky

and

mongoose

toward members of its own pack.

Common name

Meerkat

tailed

Scientific

name

Family

(suricate,

gray meerkat, slender-

Meerkats are the most sociable

mongoose

meerkat)

family.

breeding pair

Herpestidae

may be

may dominate

Order

Carnivora

Size

Length head/body: 12-18 length 6-12

Weight Key features

3-5

3

in

in

(30-45 cm);

in

(15-30 cm); height

in

kg)

Slender, short-legged animal; tan to gray

Social lives in colonies of up to 30, but usually 10-15, animals; sentries posted to

watch Breeding

Two of

for predators while colony

to 5

young born

is

foraging

in

rarely aggressive

pack

in

baby-sitting, tunnel maintenance,

their

meerkat

group members take part

other. All

the various tasks involved

duty.

male and

life,

such as

and sentry

Both females and males without young of

own

will

help guard and provide food for

other meerkats' young.

produce milk even pregnant and

will

if

Some females

will

they have not been

suckle other females' babies.

after gestation period

years

at

Weaned at 9-10 weeks; sexually about 12 months. May live 13 up to 10

in captivity,

in

the wild, but

more commonly 6 Voice

A

Diet

Insects, scorpions,

trills,

growls, and barks

and grubs; occasionally

small snakes, birds,

and mice

Africa,

in

Angola, Namibia, South

some

areas

but most

warmer, wetter months. to take adult food

when

they are about three to four weeks old. Toward

reluctant to share food with each other, they will

Population abundant Not threatened, but fallen in

Young meerkats begin

year,

the rest of the group. While adult meerkats are

and southern Botswana

numbers have

Meerkats breed throughout the

the end of weaning the young begin to follow

Dry savanna, open plains, and scrubland

Distribution Southern Africa

Juvenile Followers

births occur during the

variety of chirrups,

lizards,

Status

often give food to juvenile

group, even

if

will

particular adult

and follow

it.

associate

The adult

quality prey to

its

When

will

follower,

fed, but will also learn

meerkat.

members

they are not their

Each youngster

food from

how

it

own

itself

of the

offspring.

with a

around, begging

pass on

who to

will

its

best-

not only be

behave as a

the young start following, the

juvenile meerkats

from

earlier litters

confused between begging

for

and

arrivals:

giving food to the

new

may

get

food from adults Adolescent

meerkats have been seen to beg food from an adult only to pass litter,

100

may

75 days.

mature

Habitat

been observed

and group members are

toward each

with broken brown bands on back and sides; black eye rings, ears, and tail tip

Habits

packs,

larger than the

at

(10 cm)

lb (1.5-2. 3

packs, which

him. However, no distinct

hierarchies have tail

live in

of the

include several breeding pairs. The female of a

Suricata suricatta

shoulder: 4

They

members

SEE ALSO

-varf

1:106,

it

onto young from the next

or to give food to a youngster, then steal

Mongoose, Banded 1:110

species


MONGOOSES MEERKAT

it

back again. Meerkat

life is

complicated, but

also highly social.

Meerkats have strong, muscular forelimbs with large claws, which

make them

excellent

diggers. Their long, slender, short-legged bodies

underground

are perfect for traveling inside

tunnels. Although they are quite capable of

digging their

own burrows

(or warrens),

meerkats often share dens with African ground

mongooses.

squirrels or yellow

such cases the

In

dens are usually dug by the ground

and the meerkats move

squirrels,

Neither species

in later.

appears to object to the presence of the other

—

in fart,

the meerkats ignore the ground

squirrels altogether.

Underground Refuge Meerkat warrens

an area about

typically cover

16 feet (5 m) across and, depending on ground conditions,

may have up

to three levels, with

the deepest tunnels about 10 feet (3 m) below

may have up

the surface. The largest warrens

90 entrances, but about

more

usual. Within the

1

to

5 entrance holes are

warren a

series of

tunnels connects chambers of about 12 inches

(30 cm) across. Meerkats usually stay within

easy reach of a warren, which

their

is

main

refuge from predators. The temperature

deeper tunnels may vary by only

a

in

degree or

two, making the warrens important shelters

from the extreme desert temperatures above ground. Meerkats spend nighttime within their

warrens

—the coldest

the hottest parts of inside too.

The

parts of winter days

summer

territory of a

be up to 6 square miles contain five or

six

(1

and

days are spent

meerkat group may

5 sq. km) and will

warrens, occupied

in

rotation.

Meerkats spend from a few days to several

months around one warren, moving on when the surrounding food supply has been used up.

A meerkat territory.

(ÂŁ

pack

Marking of

is

very protective of

its

territories using scent

A male meerkat on guard duty

in

southern Africa. Sentries are posted

from

the Kalahari Desert,

in

high places to

watch out for predators while the group

is

foraging.

101


rc

I c

Leash

rh

v.

e

M

een.ers

m.—

3*r an venomous

w»il

v font

oorena source

r

ner

oe~ sres

roe

rr

—erne*

mongoose

—iiv

~=i

njmans ire _ tee'kars

-owe>e

-

is

~s~

e~raz«

ne

*

arcfer rr

us ..a

rrne *o foraging oe'rra etaors o r

tnrtD

a~?

rnan

rne razK

noaxiaptior o*

skil

rher

tv'iai'-

a so oartcutotk

r.?>

Kers raKe a 0

’etummg rr

numa~ nano

m

~ e

mre;

re—no

" ee*e*

-ag

x

*tf >o.-c :,•

r

-XH.r

n»rwc

oir

~g

**

_ ce: ~rr~ ~g

noozmn-.

;

>5

anmais such

px

ooirtt in

can Lore ne hpn s ^01

treed rnp rcscus

xi

5 *2^

tar~g*o*

rrooe -

arc

gnucs

are^cc

ra*.e

xmg

tfrat

*'

~ais me* for—

oar arouna without a

i~.ar« ,nr esaitoisihBd

rr cr<ig out

mer ~er

me

“xeme'

*0 eaie mg'?-

~»*errc

x rs

pre

me

~ a» a tar

croup

pao*. cffKr

~ae

'

groups and

— lXi ir,a~*

~g>r omr-

a*

—a is

Ziti;i

.o-rig 'e1 ae:-

rer

mfetar 3ra:»f

n«^

eguTij

•i

sdaatsied

*02

:

asr

e~'aie5

gox

SEE ALSO

1

armro-igT'

<

**> **0 110m

a icmg

"na»

: -

<

*•:

an

2.

*•

n* tiippn;

rer

mpi

npr

nrr

Df>n;

ax ?

—ca

mint 1'i'nr

nr

a*? n./n?

dm r

n#f, c*e*

oaev-stneri

rr

JUrtriziupt

— *r* aes

—a—

pan

rn ~immj\n dopn; f

oare^

•v*~ rarreerr :

*esidemt

racks

'a“"ag5.

a,rc 'e~ae ~egnca5.

“rr— ir*c ra:«

"ense'ies as

rr -esoabishrig

sememes

-"rr*

r ar

w«i

wmr a group

mwkaes ~riri; meexm

treedre an mas

*e-

o»e*

1

oai*e r*g

-O’

akamoe

p-.t.n: souirmlk

H’ltma

r*H

mem k

mmmxra

«-tnppmpT

"nice Like

03 ge

anc 00 not

food ~ne g’our

s soal rnae in

it

ne

wil accept

ne e~on

me

hatf an is

no

ekceptions,

izan out o4

mav taka

since

o^mg

o’oa’ to

ae soma

is

t a maerkart gats

gecko n

unroxe T the gecko

*

pa animats "here

oj"o\^ Arnougn

rr 'm:ro«g ~~er

ro

o*av

-rm: soe

~g osrc ~ag5 anr ra«.ng :r. anc ner *e— site. er re

~2cec .e*

own

sea"*, o* a targe

n?» rare the

and

oaa''kats *aar as a

tot

another anima to dig

roasrc 0

o>.r.

racks separate iow~*ankmg

or*

I

hunting together

,a'

noweve’

out pra\ anr ro noid

as sands

r ookng

rne maerkats

rr

o jsruia?

proramon against pnaoarors out

rrrvioas

down

do

~ongrosas

rur so’ead

ronoa-are

ng ~wc :r?- ©o Oi ne ~g car* a«Geor

ave'\

anane rnar *ood

a^ea rr

are _:sar rr

nreer roes nor

opre

~e~Or'"

-.1

goug

os appioachsa r a

~e ~esras3ng

rare ne> and a*p=5r

i

manna*

osro-rap?

ne no

o>

5 sperrer ?'".t ’n:

ran

- wil ~rr.

— er

a'-

so.rres re ne laoensnar ~na r.av\er

othaT sor.a 5 pe*e‘al'«

- a-a"r 'o ragng orte

”nav Lsuahv alow about a weak re'ne

ia-pa-

panes

~ oe's

-

5.s*er«

roni a —

**>e er-e

ne>’

^i:r

sm-orans anr can withstand sengs

oe_ o

with inre

e

fond

*

v-^eeio,

T

nolens ~g annais ~o~

kil

"ooed to

reza~ne ess savee anr sne was *jlu area or an

Vieaitkars a-e

v-\as

p?-t?n n?vg 5

o~.e"

~mp -anoucr

rrornje nev oie *o severa

nd

genre o*

5 ne

e raov-sore'

in

speed anr

’>

erraaisner:

:

-

Wente-s

jn..ng rreaarrrs

o*

a~e* 'e'STranre up gna<

*

r

or

j~ri

anr rne one *?nae was

ra»<e dirlr.

rak?^ on as

?.s

one '?~aie tolowed

r>-g

:ar4v ra* -^c w'eeo

_ 'a ,ae

snakeis

na~ na-fseues

.ape*

a? arzepre:

r -ne rr

me heir of

its

hour to

worthwhile


MONGOOSES

since geckoes are ’-’e€ riats <v

7>e

smce each

»

ve

coace''a:e

rr

- a

by a

t s

ca

s.>:~c

rs.

—.me

a

t\>

<sy an attacker

.v

c'ecatm

dher

ng grout

oesccttec a

ar

seni»y

gnes an alarm

me pack

spots a predator the

t

call

that ta

1

s

the rest of

to run for cower Veerkats ca~-ct

r-e

a

-v

oe ava

foed ng St farag -g - a grout a

a.s

can

-

nd mare food and

teach e r than an —a

s

ace

—eerkat oack

for

forag ng a cne.

member o'

a meerkat group

e;<

coses

stroke

t to

3emg

as

danger

the group but the a

-s

the ns*

-vei

as forac

ts position

or

a female

tac:<

.'.as

see'

-

az-c n

ng

-

a 'group

and havng

mongoose neenatis a

attacks

ndiv 'duals

turns ana sra-e

Meerkats that rema n beh nd bafcy-

aga nst

—as mat

as sra<es

arc

are

so as

n

me'

mer ta

backs ~a< tg the' "a

grcw-'ing

and

spittirg

ta< a

so protecting teeth

me

s

.area

r

stand

ly

or-

a'cr ng.

end and

<e a cat "the meerkats

oo not work and

defense positian

iVtth

mate themselves oax mare

by *aisng

predator, t wi

predators ana 'eat

s t are

rva

~a<a ~ :c

mom a

a

observed

as

defenses Lie the banded

tactics

.vn

,v

encounter with predators or other meerkat

a sentry s a cc cere on ze r a- or 1

others

,

me srace rather mar -epirg -er compen ors -~e r me oac< -ad chased t~ me otruders they turned on me die fomale—seeas a

oc<s or tushes a so

r

t

a hght

mpasmg

cable feed ng time

o

-c ae~:s

—such

By perform -g guard out- a sentry meerkat s

gn ports such

even

cctemta threats

te

Sentry Duty

-

*eic

mat during

gangs

sacrificing -a

me

sentnes meerkats na<e other antipredator

~cvdua ,v

of

-fewevep such cens/tm s not tolerated n a

sa-e time Therefore are mare

esst~e

tasks, t

cu- shment for ~e r se-'st tens /c*

meerkat scenes ae* ng cut "or

oredators, are

me

irrgJn

concentrate on -orag ng ana -vatdn for oredators at the

not ea.e to 'set

rr

coulc encourage ar nd vidua to take art- antage

testae n - g- olaaesto ieec n atari ov-gr the forgg

oe'

meerkat team perform such 'se~ ess'

fo-mer mprc'<ed smce guards are

Sc'et- s

-

the

at

:

stca

grauc a* meerkats than by a

-omc

themseses Since nary "erne's of

this atuatiar

graua reduce the danger a-

one meerkat ceng caught

smce

~

guaranteed a cccc -ea

s

Farag ng ar

i

'c and

-

'EE*

t

is

threatened by a

up the dassc mongoose

t -v

-'

mm.', tse or

sensitive

nape of the

and daws faeng

me

ts

bao

~et>

predator Tne

-

-eertat "*eoeh'

*~ as .

i

racy -shlB' i

kr

Z'.rg ar -ail. Weerirats

sra

re -cm soca/

of

re

noncocses arc -enters take turns

r re

csdt

«.

cercm ~g

group duties sum

as

catiy-sittrg arc tur *e

ma

'

‘era *c=


Cus

-

o>/e r

me

*"

:ai

arc "eaC An attack from the sky

Dev‘v

by ar eac e or for cover

away

*

r

f a

om

stare 'g up rrav be placed

*airs

hawk causes

pack

s

the rree r kats to run

attacked

the open,

:n

their warren, the adults will

memseives over the

juvenile

members

throw of the

oack to protect them. Like banded mongooses, *

member

a

w

of the oack

captured the others

is

launch a rescue attempt.

Nursing Care *

a

memoe

o T the pack

left

is

may be nursed back

*

a~ack.

1

njured after an to health by the

othe r me'~'oe rs. A femaie meerkat that had

peer caught and njured by an eagle was

oemg surrounded

observed

by ner pack and

-e'oed back to the warren The female had

produced a

and was

tter of kittens

eft with

them

the burrow while the

pack went to forage She was fed

"est of the

-v’n grubs by the other

and was

m

a few days earlier

memoers

of her pack

also helped to stand upright during

me mommg

sunbathing session Thanks to the

s^ooort of the group both the female and her

"er survived invalid care

is

only performed for

adui* meerkats since the loss of a baby

damagmg

to the

group than that of an

J neerlrar grouo basks n Vfe*?nrjts ree<; *c

if

me

me ^aming

sunoame earn iay

is

less

adult.

sunshine.

keec memse/ves

to

correct ootfy temperature.

Sun Worshipping Along

with underground dens and group

nesting, sunbathing plays an important part in

*eeomg meerkats

When mey

a: their correct

leave the burrow,

body temperature

members

sit

or stand

an upngh * position with their stomachs facing the sur

They may then

backs or spread

ie

down and bask on

th emselves over rocks to

their

absorb

heat Dur ng the hottest part of the day, however.

104

SEE ALSO

1 2: ?

in



SK'AA.

CARV.CRES

Dwarf

Mongoose The

tiny'dwarf mongoose

and

scurries

around

He legale par.ula

resembles a short- legged cat

groups seeking small prev.

in

specializes in eating the large insects

invertebrates that are so

MONGOOSES

Z

tne

mongoose

- :S

abundant

wh ch

.

"ore vulnerable

size aiso

i

to predators

than the they

groups Living n a group means

pa

- 4

sma

means they

wry

speoes arc s ore reason

"3

r

a e

3'ge r

m

ive

the re

are

one mongoose

s

r

pac< The dominant

w

mae

numoer

He. a'org with a

watch

msehes or a

the

log.

Dwar* mcrgccses

are

cm rent rc

themsefres against predators. They are

mow’'

o'.er

to srte tretr

near pee or *aso

the

vg ant

extremely

s

^crgooses sun

many

ecuced by the

to keec

dwarf

acect a:

presence of cither. Uke the ^eer<at. dwarf

mongooses oost guards

pair of

'arge

of eyes to watch for oanger and the nsk

to ar_,

of Africa.

a~-e

arger scenes a r e unable to do

However the r sma

in parts

i

*r

allows tre^ to enter the tunnels o* ’'Oder's,

and other

SMALLEST rr’e ''"Cer5 of

i

’a~'

It

aers

n ests.

which outs off potential attackers.

of immature ma:es,

often be seen keeping watch over the pack

from a hign vantage point. such as the top of a b<g rock or high

m

a bush

Pack Hierarchy Dwarf mongoose packs have a

stnct hierarchy

with only one dominant breeding

The

-

pai'

dominant femae or "matriarch" leads the pack,

and only she A. though

rs

ao<e to produce

other fema es

m

young

the pack

may mate,

they ether do not apcear to

become pregnant,

or ther fetuses abort,

rf

do

young are born oead.

In

suoordmate females

hep

stii

they

such cases the

produce milk and win

suckle the matnarch's

members

are involved

ang caring

for the

give birth, ther

young

AJ

m grooming,

young when the

!

oack

baby-srtting. rest of

group s out foraging After the dominant

wh ch

are usually the oldest

pack, the youngest

highest

rank

m

ran*

in

members

members

the pair,

of the

are the next

any age group the females

more ngnry than the males '

106

SEE ALSO

1


MONGOOSES DWARF MONGOOSE

Group Hunting

dwarf mongooses can spend more time looking

Dwarf mongooses also forage as

a group.

Because they hunt prey by chasing and

pouncing and not by

stalking, they

individual

is

do not

not hampered by the presence

of others. Foraging

in

a

group also provides

protection against predators. The dwarf

mongoose has developed to keep

wide

a

group movements, and warn against danger.

Dwarf mongooses have taken group feeding a step further than

for the

many

other species,

is

When

there are a

source, a dwarf of

its

lot

hornbills' presence.

of hornbills at the food

own members on is

the hornbills arrive

in

guard at

is

air

vents,

of hornbill, such as the yellow-billed, red-billed,

rise

among

The

birds feed

the packs on insects disturbed

by foraging mongooses and

keep watch for predators.

The

birds

are particularly effective at

spotting attacks

from the

When

air.

they see a

predator, they fly

up into the

mongooses

is

calls

and

trees. Their actions alert

to the danger so they can

their escape.

predator

make warning

The

hornbills

wT

too small to be a

the

make

even cat when a

risk

to themselves.

ch additional curve lance means that the

their den, they

and the mongooses

appear almost immediately.

hornbills.

If

the morning before the

often forming partnerships with several species

and von der Decken's

for

mutually beneficial:

mongooses have emerged from the

all.

some competition

food, the arrangement

down

get

mongoose pack may have none

Although there

call

for

particularly important

mongoose guards who might not

enough food without the

variety of calls

contact with others, coordinate

in

and eating rather than watching

danger. The relationship

require stealth, so the foraging success of any

one

for food

If

the

mongooses

before the hornbills arrive, they

will

the hornbills before they start foraging.

wait for


SMALL CARNIVORES

Indian Gray

Mongoose

Herpes tes edwardsii

Mongooses have a reputation as snake

Common name

killers that is well deserved, but the Indian gray mongoose, like most predators, generally prefers less

mongoose

common

Family

Order

name

mongoose, common Indian gray mongoose,

(Indian

mongoose,

Scientific

dangerous

Indian gray

The Indian gray mongoose

common

Although

Bengal mongoose)

ground,

Herpes tes edwardsii

Length head/body: 16-18

16-18

in

(40-45 cm);

in

(40-45 cm); height

shoulder: up to 8

in

Weight 2.2-4 Key features Gray

tail

Unlike

good

2-4 young born

nest.

mammals,

at

about

at 2 years.

8-10

in

threatened, she

1

May

another

live

the wild

birds,

and

itself.

species, the Indian gray

and males and

year—the

or thin

West

woodland

islands of

them

alone.

will carry

them

her

in

mouth

Newborn mongooses have

hair,

to

a fine

but they are blind for several

to catch their

mongooses

own

trips

for a

few

where she teaches

food. Although

are instinctive foragers

and the the mother's role

Japan

A common

young

kittens are born in a well-hidden

mother on hunting

their

Indies, Hawaii,

Population abundant

their

weeks but then weans them onto adult food. Once weaned, the young mongooses follow

some

and eggs

Ryukyu and Tonaki Status

digs for

The mother suckles her young

days.

lizards;

site.

covering of

Distribution From eastern and central Arabia to Nepal, India, and Sri Lanka. Introduced to Malaysia. Mauritius, the

an old

the mother feels her young are

If

after gestation

Weaned

in captivity,

Open country

it

occurs at any time of

mothers to care fo r

Mongoose

insects, snakes,

Habitat

den that

climber; hunts during

Angry chatters and chirrups Small

a

in

in

males search for other females, leaving the

period of 60 days.

Diet

active during

is

females only form pairs for breeding. After

5 lb (1-2 kg)

month, sexually mature

Voice

It

a solitary animal,

is

mating— which

Litters of

or

some mongoose

to light-brown coat, finely speckled

Solitary; agile;

20 years

mound

mongoose

(20 cm)

the day

Breeding

walls.

at

with black

Habits

time on the

its

also a skillful climber, able to

is

the day and spends the night either

Carnivora

length:

an agile animal.

spends most of

ascend trees and scale

Herpestidae

termite

II

it

it

is

skills

to her

in

young

is

passing on her hunting vitally

animal

future survival.

and hunters,

important for their

Once the young mongooses

are

able to hunt for themselves, they will leave their

mother

to find a territory of their

own.

Stealth Tactics

mongooses

Indian

some

are

more

active killers than

related species, such as the

meerkat or

dwarf and banded mongooses, which mainly forage for small invertebrate prey. The Indian

gray

mongoose

takes a higher proportion of

vertebrate prey, such as

rats, birds,

and

Hunting prey requires a high degree of stealth,

108

SEE ALSO Meerkat 1:100; Mongoose, Dw^rf

1:

1

06.

and could be a reason why

Mongoose, Banded 1:110

this

lizards. skill

and

species

prey.


MONGOOSES

is

solitary.

group would disturb

Traveling as a

larger prey

and hence reduce hunting success.

the ground and turning over rocks and stones. is

killed

the neck or head.

by a bite to the back of

Mongooses

fond of eggs. They break the the egg

in their

between

their

front

back legs

of the

The mongooses'

such as mice and

at a rock or wall.

mongoose genus

their snake-killing ability.

immune

sensitive to size.

it

suffer

receive them.

known

mammals

from snakebites a duel

will

less

of a similar

why mongooses

is

between

mongoose the mongoose strikes

and

their diet

ability to

rats

kill

kill

dangerous

pest species

make them popular

species, including the Indian gray,

mongoose

for

venom, they are

to snake

than other

In

in

killing

pets.

have

help reduce snake and rat numbers. But the

Herpestes,

Although mongooses

However, the main reason

do not

However,

attack.

been introduced to areas around the world to

such as the Indian gray, are well

are not

mongoose

snakes and their tendency to

it

Killers

Members

may be overcome by

although mongooses are capable of

Some

Snake

down

since there are easier things to catch.

by holding

paws and throwing

be worn

eating snakes, they rarely feature

are particularly

shells

will

MONGOOSE

by the mongoose's stamina. Even constrictor

the speed of a

also forages for insects by sniffing

Fleeing prey

snake

Eventually, the

species such as pythons

prey, the Indian gray

As well as vertebrate

mongoose

agility.

INDIAN GRAY

that they rarely a snake

and

a

avoid the snake's

by using a combination of speed and

whatever

is

it

an opportunist feeder, catching

can

been introduced

find. In it

many

areas

where

it

has

has attacked the local fauna,

often with disastrous consequences.

A mongoose

Its

fondness for eggs has also contributed to the decline of

some

the Caribbean islands

numbers

and

birds,

its

introduction to

may have reduced

of green turtles,

whose eggs

on sandy beaches. Caribbean ground useful

in

keeping

down

the

are laid lizards,

insect pests,

have

also declined since the introduction

of

mongooses.

killing a

Uttar Pradesh,

cobra

in

India.

Mongooses are

renowned snake

killers.

The author Rudyard Kipling wrote about a

duel between a

mongoose named tikki-tavi

and

Rikki-

a cobra in

The Jungle Book.


SMALL CARNIVORES

Banded

Mongoose Banded mongooses

Scientific

thcii large den, often

name Mungos mungo Herpestidae

Order

Carnivora

The banded mongoose

Length head/body: 12-18 tan length:

6-12

shoulder: 6

I.

in

groups— in

(30-45 cm);

mongoose

(15-30 cm); height at

in

(15 cm)

in

Weight 3-4.5

(1-2 kg). Male slightly heavier

lb

brown

to black; black-tipped

tail

Up

mature

at

10-12 months. May

up to about 12 years

in captivity,

5-10

in

up

in

Mainly invertebrates, particularly termites and beetle grubs in open grassland and various

mongoose

more

such as beetles and

forested areas;

some

vertebrates such as lizards, snakes,

eggs and some

small

and mice;

leaf

poking

litter,

Range of habitats from forested or cultivated areas to dry scrub and open grassland

Distribution Africa south of Sahara Desert, but not

mongooses smash

beetles,

dung

catapulting rocks.

An

them between

individual

when

in

the

make such

it

is

not inclined to

finds a

food supply, a mongoose

excited twitters that

it

have attracted the attention of others, then

try to

up to 0.5 square miles

does happen,

between two

it

(1.3 sq. km).

own

territories,

found

in

territories,

usually

mongoose dens

are

termite mounds.

A den on

times by two different packs. at the

larger

Mongoose, Dwarf 1:106

same den on

pack

it

and both packs

boundary may occasionally be used

up

When

tends to be at the boundary

run away. Most banded

1:100,

who

grab a share.

so meetings between packs are rare.

/

soon

will

Each group of banded mongooses uses a territory of

risk

large eggs by

their hind legs at

mongoose

share food. However,

will

and

balls, snails,

Packs usually stay within their

Mm

by a

led

and turning over stones and dung

search of prey. Banded

southwestern Africa

Population abundant Not at

group

noses and front claws

their

particularly interesting

fruit

River Basin or

as a

tough objects such as large millipedes and

and churrs

leaf litter invertebrates

SEE ALSO

the

senior female. They busy themselves scratching

the

Diet

110

in

individual characteristics.

into crevices,

Chirps, twitters,

Status

with dwarf

live

Voice

Congo

Banded mongoose

pair.

Banded mongooses forage

wild

Habitat

has three or

Busy Foragers

4 young born after gestation period of 2 months. Weaned at about 1 month;

snails in

typically

packs of up to 40 members, but

between

to

sexually

pack

lives in

1 5 and 20; each pack has a dominant male and 3 or 4 breeding females

Breeding

A

the whole

in

females. Instead, rank appears to be based on

age and Active only during the day; social: erritorial

family.

social hierarchy, as

back;

usually

the biggest

females do not necessarily outrank males

usually brownish-gray, but color varies with habitat; dark-brown bands across

Habits

large social

lives in

than a single breeding

Key features Coat

feet are dark

fact,

an old termite mound.

four breeding females and several males, rather

than female

.

They return at night

prey.

Banded mongoose

Family

Size

busy gangs that scurry

live in

around looking for small

Common name

Mungos mungo

will usually

the

If

a territory at different

two packs

same

turn

night, the

chase off the others.

to


MONGOOSES BANDED MONGOOSE

Mongoose packs have territory

several

and may use one

in

months

the group

their

for anything

single night to a couple of

The breeding

dens

from

at a time.

all

'

at three to four

of the pack

weeks

cycles of females within a

pack are synchronized so that

den

a

the young are

out foraging. The young leave the

is

old,

in play.

weeks

old

Once they

the kittens

and are

join the rest

about

five

begin to accompany

will

the adults on foraging expeditions.

born within a few days of each other. This reduces the

risk

to any one. mongoose kitten of

being caught by a predator:

If

births

were

spread out over a longer period of time,

find refuge in a termite

newborn mongooses one by one. Another

a

may be one

of

number of dens

scattered over the pack's territory

the

need

and used when arises.

for predators to pick off the

advantage of simultaneous

births

that

is

all

for cover

the

reproductive females are producing milk at the

same

time,

and the

any female, not

kittens

may be

just their mother.

suckled by

Hence

mothers are free to go and look for food turn while the

females. Other involved

in

in

of the pack are also

caring for the young.

An

mongoose

—whose is

to run

— banded mongooses may club mobbing

together and take part

in

During such attacks

members

all

bunch

closely together

heads

raised.

The

attacks.

of the pack

and advance with

resulting growling

their

and

snapping mass can give the impression of a

young are nursed by other

members

Numbers

only response to potential predators

it

would be easy

It

in

Unlike the smaller dwarf

Š Banded mongooses mound.

Safety

adult male

usually remains as a baby-sitter while the rest of

single large animal.

The displays can be so

impressive that they intimidate large dogs and

other predators such as jackals, eagles, and vultures. But

banded mongooses may

also

mob

harmless animals, such as bushbucksand geese, if

they seem to pose a threat.

on the defensive, in

it

When

a

pack

is

bunches up with the young

the center. Protection of fellow group

members may even extend individuals. For

example,

to rescuing captured

when

mongoose was caught by an into a tree to

the pack

a

banded

eagle and taken

be eaten, a dominant male from

was observed

to climb the tree

force the eagle to release

its

catch.

and


SMALL CARNIVORES

of Species

List

The following

lists all

Gulo

species of

M.

G. gulo Wolverine; circumpolar, in

small carnivores, including their

N.

distribution

America and Eurasia

(Part)

I.

L.

SUBFAMILY PROCYONINAE

Bassariscus

B astutus

Lutrogale

L

polecat); semiarid regions of Africa

islands; N. Africa

nngtailed

cat);

W.

U.S.

from

-Oregon and Colorado south and throughout Mexico B sumichrasti Cacomistle; C America

Nasua

N

nanca White-nosed

coati; S.E.

Arizona, Mexico, C. America,

W. Colombia, and Ecuador

N

Nasuella

Mountain coati, Ecuador, Venezuela, and Colombia Procyon P cancrivorus Crab-eating raccoon; olivacea

W

Costa Rica to N Argentina P gloveralleni Barbados raccoon; Barbados P msularis Tres Marias raccoon; Maria Madre Island, Mexico

P

Common raccoon; Canada. US, C America;

lotor

S

introduced

in

parts of Europe

M. foina Stone marten (beech or house marten); S. and C. Europe

SUBFAMILY POTOSINAE

Bassaricyon Olmgos

B alien/ Amazonia B beddardi Guyana B gabbi C America and N South America B lasius Costa Rica B pauli Panama Potos P flavus Kmkajou, E Central and S America from S Mexico to Brazil

W

SUBFAMILY AILURINAE

Ailurus fulgens Red panda, Himalayas to

China

P.

to N. Nigeria

V.

Eira

E barbara Tayra; C and S. America, Trinidad

africana Tropical weasel;

c u/a Little gnson,

C and .

N

New

Zealand eversmannii Steppe polecat; steppes and semideserts of Russia, Kazakhstan, and Mongolia to China

M M

felipei Colombian weasel; highlands of Colombia

M

(huron).

C and

S America from Mexico to Brazil

capensis

and

S.

China,

clawless otter; 1

of the

Congo Sea

Islands

SUBFAMILY ELLIVORINAE

Mellivora M. capensis Honey badger; Africa from Cape to Morocco in west and Ethiopia, Sudan, and Somalia in east;

Basin

otter; Kurile

and

Gulf of Alaska, reintroduced into parts of former range along Pacific

SUBFAMILY TAXIDIINAE

Taxidea

S.W Canada and

Sumatra

L

felma Marine otter; coast and

Conepatus Hog-nosed skunks C. c hinga Andes skunk, Argentina,

and Peru

longicaudis Neotropical river

humboldtn Patagonian skunk; S.

C.

Chile, Argentina

leuconotus Eastern hog-nosed skunk, E. Texas, E Mexico

and S. America from Mexico to Argentina provocax Southern river otter;

C mesoleucus Western hog-nosed

Argentina, Chile

C

lutra

skunk; S U.S semistriatus

S

Lutra

European

river otter

(Eurasian river otter), Eurasia south of tundra line,

reintroduced

Paraguay, Peru

Bolivia, Chile, C.

otter, C.

L

N.C. U.S. south

Mexico

SUBFAMILY MEPHITINAE

canadensis North American river otter (Canadian otter); Canada, U S including Alaska coastal islands of Chile

L

to

and Russia

Lontra

confined to a few declining populations in E Europe and Spain lutreolma Indonesian mountain weasel, high altitudes of Java and

taxus American badger; from

T.

coast of N. America (notably

L

Arabia to Turkmenistan,

Nepal, and India

Aleutian Islands, Alaskan coast,

L.

Myanmar

M. marchei Palawan stink badger; Palawan and Busuanga N.E. of Borneo

En hydra E. lutris

badgers

5째N from Senegal

absent only from desert regions of Namibia A congicus Congo clawless otter; African forest; streams and rivers

European mink,

mgripes Black-footed ferret, western prairies of N America; was rated extinct in the wild, but maintained in captivity and being

Cape

to Ethiopia south to the Cape;

California)

kathiah Yellow-bellied weasel;

China

Africa south of

L

112

Lanka, S.E. Asia,

Aonyx

A

N America from about 50째 north to Panama extending through northern S America along Andes

locally

Ferret

javanensis Teledu (Malaysian or Indonesian stink badger); Borneo, Sumatra, Java, and N. Natuna

Indonesia, Borneo, Palawan

frenata Long-tailed weasel;

lutreola

Melogale

M. everetti Everett's ferret badger; Borneo M. moschata Chinese ferret badger; China, Taiwan, Assam, Myanmar, and S.E. Asia M. orientalis Oriental ferret badger; Java, Bali, and S.E. Asia M. personata Indian ferret badger;

A. cinereus Short-clawed otter;

Africa; introduced to

W

Metes M. metes European badger (Eurasian badger); N. Europe up to S. Scandinavia, European Russia up to Arctic Circle south to Palestine east to Iran, Tibet, and S. China

Amblonyx

Islands, S.

in

China and

Indochina to Thailand, and

My da us

India, Sri

S.

Sumatra

India, Nepal,

not semideserts of Kazakhstan and Mongolia, Mediterranean region, or

Hog badger; Peking

A. collaris

SUBFAMILY LUTRINAE

M

S America

peregusna Marbled polecat;

Peru, Brazil altaica

M

Galictis

and Sudan

steppe and semidesert from S.E. Europe (Romania) east to W. China, Palestine, Baluchistan

N Myanmar

SUBFAMILY MUSTELINAE

banded

Poecilogale P albinucha African striped weasel; Africa south of Sahara

Japanese islands

Mountain weasel; forested mountains of Asia from Altai to Korea and Tibet M. erminea Stoat (ermine or shorttailed weasel); N America and Eurasia south to about 40째N, including Ireland and Japan, but

M

libyca North African

weasel; semidesert fringes of the

Vormela

zibellina Sable; N. Asia,

Himalayas,

Weasel Family

Gnson

(Sierra

(Appalachians)

M

naturalized throughout

Sahara from Morocco and Egypt

N America to California Nevada) and W. Virginia

SUBFAMILY MELINAE

Arctonyx north throughout

Poecilictis

Fisher (Pekan or

Virginian polecat); northern

M.

Siberia, Korea, China,

Europe, C. and E. Asia, and southern S. America

to Bolivia

FAMILY MUSTELIDAE

vittata

now

otter;

Borneo Pteronura P brasiliensis Giant otter; in all countries of S. America except possibly Chile, Argentina, and Uruguay

M. vison American mink (eastern

Japan, Korea

M

European

Smooth-coated

lower Indus, India, S.E. Asia, Myanmar, S.W. China, Malay Peninsula, Sumatra,

to Indochina

Nilgiri

W. Asia M. melampus Japanese marten;

M.

E.

Japan, and Taiwan M. strigidorsa Back-striped weasel; Nepal east through N. Myanmar

marten (yellow-

N. Europe,

M. pennanti

sibirica Kolinsky;

Russia to

perspicillata

Iraq (Tigris River),

mink); originally N. America, but

Mountains of S. India M. martes Pine marten; C. and

M

G

Scandinavia to Urals

M.

C. Asia

Nilgiri

throated marten);

E

Mexico

G

Denmark and

M. gwatkinsi

introduced in New Zealand M. nudipes Barefoot weasel; S.E. Asia, Sumatra, Borneo M. putorius European polecat; Europe except most of

Mustela and

P maynardi Nassau Island, Bahamas P minor Guadeloupe raccoon; Guadeloupe P pygmaeus Cozumel Island raccoon, Cozumel Island, Yucatan,

S.

marten; S.E Asia to Korea, Java, Sumatra, and Borneo

N.

Asia

C

M. americana American marten; northern N. America to Sierra Nevada and Rockies in Colorado and California M. flavigula Yellow-throated

to

nasua Ringtailed coati; S America east of Andes south to N Argentina and Uruguay

N

patagonicus Patagonian weasel; pampas of Argentina and Chile

and Egypt e$st

across Asia north of Himalayas;

-

Martes

Ringtail (civet, miner's, or

about 40째; Europe from

sumatrana Hairy-nosed otter; Sumatra, Java, Borneo, Thailand, Vietnam, Malaysia

including Azores, Mediterranean

Lyncodon

Raccoon Family

Arctic to

L.

striatus Zorilla (African or striped

t

south of Sahara

FAMILY PROCYONIDAE

weasel); N. America from

Atlantic seaboard (except Ireland),

Ictonyx

Order Carnivora

weasel (European

nivalis Least

comfnon

N

Africa

Spot-necked otter, Africa south of Sahara, absent from desert areas like Namibia maculicollis

,

Nicaragua

Amazonian skunk;

Mexico, N Peru,

E

Brazil

Mephitis macroura Hooded skunk,

M

S

M

W

U.S.

mephitis Striped skunk,

S Canada,

U

S.,

N Mexico


OF SPECIES

LIST

Peninsular Myanmar, Malaya, Sumatra, Borneo, Sipora and

Spilogale Sported skunks Western spotted skunk;

5. gracilis

W.

U.S. to C.

Mexico

S.

S.

Pagai Islands

Sumatra, Java; introduced to

W.

S.E.

W

Civetticcis C. tivetta

C.

FAMILY VIVERRIDAE

and

Genetta

Nandinia

G

palm civet twospotted palm civet from Gu nea including Fernando Poo island to S. Sudan in the north, to Mozambique, E. Zimbabwe, and C. Angola in south 1

:

(Mozambique genet, Hinton's Mozambique. S. Democratic Republic of Congo,

genet'; N. Angola,

N.W. Zambia,

Thailand, Malaysia, Indochina,

-toothed palm

Forest genet; southern part of

Sumatra, Java, Borneo, RiauLingga Archipelago, Bangka,

genet);

Cape

Afnca G. victoriae Giant genet (giant forest genet); Uganda, N. DRC Poiana

civet; India,

Nepal, Tibet, China north to

Taiwan, Flainan,

P.

Myanmar, Thailand, Malaya,

Paradoxurus

China, Malaya,

Sunda

Nusa Tenggara

Islands) as far east

as Timor, Philippines P.

jerdoni Jerdon's palm S. India

(Palm and

Java, P.

Indochina, Sumatra, Java, Borneo, (Lesser

civet;

Nilgiri Hills,

Sri

DRC

V.

civet,

Myanmar, Thailand, formerly

SUBFAMILY HEMIGAUNAE

S.

Chrotogale C. owstoni Owston's banded civet (Owston's palm civet); north of

the coastal distnct and W. Ghats of

V.

Indochinese peninsula

Sulawesi, Karlinata, Bangka, Buru,

Ambon and Langkawi

Islands,

Philippines

N. Vietnam Diplogale

V.

palm

— Mount

civet;

Borneo,

Dulit to

3,900

civet;

Myanmar,

Thailand, Indochina, Malaya,

ft

S.

China

Viverricula

Hemigalus derbyanus Banded palm

zibetha Large Indian N. India, Nepal,

(1,200 m) H.

civet,

Riau-Lingga Archipelago, Borneo,

Sumatra, Borneo, Malayan and Indochinese Peninsulas C. lowei Lowe's otter civet;

Sarawak

ground

tangalunga); Malaya, Sumatra,

bennettii Otter civet (water civet);

and

mongoose;

China, Nepai, Assam, Myanmar,

Hainan, Sumatra, Borneo,

FAMILY HERPESTIDAE Mongoose Family

Philippines

H. vittkollis Stripe-necked

SUBFAMILY HERPESTINAE

mongoose; Ichneumia

Atilax A. paludinosus Marsh (water

mongoose

:

mongoose Gambia east

to Ethiopia south to

I.

Afnca

S.

albicauda White-tailed mongoose;

W. African

crassicauda Bushy-tailed

forest regions

and S.W.

Liberiictis

L kuhn: Ubenan mongoose, Ubena Mungos M. gambianus Gambian mongoose, Gambia to Nigena

M. mungo Banded mongoose,

Nigena to N. Angola, C. Kenya,

Afnca south of Sahara, except Congo and S.W. Afnca

Uganda

Crossarchus C alexandn Alexander's mongoose (Congo mongoose); DRC, Uganda, Mount Elgon, Kenya C. ansorgei Angolan mongoose; N. Angola, S.E. DRC C. obscurus Kusimanse (dark mongoose, long-nosed mongoose); Sierra Leone,

Paracynictis P.

W

Cameroon C. penicillata

meerkat);

Yellow

S.

mongoose

(red

selousi Selous's

mongoose

(gray

meerkat'; southern Afnca from

Angola to northern

S.

Africa

Rhynchogale R.

mellen Meller's mongoose; DRC, Tanzania, Malawi, Zambia.

S.

C.

and

N.

Mozambique

Suricata 5.

Cynictis

S.

Sn Lanka

Africa; S. Arabia

mongoose; Mozambique, Malawi, Zambia, Tanzania, Kenya B. jacksoni Jackson's mongoose; C. Kenya and S.E. Uganda 8. nigripes Black-legged mongoose; S.E.

S. India,

Sub-Saharan Afnca except C. and

Bdeogale

suricatta Suricate (meerkat, gray meerkat, or stoksterjey Angola, Namibia, S. Africa, S. Botswana

Africa, Namibia.

SUBFAMILY GAUDIINAE

Angola, Botswana

Galidia

Dologale dybowskii Pousargue

mongoose

s

(Dybowski's or African tropical

G. elegans Ringta led

mongoose;

Madagascar

savanna mongoose); N.E. DRC, C. African Republic, S. Sudan,

Galidictis

W. Uganda

(Madagascar banded mongoose), Madagascar G. grand idieri Giant stnped mongoose; desert of S.W. Madagascar Mungotictis M. decemlineata Narrow-striped mongoose; W. Madagascar Salanoia S. concolor Brown mongoose, E. Madagascar

Helogale

Dwarf mongoose;

H. parvula

Ethiopia to northern

S.

Afnca west

H.

brachyurus Short-tailed

mongoose; Malaysia, Sumatra, Java, Philippines

ed wards: Indian gray mongoose; and C Arabia to Nepal, India, and Sn Lanka H. (Galerella) flavescens N. and C. Namibia, S. Angola H. fuscus Indian brown mongoose; H.

E

tangalunga Malayan civet (Oriental or

D. hosei Hose's

Indochina, Malay

Peninsula to Penang

Cynogale C.

S. India;

S.

Herpestes

megaspila Large-spotted

js; S.

H. un/a Crab-eating

Madagascar

to N. Namibia

Viverra

Lanka

swa

C. Namibia

Indochinese peninsula, Taiwan,

D.

pardicolor Spotted Iinsang; Nepal,

of

zeylonensis Golden palm civet;

Borneo

Assam, Sikkim, N. Myanmar, Indochina Osbomictis O. piscivora Aquatic genet (fishing genet or Congo water civet); Kisangani and Kibale-lturi districts

Travancore, and Coorg) P.

richardsoni African iinsang (oyan);

Cote d'Ivoire, Gabon, Cameroon, N. Congo, Fernando Poo Island

Nepal, Assam, Bhutan, Myanmar,

Kei Islands,

S.

Prionodon P Iinsang Banded Iinsang; W. Malaysia, Tenasserim, Sumatra,

palm

civet (toddy cat); India, Sri Lanka,

Ceram,

region of

Sierra Leone,

Sumatra, N. Borneo, S Andaman Islands; introduced to Japan

S.

Africa

(blotched genet or tigrine genet);

(Celebes)

Thailand,

W.

G. tigrina Large-spotted genet

Paguma

Common

region;

spotted genet); C. Africa, with restricted range in E. Afnca G. thierryi Vlilier's genet (false

Natuna islands Macrogalidia M. musschenbroekii Sul awes oalm civet (giant civet or brown palm civet); N.E. and C. Sulawesi

hermaphroditus

Cape

G. servalina Servaline genet (small-

Bilitung. N.

P.

Africa, C. Africa, S. Africa

(except

ferox Fossa;

C.

B.

G. maculata (formerly G. pardina

,

Flopei, Shansi,

Common

johnstom Johnston's genet

W.

Sn Lanka H. (Galerella)

Tanzania

(Lehmann's genet); Libena

palm rivet); Assam, Myanmar, Tha and, Malayan and Indochinese peninsulas, China (Yunnan civet (three-striped

Madagascar

Cryptoprocta

Peninsula, France, Palestine G.

rivet);

SUBFAMILY CRYPTOPROCT1NAE

Africa (north of Sahara), ibenan

Sumatra, Java, Borneo, Paiawan

Masked palm

S.

genet isma IIspotted genet or European genet);

G. genetta

India, Nepal,

larvata

Malagasy

Cape

sangu neus Slende" mongoose; Afnca south of Sahara H. smith ii Ruddy mongoose: India,

fossa Fanaloka (.Madagascar or

F.

Fiji

H. 'Galerella)

Angolan genet

of the Sahara except for ra n forest; S. Arabian Peninsula

Arctogalidia

goudotii Falanouc; E.C. to N.W.

E.

:

G. angoiensis

Arctictus

binturong Binturong ibear cat Bhutan, Myanmar,

Eupleres

Fossa

abyssr ca Abyssinian genet

ian Islands,

gray mongoose, S. Angola, Namibia, S. Afnca

Madagascar

G. felina Fei.ne genet: Africa south

SUBFAMILY PARADOXURINAE

P.

and

Ethiopian highlands, Somaiia

N. binotata Africa"'

Sma

Africa to KwaZulu-Natal,

Hawa

H. (Galerelia) pulverulentus

SUBFAMILY EUPLERiNAE

Namibia n south

SUBFAMILY NANDINIIAE

A. trivirgata

north through

Transvaal, N. Botswana,

and Genet Family

N.

A

E.

in

Indies,

naso Long-nosed mongoose; S:e Nigena to Gabon and DRC

H.

Islands

African over; Senegal east

to Somalia

Civet

S.

Sn Lanka, Bhutan; introduced to Madagascar, Sokotra, and Comoro

SUBFAMILY VIVERRINAE

and C. U.S. to E. Mexico and S. pygmaea Pygmy skunk; S.W. Mexico

mongoose;; N. Arabia to China ana Malay Per nsula;

Ball,

-Hainan, Taiwan, Indochina, India,

putorius Eastern spotted skunk;

5.

China, Myanmar, W. Malaysia,

Thailand, Sumatra, Java,

S. India,

G. fascia ta Broad-striped

mongoose

Sn Lanka

ichneumon Egyptian mongoose (Ichneumon); most of Afnca except Sahara, C. and W. African forest regions, and S.W. Africa; Israel, S. Spam, and Portugal

H.

H. javanicus (includes H.

auropunctatus Small Indian )

civet;

V.

indica Small Indian rivet (rasse);

mongoose

(Javan gold-spotted

113


SMALL CARNIVORES

Glossary Words

small capitals refer to

in

other entries

in

the glossary.

Adaptation features animal that adjust

it

to

its

may be produced

environment;

—

by evolution

of an

camouflage

e.g.,

Adaptive radiation when

(e

g

,

closely related animals

members

of a family)

have

other so that they can survive

in

different niches

grown animal

fully

that

Amphibian any cold-blooded .ertebrate of the class

in

Amphibia,

on land but

typically living

the water, e

g.,

and newts

frogs, toads,

Anal gland anal

anus or on either

it

Aquatic

that conveys specific information

by pines, spruces, and cedars

to others, usually to

of teeth especially adapted to

Corm underground food

the

shear with a cutting (scissorlike)

storage bulb of certain plants

visual or vocal elements, as in

Crepuscular active

threat, courtship, or greeting

edge;

living in

Arboreal

living

water

among

the

branches of trees

Arthropod animals with

mammals

in living

unique to

is

a

total

them

weight of

living material

used as a food source by

cutting

scavengers

trees for timber or to create

down from

open space

generation

a blind sac in the

for activities such as

animals

fingerprinting," a technique that

and

Delayed implantation when

allows scientists to see

the development of a fertilized

related to

large intestines. In

herbivorous

mammals the

is

it

end of the cecum appendix;

often

egg

site of

on

bacterial action

is

it

cellulose. is

in species

The

the

with a

is

suspended

period before

Domestication process

taming and breeding animals to

Births ere thus delayed until a

provide help and useful products

may

favorable time of year

for

function

Den

Dorsal relating to the back or

Cellulose the material that

constructed, used for sleeping,

forms the

giving birth,

retain

an antibacterial

walls of plants

cell

is

laid

that,

hard material that

some

down

mammalian

species

cementum

and

of individuals

numbers in

Ecosystem for

of

all

types of tooth

in

upper and

or

each

for

lower jaw are given. The

teeth to be found

International Trade in

typical

leaves of

molar (M). The

number

the total

example

in

the

name

nowhere

final

for Carnivora

A is

M3/3 = 44

also

and armadillos only

one

in

else

Estivation inactivity or greatly

decreased

of

skull.

is

for

small geographical area,

activity

during hot or

dry weather

Estrus the period

when eggs

Endangered Species. An

13/3,

Dentition animal’s

set of teeth

ovaries,

Desert area of low

rainfall

available for successful mating.

trade to

Cl/1, P4/4,

are released from the female's

and she becomes

Estrous females are often

permitted levels through a

dominated by

also (verb) to hide food for

system of licensing and

plants such as cacti

future use

administrative controls. Rare

Digit a finger or toe

"receptive" to

of food,

in

their

agreement between nations that restricts international

Cache a hidden supply

is

in

canine (C),

(I),

and

toothless, but

used as group

Endemic found

figure

(P),

whole system

a

plants, animals,

anteaters, sloths,

CITES Convention on

teeth lying behind

which

Edentate

which the

half of the

and Scats

Feces

environment interact

summarizing the dental in

Droppings see

raise

young, or for winter shelter

arrangement,

pocket

the upper surface

young;

Dental formula convention

under a microscope, can

a

and

to give birth

be counted to estimate the age

mouth used

raising

humans

spinal part of the body; usually

also (verb) the act of retiring to a

den

annual layers

in

a shelter, natural or

premolars and molars

and shrubs

of

reduced cecum the appendix

premolar

trees

is

completes normal pregnancy.

the order: incisor

Browsing Ceding on

who

for example,

the wall of the uterus and

the canines, consisting of

independence of young

whom,

particular offspring

Cheek teeth

through nesting to

analysis "genetic

which male was the father of

for a variable

implants into

it

from courtship,

territory),

handed

generation to

the junction between the small

digestive tract

the temporary storage of food

formation

is

DNA

opening out from

cycle of reproductive activity pair

contains the

living things;

all

genetic code that

numbers are always presented

(and often establishment of

specially

adapted

referred to as "in heat" or as

males

Callosities hardened, thickened

animals and plants are assigned

Digitigrade method of walking

Eutherian mammals that give

areas on the skin (e

to categories: (for instance

on the toes without the heel

birth to babies, not eggs,

touching the ground. See

rear

callosities in

Canine

some

g,, ischial

primates)

a sharp stabbing

(tootf

Appendix

1, 2).

See Volume

Cloven hoof

Canopy

formed from two

ont

-

jous (closed) or layer in forests

1

foot that toes,

produced by the intermingling

Congenital born with

< <

and

a

Plantigrade

pouch on the mother's

Dispersal the scattering of

Extinction the process of dying

each

young animals going

out

within a horny covering

of branches of trees

them without using

is

page 17

tooth usually longer than rest

broken (open)

114

down and removing

growing crops or grazing

Cheek pouch

entire

(deoxyribonucleic acid) the

substance that makes up the

main part of the chromosomes

alongside the

Breeding season the

DNA in

of

Biped any animal that walks on See Quadruped

lower molar

Deforestation the process of

two

legs

displays

winter (or the dry season)

teeth. In

variation within

of

Carrion dead animal matter

coats the roots of

Biomass the

members

can involve

species;

Carnivore meat-eating animal

crabs and insects

species

same

Diurnal active during the day

by trees that lose their leaves

Cementum

and the

Cursorial adapted-for running

premolar and

first

relatively

conspicuous pattern of behavior

twilight

in

Deciduous forest dominated

jointed outer skeleton, e.g.,

Biodiversity a variety of

the

involved are the fourth upper

very large;

sac) a gland

opening by a short duct either just inside the

side of

mountainous areas dominated

Carnassial (teeth) opposing pair

Cecum

has reached breeding age

breeding

organs from arteries to veins

Display any

and

Carnivora, and the teeth

a

evolved differences from each

Adult a

Coniferous forest evergreen forests of northern regions

arrangement

coloration

group of

Capillaries tiny blood vessels that convey blood through

mdition animal

is

to

live

in

which every

and the

away from where they were

dies,

born and brought up

forever

belly

last individual

species

is

lost


GLOSSARY

Eyeshine when eyes of animals

Gestation the period of

Inbreeding breeding among

(especially carnivores) reflect a

pregnancy between

closely related animals (e.g.,

beam

of the

is

of light shone at them.

caused by a special

layer (the

tapetum)

It

Gregarious

back

of the eye characteristic of

baby

birth of the

Grazing feeding on grass

reflective

at the

egg and

fertilization

many

living

together

in

loose groups or herds

nocturnal species and associated with an increased in

Harem

see

ability to

the dark

in

a

group of females

same

the

territory

living

and

cousins) leading to

Larynx voice box where sounds

Incisor (teeth) simple pointed

Latrine place where feces are

teeth at the front of the jaws

left regularly,

used for nipping and snipping

added

Indigenous

Leptospirosis disease caused by

living naturally in a

introduced

of closely related species that

plants (grazers

often also look quite similar.

thus herbivores)

on

Heterodont

Also used as a group

Volume

and

for

are created

region; native

dentition specialized

mammary glands

reduced survival rates

Herbivore an animal that eats

names always

in

offspring

genetic composition and

consorting with a single male

Zoological family

milk

weakened

Family technical term for group

and browsers are

Lactation process of producing

not an

(i.e.,

often with scent

leptospiral bacteria in kidneys

and transmitted

species)

via urine

Insectivore animals that feed insects

and

Mammary

similar small prey.

name

of

for

glands

characteristic

mammals, glands

for

animals such as hedgehogs,

production of milk

premolars, each type of tooth

shrews, and moles

Marine

within a species consisting of

having a different function. See

Interbreeding breeding

Matriarch senior female

parents and their offspring

Homodont

between animals of

member

Feces remains of digested food

Hibernation becoming

end

in

page

1

"idae." See 1

.

Also, a social

body

into canines, incisors,

1

group

inactive

body

different

of a social group

Metabolic rate the

within a single

species or varieties

the sea

living in

rate at

family or strain; interbreeding

which chemical

temperature to save energy.

can cause dilution of the gene

within animals, including the

Feral domestic animals that

Hibernation takes place

pool

exchange of gasses

have gone wild and

special nest or den called a

Interspecific between species

and the

Intraspecific between

from food

expelled from

as pellets,

in

often with scent secretions

live

in

a

hibernaculum

independently of people Flystrike

winter, with lowered

Homeothermy maintenance

where CARRiON-feeding

of

and constant body

same

individuals of the

activities

in

liberation of

Metabolism the chemical

species

Invertebrates animals that have

activities

temperature by means of

no backbone

turn food into energy

and

have

eggs

laid their

a high

burrows or

other true

(or

internal processes; also called

bones) inside their body,

underground tunnels

"warm-blooded"

mollusks, insects,

Frugivore an animal that eats

Home

crabs

living in

main part of the

fruit as

range the area that an

animal uses

diet

in

the course of

Fur mass of hairs forming a

normal periods of

continuous coat characteristic of

Territory

mammals

Homodont

Fused joined together

the teeth are

activity.

dentition in all

its

See

which

IUCN

and

place to another

and back

again, usually seasonal

Molars

International Union for the

large crushing teeth at

Conservation of Nature,

the back of the

mouth

responsible for assigning animals

Molt process

which mammals

and plants to

shed

internationally

agreed categories of

similar in

appearance and function

within animals that

Migration movement from one

e.g.,

jellyfish,

table

rarity.

hair,

in

usually seasonal

Monogamous

See

below

animals that

have only one mate

Gape wide-open mouth Gene the basic unit of heredity

unbranched prongs projecting

Juvenile young animal that has

environment

enabling one generation to pass

from the head of cloven-hoofed

not yet reached breeding age

Musk mammalian

on characteristics to

animals. Horns have a bony core

its

Horr s

offspring

Generalist an animal that

is

capable of a wide range of activities,

Keratin tough, fibrous material

Hybrid offspring of two

The 1

made

related species that can

species.

Volume

Kelp brown seaweeds

not specialized

a

plural

page

closely related is

genera. See

closely

interbreed, but the hybrid

is

in

a

at a

time

mountain

scent

Mutation random changes

with a tough outer covering

group of

Genus

Montane

a pair of sharp,

of keratin like fingernails

respiration

energy

Fossorial adapted for digging

flies

occur

in

genetic material

that forms hairs, feathers,

and

Native belonging to that area

protective plates on the skin of

or country, not introduced by

vertebrate animals

human

assistance

sterile

1

IUCN CATEGORIES EX

Extinct,

when

there

is

no reasonable doubt that the

last

VU

EW

when

Extinct in the Wild,

a species

is

known

only to

LR

outside the past range. Critically

DO

Endangered, when

extremely high

risk

a species

is

facing an

of extinction in the wild in the

immediate future.

Lower not

survive in captivity or as a naturalized population well

CR

Vulnerable, the wild

individual of a species has died.

NE

in

Risk,

satisfy

when

a species faces a high risk of extinction in

the medium-term future.

when

the

been evaluated and does CR, EN, or VU.

a species has

criteria for

Data Deficient, when there

is

about a species to assess the

risk

Not Eva U ated, IUCN cr t eria. |

not enough information of extinction.

species that have not been assessed by the

j

EN

Endangered, when extinction

in

the wild

a species faces a very high risk of in

the near future.

115


SMALL CARNIVORES

when

Natural selection

Placenta the structure that

an embryo to

animals and plants are

links

mother during

its

Roadkill animals traffic

weather) to ensure survival of

Plantigrade walking on the

Rumen complex stomach

the

soles of the feet with the heels

in

touching the ground. See

digesting plant material

and bad

fittest

New World

the Americas; Old

by road

killed

Ruminant animals

vegetation and later bring

Australia)

have more than one mate

Niche

part of a habitat occupied

by an organism, defined of

all

aspects of

Nocturnal

Nomadic fixed

in

terms

single

mating season.

Monogomous animals have a single

its lifestyle

animals that have no

Population

a distinct

Old World non-American

New World

again ("chewing the cud" or

Terrestrial living

"rumination") to

Territory defended space

assist

its

the

in

stomach

group of

species or

all

either by adjustments to

with scattered trees and low

metabolism or by moving

warm

rainfall, usually in

Scats fecal

areas

pellets, especially of

Posterior the hind end or

carnivores. Scent

deposited with the pellets as

Omnivore an animal

Predator an animal that prey for food

vegetable

Prehensile grasping

Opportunistic taking advantage

fingers

of every varied opportunity that

Premolars teeth found

arises, flexible

Opposable

behavior

fingers or toes that

can be brought to bear against others on the in

same hand

or foot

order to grip objects

Order

a subdivision of a class of

animals consisting of a series of

the female's ovary prior to

dominated by shrubs plants usually with

Promiscuous mating often with

one stem

many

Secondary forest

made up

it

on or

in

on two

Range

takes to

a "pair bond"

body

all

fours (a biped walks

legs)

the total geographical

area over which a species

is

distributed

Receptive when a female

of another

birth

Reproduction the process furry coat of a

for the next

Pelt furry coat, often refers to

Retina

ent

fur

produced by

is

(in estrus)

breeding, creating

mammal

Pheromone

—woody

more than

or shrub-

covered lands of the

far north

fine hairs

skin

and underneath the outer

coat of

stiff

trees that

mostly HERBIVORES

Uterus

Siblings brothers and sisters

of

womb

in

new

offspring

backbone

produce

reptiles), usually

fertile

offspring

May

include

beam

with a skeleton

made

( see above) of certain carnivores,

softer cartilage

especially otters

Vibrissae sensory whiskers,

is

in

parts

where the climate

too harsh for trees to grow all

Subspecies a

locally distinc

usually

on snout, but can be on

areas such as elbows,

t

birth to active

laying

young

such as barking and croaking

from the normal

Zoologist person

workings within plants and

withdrawn, as

animals

animal bodies, e g

typical cats,

digestion.

Keeping a warm-blooded state

is

in

of

the claws of

which can be folded

slightly

appearance of the

species live

species;

often

back into the paws to protect

mutual benefit more successfully

from damage when walking

than either could

on

its

who

Zoology the study

together for their

live

rather than

eggs

Symbiosis when two or more

a

eyes

or

Vocalization making of sounds

differ

called a race

at the

tail,

eyebrows

Retractile capable of being

shone

sometimes

Viviparous animals that give

parts of At

south of the Sahara Desert

group of animals that

of bones, but

Physiology the processes and

is

a

mammals,

(e.g., fish,

Spraint hunting term for scats

lying

a tapetum, a reflective layer

when

Vertebrate animal with

look similar and can breed to

Sub-Saharan

light-sensitive layer at

underneath

belly or

of an animal (opposite of dorsal)

courtship

species, e.g.,

Ventral the

Species a group of animals that

of the world

of

generation

the back of the eye

same

individuals within the

light

physiology

which embryos

mammals develop

causing eyeshine

mammal

and horses;

as pigs, deer, cattle,

and recognize them

a part of

mammals

hairs in

animals to enable others to find

,

forming a

Ungulate hoofed animals such

Steppe open grassland

ready to mate

skm removed from animal as

Tundra open grassy

on cleared ground

between

animal that

Parturition process of giving

Pelage the

one

Social behavior interactions

Parasite animal or plant that lives

of a species from

dense, woolly mass close to the

is

have been planted or grown up

of

acids. Essential in the diet

male and a female together

is

interpret

Underfur

that includes monkeys, apes,

mates, not just one

rate

testicles are

and ourselves

walks on

mate, marriage

and

of skin within

Scrub vegetation that

its

behavior that keeps a

beyond the time

Scrotum bag

Primate a group of mammals

Quadruped an

bond

location to another

located

of animals

fertilization

Pair

members

Pride social group of lions

Protein chemicals

Ovulation release of egg from

Scent chemicals produced by animals to leave smell messages

which the male

ammo

1

front

in

and reduced metabolic

Translocation transferring

markers

territorial

between sunshine and shade Torpor deep sleep accompanied by lowered body temperature

often

of molars, but behind canines

related animal families See

page

or

is

for othe s to find

Volume

1

kills live

tail

a relatively

tropical grasslands

behind another structure

that eats

on land

Thermoregulation the maintenance of

the animals of that species

almost anything, meat or

page

1

11

Olfaction sense of smell

continents. See

GENUS, FAMILY, ORDER, Class,

and phylum. See Volume

back from the stomach to chew

Savanna

animals of the same

order of

in

constant body temperature

breeding season

continuously

The categories,

SPECIES,

it

one

in

structure, origins, or behavior. •

that eat

digestion by microbes

mates

a male

with several females

home, but wander

only

mate

Polygynous when

active at night

a

groups

similarities in their

increasing broadness, are:

Polygamous when animals in

found

ruminants specifically for

Digitigrade

non-

refers to the

classifying organisms into

according to

American continents (not usually

World

the branch of

biology concerned with

pregnancy, allowing exchange of

(including predation

Taxonomy

beside rivers and

living

chemicals between them

challenged by natural processes

116

Riparian lakes

own

studies

of animals


FURTHER READING AND WEBSITES

Further Reading General

MacDonald, D

Mammals, Cranbrook, G., The

Mammals of

Southeast Asia, Oxford

New

The Encyclopedia of

,

New

Barnes and Noble,

York, NY,

2001

J.

Nowak,

and Redford,

F.,

Chicago

M., Walker's

R

Mammals

of the

The

K. H.,

MD, 1999

Baltimore,

IL,

1999

Skinner,

The Behavioral Guide to

D

African

Mammals,

,

D.,

J.

University of California

Specific to this

Buskirk,

and Smithers,

R.

volume

The

H. N.,

and and Conservation,

W., Martens, Sables,

S.

1994

Cornell University Press, Ithaca, NY,

of the Southern African

Subregion, University of

Pretoria, Pretoria,

South Africa, 1990

Strahan, R

The

Madagascar,

Mammals

of

Reed

Pica Press, Sussex, U.K.,

and

L.,

Bates, P P

Mammals of Arabia,

J. J.,

,

New

Dennis, N., and Macdonald, M., Meerkats, Struik,

Mammals

The

of Australia,

Holland, Australia,

1998

South Africa, 1999

The

Sevenoaks, U.K.,

Gittelman,

L

J

and

Ecology,

1999

,

Carnivore Behavior,

Evolution, Cornell University

Press, Ithaca, NY,

Whitaker, Harrison, D.

1975

CA, 1991

Press, Berkley,

Garbutt, N.,

Physiology, Oxford

Fishers: Biology

Mammals Estes, R

of Mammals: Their

Press,

Neotropics, University of

Chicago,

Press,

Anatomy and

Life

University Press, Oxford, U.K.,

World, The John Hopkins University Eisenberg,

The

Z.,

J

University Press,

York, NY, 1991

Mammals of the

Young,

Field

J

National

0.,

Audubon

Guide to North American

Mammals,

Alfred A. Knopf,

New

1996

Society Griffiths,

York, NY,

H

I.,

Mustelids

Modern

in a

World, Backhuys, Leiden, Netherlands, 2000

1996

1991

Jackson, P, Weasels, Badgers, Civets King,

C M., The Handbook of

Zealand Mammals, Oxford

New

University Press,

Oxford, U.K., 1995

Wilson, D.

E.,

The Smithsonian Book of

North American Mammals, Smithsonian Institution Press,

Mongooses, and

Switzerland and Cambridge, U.K., 1990

Washington, DC, 1999

MacDonald, Kingdon,

African

J

,

The Kingdon Field Guide to

Mammals, Academic

Press,

San

Wilson, D.

E.,

and Reeder, D.M.,

Species of the World.

Institution Press,

MacDonald, D

,

Mammal

Collins Field

D.,

The Velvet Claw, B.B.C

Books, London, U.K.,1992

A Taxonomic and

Geographical Reference Smithsonian

Diego, CA, 1997

and

their Relatives, IUCN,

Washington, DC, 1999

Neal,

E.,

Poyser,

and Cheeseman,

C.,

Badgers,

London, U.K.,1995

Guide to the

Mammals of Britain and Europe, Collins, New York, NY, 1993

Harper

Useful Websites General

http://www.nmnh.si.edu/msw/

http://www.iucn.org Details of species

http://animaldiversity.ummz.umich.edu/ University of Michigan

Museum

of Zoology

and

their status; listings by

Mammals

list

on Smithsonian Museum

site

the International Union for the Conservation of Nature, also

lists

IUCN publications

Specific to this

volume

animal diversity websites. Search for pictures

and information about animals by family,

common name.

and

class,

Includes glossary

http://www.panda.org World Wide Fund

newsroom, press http://www.cites.org/

IUCN and CITES by

scientific

species, or

listings.

name,

reports,

http://www.carnivoreconservation.org/

for Nature releases,

(WWF),

government

common name.

Location by

links,

recent books,

etc.,

on carnivore

ecology and conservation

campaigns

http://www.defenders.org/

Search for animals

order, family, genus,

News,

http://www.aza.org

Active conservation of carnivores, including

American Zoo and Aquarium Association

wolves and

http://www.ultimateungulate.com

http://www.wwfcanada.org/en/res

Guide to world's hoofed mammals

links/pdf/projdesc.pdf

grizzly bears

country and explanation of reasons for listings

http://endangered.fws.gov

Carnivore conservation

Information about threatened animals and plants

from the U.S.

Fish

and

Service, the organization in

Wildlife

American

in

the Rocky

Mountains

Website of the Wildlife Conservation Society

http://www.5tigers.org

charge of 94

million acres (38 million ha) of wildlife refuges

http://www.wcs.org

http://www.nwf.org

Comprehensive information about

tigers

Website of the National Wildlife Federation

117


Set Index A bold number shows

the volume and

page numbers

relevant

Common names

in

bold

(e

g

.

page numbers

animals

(e

g

parts of the set

in

Page numbers information

refer to the

:

B.

1

88

:

8.

aijfeus

.4

calabarensis 4: 106

— locate

parentheses e g., At-a-Glance boxes.

1: (24)

bandicoot 10

gazella 3: 9 A. pusillus 3: 9, *16-17

large short-nosed

trivirgata

Shan Lop Nur Nature

Resen/e 5: (102)

:

65

aardvark 1: 10. 5: (10), (12); 9 : 64, 65, (66), 67, 78-79 African 9 65 aardwolf 2: 102, 103, 110-111 Abrocoma bennetti 8: 30 Abrocomidae 8: 31 acacia 6: 87 Acinonyx A jubatus 2: 10, 26-29 A rex 2: 29 Aconaemys fuscus 8: 30 :

acouchi 7: 12

alpaca 5 92, 93. 105, (106) ambergris 3: 89 Amblonyx cinereus 1: 32, :

70-71 American Sign Language 4: 13. (16), 27 Ammodorcas darkei 6: 62 Ammotragus lervia 6: 62 angwantibo 4: 106, 106

Anomaluridae 7: 12, (19) anteater 1:9, 14, 9 64-67

84-85 Acrobatidae 10 (76) 10: 74,

:

6:

86-87 Aepyprymnus rufescens 10: 48 Aeromys tephromelas 7:

Africa, national parks/reserves 2: 16. 31, 5:

34

Afrotheres 5: (10) Afrotheria 9 : 10

agouti

W.

7: 8,

12, 14, 8:

28

42 43

8:

spotted (common) 8: 42-43

10: 27,

40-41

d'jr r jlture 1:

30

40-41 swainsonn 10: 27 antelope 5: 10, 11, 12, 13; 6: 60-63 American pronghorn 2: 26, 6: 63 four-horned 6 60, 62

pygmy roan

46, 2: 21, 28,

44, 69. 77. 79

A / dae

:

20

2 : 82 98-101 ,

10

:

Ailurus fulgent 1: 20. 3Q-31: 2: (99)

alarm

4 46. 57, 89, 100,

calls

:

6: 79. 7: 53.

albino 8: (84)

A

bu's-laphus 6 62 Ik htpnslcinn 6: 62

A

A akrs ai<

anrinrsom 6:

1

1

1

A aid-, shlrasi 6: 15 Allenopithecus ngroviridis A 1

Alopcr lagopus see Vulpes lagopus

118

fusca 4: 72.

74-7S

into 2

cinereus see cinereus i

nnr/it

US

1:

Amblonyx 32

nigrkeps 4 72 A trivirgatus 4: 77, 84-85 ape ape family 4: 12 13 :

Barbary see macaque, Barbary red see orangutan into nila 7

Aplodontidae

Apodemus

7:

12. 28,

A. laniger

4 96 :

sylvaticus

10 porcinus 6: 10 axis 6:

5: 74,

75,

58-59 olive

mangabey

4 10-11 :

sacred (hamadryas) 4 40, :

42 43,

4

5:

archaeocetes

59 3:

56

:

1: 32, 34,

35,

honey

9:

82,

7:

beetles,

free-

9 82, (83), 84-85, 86, 100-103 hairy big-eyed 9 85 hairy-legged vampire 9: 94 hammerheaded 9 80, 86 horseshoe 9 80, 87 house (Mexican freetailed) 9 82, (83), 84-85, 86, 100-103 :

:

vampire)

(false

9 82. 98-99 Kitti's hog-nosed

1:

1:

32,

82 83

1Q-111

B acutorostrata 3 1Q6-1Q7

little

brown

:

1

Mexican (83),

beluga 3 55, 80-83 bettong burrowing 10: 48, 51 Tasmanian 10: 8 Bettongia lesueur 10: 48 bilby 10: 44 45 :

greater 10: 27, 44, 45 lesser 10: 27, 44

binturong 1: 88, 89, 90, 91 biomedical research see medical research bipedalism 4: 10

wood

60, 62,

6:

6_4

69

67

68

B bonasus

6: 62,

66

blackbuck 6: 61 62 Blarma brpvicaud.i 9 28,

LQ 111

free-tailed

6:

Bison B bison 6: 62, 64 62 B bison athabascae 6: 68

:

30-33

:

55,

76-77

6: 62, (66),

9: 83,

104-105 long-eared 9

2:

62

American

long-tongued 9 86 :

beira 6: 60,

:

9 :

13

12,

bison

1 1,

86

long-nosed

lesser

83

see coypu

dung

9 108

lesser bulldog

83

28-29

swamp

86

greater horseshoe 9:

guano (Mexican

:

:

:

Palawan stink 1: 32 Balaena mysticetus 3 55, 1

vampire)

giant

:

horseshoe 9 106-107

Indian ferret 1: (7

82

:

lesser

78-81

2:

Canadian (American) 7 28, 29. 30-33 Eurasian 7 28, 29 mountain 7: 12, 14, 28, (29) mountain beaver family

98-99

(11), 9: 38, 80.

European

Malaysian sun

native Australian see koala

:

:

2: (60)

koala see koala

:

funnel-eared 9: 87, 87 greater false vampire

vampire

grizzly 2: 82, 83, 92,

American 7 28, 29, 30-33 beaver family 7 28-29

Indian greater false

5: 74, 86-87 badger 1: 34 American 1: (2, 76-77

dancing 2: (97) "dawn bear" 2: 82

sun 2: 82, 83 Beatragus hunteri 6: 62 beaver 7 8, 9, 70, 11,

:

42-43. 54-57 Babyrousa babyrussa

9

2: 9, 5:

spectacled 2: 82,

:

40,

:

98

American black 2: 82, 90-93 Andean 2: 82, 83 Asian black 2: 82, 83 bear family 2: 82-83 big brown (brown) 2: 82, 83, 92, 94-97 brown 2: 82, 83, 92, 94-97

sloth 2: 82,

:

4: 40,

:

:

:

(false

1

98

:

polar 2: 9, 82, 84-89 3 83 skunk see wolverine

diadem roundleaf 9: 86 disk-winged 9 87 Egyptian fruit 9 86 Egyptian rousette 9: 92-93 false vampire 9: 82, 98^99 fisherman 9: 108-109 free-tailed 9: 87 fruit 9: 58, 80, 81, 86

tailed)

4: 40, 43,

Balaenoptera

7

Appaloosa

96

A. occidentalis 4:

Axis

1

panda see panda,

:

1

29

:

:

introduced

9

94-97

Brazilian (Mexican) free-

Daubenton's 9 87

SO; 5: (97); 8: 72

:

yellow (savanna)

32

1:

A

m anus 6:

es gigas 6:

capensis

Aotus

1

A

Aonyx

:

a akes 6: io. 14-19 A akes akes 6:

a

:

:

pallid 9:

108-109

98

1:

Avahi

savanna 54=51

ants 9 64, 66, 69, 76

little

:

43. 58-59

60

84

vampire

false

bulldog (fisherman) 9 87,

:

12,(15), 24-25,

34, 38, 40,

A A

Alcelaphus

9,

American 9: 86 American American

9 82, (83), 84-85, 86, 100-103

mammals

Australia,

long-tailed see

:

:

10, 14; 7: 12, (37);

tailed

6: 62, 110-111 Antilocapridae 6: 63, 110 Antilope cen/icapra 6 : 62

6

76-77

geoffroyi 4: 72,

86-87 baboon 4: 8, 40, 42, 42-43 Chacma 4 56-57 gelada 4: 40, 42, 43, 62-63

96-97

:

1: 8,

jacksoni

bear

astutus 1 20 sumichrasti 1 20

brown 9 87 84 bat families 9 86-87 Bechstein’s 9: 80-81

belzebuth 4: 72

babirusa

62

B.

:

hamadryas

antlers

20-2 1

B

62

:

:

Ateles

A A

1

12; 8:

:

Bdeogale B crassicauda

20 20

:

frontalis 9: 21

A A

:

94-97

9: 84, (85),

B janetta 8: 56 B suillus 8: 56

9 80-87

12, 13

Antilocapra americana

111,8: 51,

55. 99 see also communication

A

62

Antidorcas marsupialis 6: 62,

:

:

vampire

pipistrelle

African slit-faced 9 82,

:

royal 6: 60,

Ailuropoda melanoleuca

bat

57

A. albiventris 9: 12,

A. algirus 9

gabbii Bassariscus B.

B.

Asian wild 5: 56-57 Asiatic 5: 42 5:

Bassaricyon B. allem 1

B.

aye-aye 4 96, 97, 102-103

sable 6: 62 Tibetan 6: 62

2: '99)

Ailurinae 1

6: 60, 62, (63),

6: 60,

6:

(29)

i

110-111 8:

:

At lax paludinosus aurochs 6: 63

stuartii 10: 27,

pronghorn

4

42. (44) African 5: 42

thumbless 9 87

Bathyergidae 7 Bathyergus

:

5:

:

spectacled 9: 87

:

108

98-99

spear-nosed 9 84, 87

:

barnacles 3 57, 92, 102,

A. africanus 8: 12

:

Agouti A paca 8: 30 A taczanowskn Agoutidae 7: 12

66; 6: 52 Arvicola terrestris 7:

:

:

:

102

A. macrourus 8: 12

dusky 10: 27 sandstone 10: 27 Antechinus

A A

black 8: 8-9 Central American 8: 30

common

brown

64

Atherurus

agile 10: 41

34,

9:

Artiodactyla 1: 70; 5: (10), 12,

A

:

Antechinomys laniger 10: 27 antechinus 10: 11, 25, 20, (25), 27

36

three-banded

artiodactyl 1: 8; 8:

Atelerix

9 65. 66

silky

66

domestic

short-nosed spiny see echidna, short-beaked

62 Addax nasomaculatus 6: 62 Aepyceros melampus 6: 62. adda>

74-77

65,

:

86-87 World false vampire 82, 87 World leaf-nosed 9: 87 World sucker-footed 9: 87 rousette 9: 86 sheath-tailed 9 87 slit-faced 9: 87 9:

9 110-111 white-winged vampire 9 94 see also flying fox;

:

:

southern naked-tailed 9: 65,

ass

banded see numbat giant 9: 64, 65, 68-71 marsupial see numbat

Acrobates pygmaeus

74-77 nine-banded 9

87

9:

Zealand short-tailed

whispering (long-eared)

:

:

9: 65,

Aspilia

animal farming 2: (97) anoa, lowland 6: 62 :

red 8: 30

65

long-nosed (nine-banded)

:

10 27 rufous spiny 10 27 Seram Island 10 27 striped 10 27 western barred 10 27 banteng 6: 62 bark stripping 8: 24 Raff ray's

lesser fairy 9:

72

4 72

:

:

64-67

9:

giant 9: palliate 4:

:

:

:

A. seniculus

(northern) 10 46-47 long-nosed 10 27 mouse 10 27 northern 10 46-47 northern brown (northern) 10 27, 46-47 pig-footed 10 27 rabbit-eared see bilby :

:

New Old 9 Old Old

:

common 9 65

A

25,27

8-9

:

88

armadillo

A

:

golden 10 27

3 98 Arjin

4 97

(10). 24.

:

eastern barred 10: giant 10 27

Arctonyx collaris 1: 32 Argentinosaurus huinculensis

names

100;

:

A

1:

9: 86 mustached 9: 87 New World leaf-nosed

mouse-tailed

:

bamboo 2 98-99,

:

Arctogahdia

bonaerensis 3 1 07 musculus 3 : 55,

98-101

1

:

A

Arctocephalus

Animals that get mam entries in the set are indexed under their common names, alternative common names, and scientific

8

06 4 106

Arctocebus 4

set.

:

in

in

the

in

103 point to illustrations of other than the main entry.

2

,

entry

9 78—79)

,

Archaeonycteris 1: binturong

Arctictis

aardwolf) mean that

mam

the animal has an illustrated Underlined page numbers (e g mam entry for that animal. Italic

followed by the

is

(e g., 1: 52, 74).

9:

82,

84-85. 86, 100-103

Blastocerus dichotomus 6:

10


SET INDEX

bubber

34

3:

72 84 85

58,

89, 91, (101)

bluebuck 6: 63 boar wild 5: 74 76-79 boat traffic 3: 50, 51, 65. 79, 103 bobcat 2: 10, 38-39. 40 body temperature 1: 9-12 control of 5 (17); 9: 24, 67, :

72. 82, (90; 10: 69 desert amma's 5 95-96 see also hibernation; :

bontebok

6:

grunmens

6:

argentata see Mico argentata C. geoffroy 4: 86 C. humilis see Mico C.

C. prevosti 7:

:

derbianus 10 14 philander 10 14 Caluromysiops irrupta 10 : 14 C.

93

72

9 12 Arabian (dromedary) 5: 92,

8: (84)

forestation in 4:

90-91

breaching 3: 96, 96-97, 103, 104. 109 breeding K strategists 7: 14 8: 21 R strategists 7: 14 selective 5 (43), 74; 6: 63 synchros zed 3: 20—2 :

6:83, 88; 7: 57 8: 79; 10: 41 see also captive breeding; reproduction; inbreeding;

6

7:

18 ,

100-103 camel family

:

:

:

’2'

dromedary

one-humped (dromedary) 5 92, 93, 94-99. 102' :

two-humped

:

Buphagus africanus

6: (73)

Burramys parvus 10: 74 burrows 7: 26-27; 9: 44-45, 67. 10: 99 ground destabilization 7: 49, 50, 59 8: 29, 57, 66-67; 9: 77; 10: 99

5 92, 93, (98), 100-103 camelids 5: 92-93

Camelus

100-103 C.

:

:

:

dromedarius 5 92, :

(

bactrianus 5 : 92,

Canis C. familiaris C. latrans C.

80-81

2: 50,

2

:

50, (53)

C.

106-107 1 06

C. ibex 6: 62,

C. ibex caucasica 6: C.

ibex cylindricomis 6:

C

catatonia 10: .20) cattle 1: 15; 5: 9. 71

1

06

humped zebu

6:

ibex nubiana 6:

Capreolus capreolus

1

6: 10,

63 79

6:

C

aperaea

39

8:

38-41

C. tschudii 8: 39 Cavndae 7: 12 Caviomorpha 7: (8); cavy 7 11. 12. (14) Brazilian 8: 39, 41

cavies 8:

and

62

sumatraensis 6: 62, 108 Caprolagus hispidus 8: 64 Capromyidae 7: 12. 8: 31

Capromys pilorides 52-53

8: 30,

30

:

:

:

:

2

:

(23). (29),

:

:

captivity 1: 22, 29, 65; 3: 59,

65, 71. 74; 4: 27, 78, 6 : 42-43, 58, 66, 91; 9: 21; 10: 83

capuchin 4: 10, 72, 83 4: 72,

73

Ceratotherium

:

simum

5: 28,

30-35

28-33;

5:

9

4:

1

cercopithecines 4: 40, 42, 43

Cercopithecus C aethiops 4: 44-47

C cephus 4: 40 C neglectus 4: 40 9, 10,

12

:

20

coati,

ringtailed

Coelodonta 5: 28 coendou see porcupine, tree 8:

1

mexicanus

8: 12,

26-27

4

:

41^12

40,

colobus 4 10, 40 40-42, 68-69, 69. 75 Angola (black-and-white) ,

4 40, 68-69 :

:

:

68-69

37

red 4 41^12,

68 4 40 western red 4: 40 :

7:

1

:

68-69 Colobus 4: 40

18

5,

48-49

7: 34,

Chiroptera 1 10, 9 : (86) Chiroptes C. albinasus 4 72 C. satanas 4: 72 :

C C.

angolensis 4 40, 68-69 satanus 4 40 :

:

colugo 1: 10, 7: (37); 8: 108-111 Malayan 8: 109, 109, 110, 110-111 108-109,

Philippine 8:

62

109, 110

commensal animals

45

chital 6: 10, 13.

Chlamydia psittac 10: 95 Chlamyphorus truncatus 9: 65 Chlorocebus aethiops 4 40 Choloepus C. didactylus 9 65 C. hoffmanni 9: 65

:

:

7 : 53, 101; 8: 16, 42, 51, 9: 61;

10:

alarm

80 calls

98 88

African palm

1:

banded palm

1

91 civet family

88 :

88, 90, 90,

7:

8: 51, 55,

57, 89,

53. Ill;

99

9 40,

cristate

:

48-51 Conepatus chinga

1: 32 mesoleucus 1 32 coney see pika, American Connochaetes C. gnou 6: 62, 82

C.

Endangered Species of Wild Fauna and Flora

:

American Sign Language 4: 13, (16), 27 see also songs Condylura

109

4 46,

6:

:

40

14

(24), 44. (49), 79;

:

chozchori 8: 29 Chrotogale owstom 1 88 Chrysochloridae 9 : 9 Chrysocyon brachyurus 2: 50 Chrysospalax trevelyani

7:

communication 1: (23); 3 (83); 4 (56). 74, 79, 85, 99-100; 5: 13, 15, 19,

:

African 1:

:

white-epauleted black ^lack-and-white) 4 40

30

48 Chironectes minimus 10: 14

1:

20, 21 1:

coatimundi see

Chinchillidae 7: 12

civet

1: 19,

30

8:

28

4 42 black-and-white 4 40,

CITES see Convention on International Trade in

40-41

94-95

:

tridecemlineatus

40 C. torquatus 4: 40 Cercopithecidae 4: 40

23

C. prehensilis 8:

Spermophilus

C. galeritus 4:

7:

black

8:

5:

88

rangiferina 6:

clompers 5 58 C amis aticeps

13

12,

chinchilla rat 8: 31

chiru 6:

88

Clethrionomys gapped

C.

Citellus tridecemlineatus see

nanus 10: 74 Cercocebus

Oadonia

satanic black

eastern

88, 91

1:

Civettictis c/v-efta 1:

colobines

short-tailed 8:

chulengo

concinnus 10: 74

Cervidae 6:

4: 11

common

9:

Cercartetus 10: 74

C

8

:

southern mountain 8: 10-11, 30 cavylike rodents 8: 8-1 Cebidae 4 72, 84 Cebuella pygmaea 4 86 Cebus 4 72 C apella 4 72 C capucinus 4 72 C. olivaceus 4: 72 cellulose 4: 41, 75; 5: 11-12, 70; 6 8; 7 103; 8 49

C.

captive breeding 1: 45, 47, 54,

brown

rock 8:

6:

Siberian 7:

:

C.

:

30

8: 28,

Owston's banded palm 1: 90, 91

Coendou

10

chipmunk

yellow-toothed

otter 1: 88, 91

:

Chilean 8: 30, 31

28-31

:

ringtailed 1: 28-29 w'hite-nosed 1 20, 29

chemical poisoning see

common

relatives

:

mountain

36-37 8: 28, 31

88 88 large spotted 1: 88 Lowe's otter 1 88 Malayan 1 88, 90 masked palm 1 88 Oriental 1: 88, 90 1:

large Indian 1:

coati

:

Chinchilla lanigera 8: 30,

:

C. crispus 6:

major 4: 96 medius 4 96

6:

90 palm

coalitions 2:

36-37 fur 8: JO)

Cephalophus 6 62, 80 C. dorsalis 6 62 C. monticola 6: 62

Capricomis

29

China, Imperial Hunting Park 6: 44, 45 chinchilla 7: 72; 8: ’0, 28,

63

Indian 1:

:

Cheirogaleus

water

:

civet oil 1: (91)

pygmy see bonobo

(^63; 8: 62

:

06 C ibex sibirica 6: 106 C ibex walie 6: 106 C.

12

:

C. ibex ibex 6:

:

C.

golden palm 1 88 Hose's palm 1: 88

small Indian 1:

chamois 6: 62 European 6: 108 Pyrenean 6: 62 charms 3 61; 8 67; 9 : 79 cheetah 2: 9 10 26-29

C.

90,

small-toothed palm 1: 88 Sulawesi palm 1: 88

3

chimpanzee

1

:

106

22-23 subspinosus Chaetomys o' 7:

catarrhines 4:

domestic see guinea pig Patagonian see mara Peruvian 8: 39

Caperea marginata 3: 55 Capra C. aegagrus 6: 62

:

chewing the cud

80-81 mesomelas 2 50, 62-63 simensis 2: 50

59, 100; 4: (17), 87,

caniventer 10: 14 fuliginosus 10: 14

48-49

6

30-33

Cetacea 1: 10 5: 10 cetaceans 3: 54-59 Chaeropus ecaudatus 10 27 Chaetodipus penicillatus

king 2:

88

1:

94-95

90.

:

6: 10. 26,

Catagonus wagnen 5: 88 catamount see puma

common

:

27 eaphus

pollution

:

:

2: 50, C.

canadensis nelsom 6: 26 canadensis roosevelti 6:

C.

common palm

Jerdon's

chevrotain 5: 8; 6: 10 greater Malay see deer, greater mouse Indian spotted 6: 10

:

2 50, 58-61

90-91, 98. 101; 5 26, 65, 6 91 7 15

C.

saber-toothed 2 : 1 sand 2 10. 13 tiger 2 10, 12 wildcat 2 10, 13,

lupus 2: 50. 54-57 lupus dingo (C. dingo)

c 20, 21 Caenolestes 10: 14

:

leopard 2: 10, 13 miner's see raccoon, ringtail

:

dingo

Cabassous unicinctus 9: 65 Cacajao 4: 72 C calvus 4: 72. 80-81 C. melanocephalus 4: 72 1:

:

C. porcellus 8: 30,

(55), (59), 71;

cacomistle

:

Cavia

100-103 Canidae 1: 20 C.

:

cattle rearing 4:

:

38-39

106-107 Demidoff's 4: 106 110-111 bushbuck 6: 62 bushmeat trade 4: 27, 32, 40, 53, 61, 85; 5 87; 6 59. 66. 72 bushpig 5 74, 75 4:

golden 2 10, 13 blackfooted 2 : 10, 13 cat family 2: 10-13 domestic 2 9, 10, (12). 49 European v. dcat 2 12 jungle 2 10, 13 Asiatic

humpless 5 92,

C. bactrian us

C.

27

Chaga's disease 8: 27 chain chorusing 5: 71

:

:

:

see also tunnels

bush baby

(Bactrian)

C.

27

canadensis nannodes 6:

8:

31 Castoridae 7 12, 28 cat 2: African wildcat 2 : 12

i.98)

:

92, 93,

5:

94-99. 102

C.

bubble netting 3: 104 bubonic plague 7: 14, 76-77 Bubulcus ibis 6 (73) Budorcas taxicolor 6: 62, 108 buffalo 1: 15 African 6: 60, 62, 70-73 see also bison. American Bunolagus monticularis 8: 64

:

C.

28

:

92-93

5:

domestication 5

C. ferus

13

bromei ads 4: 90 browsing 5 12, 13, 38; 6 9 brumby see mustang Bubalus B. depressicomis 6 62 B. mindorensis 6: 62 Bubastis 2:

98

92, 93,

5:

94-99

10, 11

:

8,

:

Bactrian

interbreeding

mound

5

93. 94-99. 102

.ariegatus 9: 65, 72-73 branding, freeze branding

brocket, red

:

camel

torquatus 9: 65

breeding

:

C

B.

,

36 36

C. notatusisi 7:

Cervus C. canadensis 6: 1 0, 26-29 C. canadensis manitobensis 6:

:

castoreum 7

36

mgrovittatus 7 :

9

capybara fa r ms 8: (50) caracal 2: 10 II G avaning 9: 39 caribou 6: 10, '2, 20-25 Carnivora 1: 10, IS; 2: 8 carnivores 1: 10 large 2: 8-9 small 1: 18-19 teeth 1: 15 'Carterodon sulcidens 8: 30 Castor 7 28 C. canadensis 7 28, 30-33 C. fiber 7:

10 48 Caluromys

Bradypus

Brar

86

4:

Caloprymnus campestris

~!2

Brachyteles arachnoides 4: B.

goe P

8: 8.

:

Callithrix

C.

Brachylagus idahoensis 8: 64,

C

Callimico

72

4:

capybara 7 8 '2, 28 10. 28, 48-51

72

4:

Callosciurus

74-75 62

38

personatus 4: 72 torquatus 4: 72

14-15

boto see dolphin, Amazon Bovidae 6: 9, 60-63 bovine tuberculosis 1:81 brachiation 4:

moloch

C.

Callorhinus ursinus 3: S

avanicus 6: Boselaphus tragocameius 6: 52 6.

C.

C.

62 Borhyaenidae 10: 26, 36 Bos B. frontalis 6: 62 6.

white-faced 4: 72

C acchus 4: 86, 92-93 pygmaea see Cebuella pygmaea

34-35

4: 12,

weeper

Callicebus

humilis

torpor

bonobo

Caenolestidae 10: 16

C.

:

C. taurinus 6: 62, 82-85 conservation 1: 47, (59), 75; 2: 33, 55. (88), 100; 3: 15, 19, 4 35, 5 26, 39, 63 6 25, 64, 7 45, 77 9 85; 10 (77), (95), (101) see also protection :

:

:

:

:

:

1:

88-91

119


SET INDEX

Convention on International Trade in Endangered Species of Wild Fauna and Flora (CITES) 1: 17 convergent evolution 9: 10, 1:

D. D. D.

26

10:

27

Dasyprocta punctata

8: 30,

42-43

03

1

Dasyproctidae

72

7:

Dasypus novemclnctus 74-77

desert 8: 64, 67, (93) Dice's 8: (93)

eastern 8: 64, 90-93 Mexican Guerrero 8: (93) New England 8: (93) Omilteneor 8: (93)

dasyure

swamp

short-furred 10: 25, Dasyuridae 10: 24, 36

8: (93)

puma

cougar see

coyote

1:

77

coypu 7: 72; 44—47

58-61

2: 50,

8: 9,

1

31,

1,

South American 8: (46) Craseonycteridae 9: 86 Craseonycteris thonglongyai 9:

102,

2:

108-109 23-24, 41, 72,

105, 8: 18-19; 9: 85, 91

Crossarchus alexandri

C,

ansorgei

1 1

:

:

Cryptomys C. damarensis C. C.

98 98

56 hottentotus 8: 56 mechowi 8: 56 8:

Cryptoprocta ferox

1:

5: 8, 9, 10, 12 barking see muntjac black musk 6: 10 black-tailed (mule) 6: 10,

34-37

7:

108, 110-111

C. vah 7: 1 08 Ctenomyidae 7: 72, Ctenomys C. frater 8: 30 C. validus 8: 30

culls 3: 15,

8:

Chinese water

10

6:

9

66; 8: 81, 10: (95) alpinus 2: 50

mouse

marsh

6: 10,

6: 10,

48

musk

.

6:

10

62 dimorphism, sexual 3: 87 dingo 2: 50, 80-81 10: (30), 37 Dinomyidae 7: 72 Dinomys branickii 8: 30 dinosaur 3: 98 Diplogale hosei 1 88 Diplomesodon pulchellum :

tufted

28

raccoon see also

Cynictis penlcillata

Cynocephalidae Cynocephalus

8:

C.

variegatus 8:

C.

volans 8:

1

see also habitat destruction 8: 29, 31 Bridge's 8: 30 mountain 8: 30 dehydration 5: 95-96 Delphinapterus leucas 3: 55,

degu

65 1: 98 108

Cyclopes didactylus

1

9:

80-83

09

09

C.

bennettii 1:

88

:

56-59

D Dactylomys dactylinus

8:

30

D tatei 10: 74 D trivirgata 10: 74 Dama dama 6: 10, 40-43 Damaliscus D. lunatus 6: 62,

Dendrohyrax

3:

69

D

88-89

8:

1

;

66-67 Dermoptera

3: 55,

3: 55,

1:

70; 8:

108

1:

Dugong dugon

3: 55, (61)

Indus' 3: 55, (61)

La Plata 3: 55, (61)

common

long-beaked (spinner)

forest 6:

76-77

common

69

dunnart

Kangaroo little

80-81

80-81

27

38-39 27 38

Island 10:

Dusicyon australis

76-77 56-57, 77

spotted 3:

white-beaked

38 38

3: 71

white-sided 3: 71 river 3: 55, (61)

2:

12

Echimyidae 7: Echimys pictus

50

10:

10: 105,

12, 8:

8:

30

przewalskii) 5: 42,

54-55

105

long-nosed 10: 105, 110

short-beaked 110-111

1

27

85 Enhydra lutris 1: 32, 72-75 Eohippus 5: 42 Eomanis waldi 9: 64 Equidae 5: 42 Equus E. asinus 5: 42, 57 E. burchelli 5: 42, 46-51 E. ca ball us 5: 42, 58-61 E caballus przewalskii (E.

E 1: 14; 8:

1

see also moose Nino 3: 22 Emballonuridae 9: 87 embryonic diapause

red-cheeked 10: 27

rollover (spinner) 3: 55,

Yangtze

52-53

6: 62,

Duplicidentata 8: 61

3: 55,

102 02

10: 12- 13, 57, 63, 69,

long-tailed 10:

echidna

1

71, 73,

river 3: (61)

short-beaked common 3: 69 short-snouted spinner 3: 76

7:

El

10:

Gilbert's 10:

quercinus 7:

Tule 6:

fat-tailed 10: 25,

white-sided 3: 55, 70-71 pink (Amazon) 3: 55, 60-61

melanurus

E.

Siberian 6:

02 46-47,

6: 62,

10:

4,

Manitoba 6: 27 Merriman 6: 26 North American 6: 12 Rocky Mountain 6: 26-27 Roosevelt's 6: 27

1

80

common

E.

Irish 6:

3: 47,

duiker bay 6: 62 blue 6: 62 bush (common)

1

elk 6: 10, 26-29 eastern 6: 26

102 102

10, 3:

5:

Eliomys

5: (12)

:

59

22-27

4: 40, 42 dromedary see camel, dromedary

:

9: 59,

Elephas maximus

Draculin 9: 97

D. sichuanensis 7:

hourglass 3: 71

3: 55,

North African

woolly 7: 102 douroucouli see monkey, Northern night

52-53

9: 59,

short-eared 9: 59, 59 Elephantulus rozeti 9: 59

102

dugong

15

62-63

Setzer's mouse-tailed

.

5: 14,

golden-rumped

102

D. nitedula 7:

76-77

40-43 Pyrenean 9: 7 7, 40, 52-53 Russian 9: 40, 42, 43, 53

102

Oriental 7: 103

68-69 71

14-1

elephant shrew 1: 10; 9: 10, 58-59 black and rufous 9: 59, 59 checkered 9: 58-59, 59 four-toed 9: 59, 59

102, 102

D. lanigerl

3: 71

spinner

9: 9, 11,

60-61

Risso's 3: 71

bennettianus 10: 48 goodfellowi 10: 48,

desman

:

Pacific

68-69

02 D. arboreus 8: 1 03 D. dorsalis 8: 103 D validus 8: 1 03 Dendrolagus D.

Dactylopsila

1 98 54-59

Fraser's 3: 71

Ganges

savanna

Dromiciops gliroides 10: 14 drought 4: 63; 5: 34, 94 drug testing 4: 13, 93

bottlenose 3: 55, 66, 72-75 clymene 3: 76

3:

D. delphis 3: 55,

lowei 1 88 Cynomys ludovlcianus 7: 34, C.

8: 30,

5:

14-15

forest 5: 14,

masked mouse-tailed

7:

22-27

family units 5: 15

Dryomys

Amazon

long-beaked

Delphinus D. capensis

Cynogale

70; 3:

1:

elephant family

02

drill

dog

patagonum

dusky

86

78-79

50

2:

73, 75, 81, 90; 5: 20, 6: 49, 51, 9: 23, 91

9:

50-53

2:

prairie

spotted waigeou

Cyanophenus

78-79

5: 14, 15, 16,

circus elephants 5: 24, (24)

106-107

7:

10, 5: 8, (10),

1: 10,

(

,

26 Asian

102,

garden 7: 102, 102-103 hazel 7: 102, 103,

7:

1

0,

12 ) African 5: 14, 15, 16-21. 11

spectacled 7: 102 spiny 7: 103

wild) 2: 9, 50, 53,

common

cuy see guinea pig

elephant

104-105

7:

1

44-45

102-103 7: (14),

6:

electrocution 4: 67

dormouse family

2: 9, 50, 53,

domestic 2: 9, 50, (53) painted hunting (African

4: (17), 27, 32, 37, 61, 69,

74 12-13, 82-83 74

Elaphurus davidianus 6:

Roach's mouse-tailed

53

2: 50,

small Sulawesi 10: 10:

9

1: 15; 5:

dolphin

76-77

(hazel) 7: 102,

Japanese

76-77

Elaphodus cephalophus

103, 106-107 desert 7: 102, 103

1

76

6: 60, 62,

6: 62,

Lord Derby's (giant) 6: 62,

garden 7: 102 Chinese 7: 102 Chinese pygmy 7: 103

common

32

1:

13

5:

common

12, 13, 15,

1,

104-105

5: (10)

sde also brocket deer parks 6: 1 3, 42-43, 47 deforestation 1: 21, 31, 48, 51, 71; 2: 58-59; 3: 61,

1

barbara

eland

giant

forest 7:

24-25

7:

Dologale dybowskii

(36)

7:

Eira

fat (edible) 7: (14), 102,

32-35

6:10

10:

17, 19

7:

104-105

Eimer's organs 9: 45, (50)

48 vanheurni 10: 48

madeayi

edible

rufescens 10: 27

egret 6: (73) Eidolon 9: 86

Asiatic

Salt's 6:

Dollchotis

13

6: 10,

48 luctuosa 1 0: 48

African 7: 103

dog family

30-33 38-39 60

48

10:

dormouse

62

Kirk's 6: 60,

pampas

6: 10, 12, 13,

E.

10:

hagem

28, 32, 37,

8,

ecotones 6: 86 Edentates 9: 64, 64 egg-laying mammals

Dorcopsis

18-23

78-79

6: 10, 13,

62

6:

48

white-striped 10:

D.

dik-dik

bush

6: 10, 12,

dorcopsis gray 10:

Dorcopsulus

14

D. virginiana 10: 14,

5: (45)

44-45

Dorcatragus megalotis

D.

muntjac see muntjac musk 6: 10

red roe

wild 5:

D.

Didelphis

wild) 2: 9, 50, 53,

6: 10, 72

22-23 80-81, 92-93, 99, 106, 109, 110-111 echymipera, Clara's 10: 27 Echymipera E clara 10: 27

donkey

D.

African wild

common

10:

36-39

Virginia

dog

Echinoprocta rufescens 8: 1 Echinops telfairi 9: 24 Echinosorex gymnura 9: 12, echolocation 9:

(72) see also cat, domestic;

African hunting (African

72

white-tailed 6: 10, 13, 18,

ground 10: 74 mountain 10: 74 peleng 10: 74

28,

dicoumarol 7: 75 Didelphidae 10: 14, 16 common see opossum,

mouse see chevrotain mule 6: 10, 34-37

spotted

spotted (spotted) 10: 12-13, 82-83

5:

Diprotodontia 10: 25, 75, (94) diseases 1: 26, 87 rodent-borne 7: 14 distemper 3: 41 canine 2: 79 Distoechurus pennatus 1 0: 74

lesser

105-106, 107; 38-39, 40,

(45), (98),

domestic

diastema 7: 10, 7 7; 8: 60 see also reproduction dibatag 6: 67, 62 dibbler 10: 27 Dicerorhinus sumatrensis 5: 28

Dtpodomys ordn

sika 6: 12

cuscus 10: 76, (77) admiralty 10: 74 bear 10: 74

85

73,

5: (43),

dog, domestic

10: 12-13, 57, 63, 69, 71,

fallow 6: 10, 12, 13, 40-43 greater mouse 6: 10,

Siberian

50

cursorial animals 2:

63

3:

Diplomys labilis 8: 30 Dipodidae 7: 72

44-45

Cuon

whale pods diapause, embryonic

deer and relatives 6: 10-13 dwarf musk 6: 10

Pere David's

43, 45, 67; 6: 32,

57

2: 50,

dialects^of

9:

6: 10,

domestication 2: 53; 6: 25, 63; 8:

;

50-51 hog 6: 10

88,

Cryptotis parva 9: 28 Ctenodactylidae 7: 12, 108 Ctenodactylus

gundi

102-103

103

48-49 Himalayan musk

96-97

C.

32-33

Daubentonia madagascariensis 9:

dhole

74

dolphinaria 3: 71,

27, 28-31

D. albiventris 10:

D. hallucatus 10: 27,

91; 6: 13, 78, 81, 7: 100,

C.

27

Dasyuromorphia 10: 24 Dasyurus D. albopunctatus 10: 27

4: 96,

Desmana moschata 9: 40 Desmodus rotundus 9: 94-97 devil, Tasmanian 10: (24),

Dlceros bicornis

deer

107 Crocuta crocuta

cria 5:

5:

9: 65,

broad-striped 10: 27 narrow-striped 10: 25, 27 red-bellied 10: 27

DDT

38

crop raiding

6:

Dasykaluta rosamondae

62

8: 51,

coruro 8: 29, 30 cotton bollworms 9: cottontail 8: 62

120

korrigum

D.

coonskins 1: (23) coppicing 7: 106

coprophagy

88 lunatus lunatus 6: 88 lunatus tiang 6: 88 lunatus topi 6: 88

D. lunatus

pygargus 6: 62 dassie, rock see hyrax, rock Dasycercus cristicauda 10: 27

40; 10: 36, (52)

coon hunting

88

D. lunatus jimela 6:

31

52-53 56-57 £. przewalskii 5: 42, 54-55 £. quagga 5: 48 E. zebra 5: 42 Eremitalpa grand 9: 40, 56-57 Erethizon dorsatum 8: 2, 20-25

£

grevyi 5: 42,

£.

hemionus

5:

42,

1


SET INDEX

Erethizontidae 7: 72; 8: 12

Erinaceidae 9:

9,

Erinaceus E. concolor 9:

europaeus ermine 1:112

14-19

see also deforestation fossa 1: 88, 90, 97, 96-97 4: 100

see also stoat Erythrocebus patas 4: 40 Eschrichtidae 3: 92

92-97 estivation 7: 19, 9: 13, 21

Ethiopia, geladas in 4:

Arctic 2: 50, 70-73. 87;

108-109

australis 3:

7: (14)

55 eucalyptus 10: 95-96, 96 Eulemur 4: 96 glacialIs 3:

E.

bat-eared

2: 50,

76-77 53

Blanford's 2: 50, blue Arctic 2: 71, (72)

E.

coronatus 4: 96

Cape

E.

macaco

E.

mongoz

corsac 2: "cross fox" 2: 64, fennec 2: 50, 52, gray 2: 50, 52

96 96

4: 4:

Eumetopias jubatus 18-19 Euoticus E. elegantulus 4: pallidus 4:

E.

3: 9,

06

Eupleres goudotii 1: 88 Euroscaptor parvidens 9:

Eutheria 10: (10)

evolution

convergent

mammals

bengalensls caracal

F.

catus 2: 10

15-16

10

2:

chaus 2: F concolor

1

2: 10,

lynx 2. 10,

42-43

64

2:

swift 2: 50, 52, 68-69 white Arctic 2:71, 72 freeze branding 8: (84) 7:

40-41

F.

rufus 2:

F.

serval 2:

F.

silvestris

F.

temmincki

F.

tigrinus

'

1

2:10

yaguarondi 2: 1 Felovia vae 7: 108 Fennecus zerda see Vulpes zerda ferret

1:

5:

1

45

black-footed

1:

32, 34, 35,

46-47 see implantation/ fertilization; reproduction

fertilization

50-51 farms 3: 65

fisher 1: fish

;

8:

20

fishing 3: 12, 15, 19, 37, 43,

105

106

107

thick-tailed 4: 106,

Thomas's

4:

Zanzibar 4:

69, 71, 77, 79, 105

flehmen 5: (11), 62, 65 flood water 4: 80 flying fox 9: 80, 86 Indian 9: 88-91 food poisoning 7: 74 food sharing 9: (96)

26

western

G

4: 36,

4: 1

06

1

G. senegalensis 4:

06

1

G. zanzibaricus 4:

06 30

1

Galea musteloides 8: Galemys pyrenaicus 9: 40,

52-53

2, (2

Graphiurus ocularis

Area little 1:

32,

7:

99

4: 8, 10, 40, 42, 43,

guiara 8:

kordofan 6: 52 Masai 6: 52, 54 nubian 6: 52

guinea pig domestic

54

56

8: 10, 30,

38-41

38-39, 39, 40

8:

wild 8: 28,

Gulo gulo

39-40 32, 56-57

Thornicroft's 6: 52, 53

desert 7:

West

felou 7: 108, 109, 109

Galidia elegans

1

Giraffidae 6:

6:

98

National Park 5: 34

62

G dama

gleaning glider

9:

feathertail 6:

62

G. teptoceros 6: 62 G. subguttarosa 6:

thomsoni

6:

62

94-95

1:

13 108, 109

7: 12,

gundi family Lataste's 7:

(1

5)

G. sabrinus 7: 61 G. volans 7: 34,

Gazella

G.

52-53

Glaucomys

98

G. grandidieri 1:

Garamba

gerenuk

giraffe-gazelle see

98

:

Gir Forest Reserve 2:

G. fasciata 1:

gaur

African 6: 52

gundi

60-61

84 (pygmy) 10: 74,

77, 84-85 greater 10: 74 mahogany 10: 74, (77) pygmy 10: 74, 77, 84-85

mzab

7:

108-109

hedgehog

10, 14; 8: 12;

1: 9,

9: 8, 9, 11

African 9: 12, 13

pygmy

9: 12,

9:

12

collared 9: 12

Daurian

9:

12

desert 9: 12, 13 dwarf (African pygmy) 9: 12.

20-21

eastern European 9: 12 four-toed (African pygmy)

20-21

7: 108, 109,

9: 12,

7:

109 108,

13

short-tailed 9: 12, 12 9: 12,

12

hedgehog family

9:

12-13

Hugh's 9: 12, 13 long-eared 9: 12, 12-13

Madagascan

9:

North African

25

9: 12,

13

western European 14-19

white-bellied (African

pygmy)

9: 12,

20-21

Helarctos malayanus 2: 82

Speke's 7: 108, 109, 109 Gymnobelideus leadbeateri 10: 74, 88-89 gymnure 9: 12, 13

shrew

1

Hawaiian Islands National Wildlife Refuge 3: 31 hawks, bat 9: 84

9: 12,

109

North African 110-111 Saharan 7: 108

Hainan

62

hairy 9: 12, 22

30

black 8: 41

reticulated 6: 52, 53,

hare hunting 8: (89) hartebeest 6: 62

9: 12,

giraffe family 6: 52-53

Galictis

32 1: 32

49,

wild 5: 93

guenon 44

8: 62, 64,

74-79

Asian

34

groundhog see woodchuck guanaco 5: 92, 108-109 .

American

pika,

20-21

50, 59; 8: 29, 57, 66-67;

54 54-57

64

African

32

9: 77, 10:

tippelskirchi 6: 52, 5: 11, 12, 6:

102

103

5:

1:

7:

ground destabilization

thornicrofti 6: 52

8:

mouse see

sustainable 3:

grass see sea grass grazing 5: 12, 13, 30, 31, 6: 67, 89 Great Gobi Strictly Protected

54

64-67 66

8:

hispid 8: 63, 64,

Lichtenstein's 6:

dryas 10: 14

Galeopithecidae 8: 108 :

26

G. agilis 10: 14

54

(Arctic) 8: 74,

harvesting 3: 28; 6: 18 controlled 3: 12, 16

Rothschild's 6: 52, 54, southern African 6: 52

G. cuja 1

1 ),

Camelopardalis 6: 52 G. Camelopardalis capensis 6: 52 G. Camelopardalis peralta 6: 52 G. Camelopardalis reticulata 6: 52,

64

82-85 hare family

snowshoe

Gracilinanus

60

demidoff 4: 106, 110-111 G. thomasi 4: 1 06 G,

1

G. Camelopardalis

giraffe

Galagoides

2,

26-27

(21),

G. Camelopardalis

06

1

G. gorilla gorilla 4: 12,

grison

8:

polar (Arctic) 8: 74, 82-85 Smith's red rockhare 8: 64

beringei 4: 12, (21)

G. gorilla diehli 4:

G

80

Patagonian see mara

20-25

52

8:

Jameson's red rockhare 4: 8, 12,

G. gorilla 4: 12, (21)

6:

86-89

8: 62,

see pika, American

64

Giraffa

antiquorum

4:

Greenland

G. beringei diehli 4: 12, (21)

angolensis 6: 52

simus

haplorhines see primates, higher hare 8: 61 Arctic 8: 74, 82-85

greater red rockhare 8: 63,

apes in 4: 50 Gigantoplthecus 4: 12

G. Camelopardalis

96 96 96

4:

European

4: 12, (21)

G. beringei beringei 4:

Gibraltar,

aureus

H. griseus 4:

Cape

Gorilla

38-39

84-85

brown

20-25 22-23

4: 12,

western lowland (21), 26-27

rothschildi 6: 52,

gallery forests 4: 101

fleas 9: (18)

37

lar 4: 36,

G. Camelopardalis

Galidictis

3: (74)

fishing nets 3: 22, 30, 50, 59,

kloss 4: 36, 37,

Syrian (golden) 7:

Hapalemur

calling

silverback 4: 13,22,

G. Camelopardalis

06 106 1

G. gallarum 4:

G. vittata

cooperative, by dolphins

forestry operations 5:

106

Galago G. gabonensis

10

F.

fermentation

4:

0,

2:

4:

54-57

margarita 2:

44-45 10, 38-39 10, 46-47 2: 0, 48-49

36-37

gibbon family

G. Camelopardalis

F.

1

mountain

crested black 4: 36

G

4:

26-27

7:

pocket 7: 12, 18 western pocket (northern

Cross River 4: 12, (21), 26 eastern 4: 12, (21) eastern lowland 4: 12, 13, ( 21 )

galago Demidoff's dwarf see bush baby, Demidoff's Gabon 4: 106

06

18

7: 17,

goral 6: 62, 108 gorilla 4: 8, 10, 12, 13

G. Camelopardalis 6: 52,

Senegal 4: 106 Somali 4: 106 southern needle-clawed

pardalis 2:

56

18, 19

H.

gopher

horned 7: 12 northern pocket

crested 4: 37

(lar)

lynx pardinus 2: 40, 41

F.

8:

9: 10; 10: (1 1),

108

30

8:

hamster 7: 11, 15, golden 7: 84-85

H.

Gondwanaland

4: 36,

white-handed

F.

1

(lar)

(72), 75. 3: 12, 16; 4: 40,

1

108-109

6: 62,

pocket) 7: 26-27

37

36, 37, 46, 61, 90; 9: 43, 10: 22, 65, (80) see also skin trade

63

6:

mountain

31-32, (31)

4:

60-63

6:

domestic

92-93

G. tigrina 1: 88 Geocapromys brownii Geogale aurita 9: 24 Geomyidae 7: 72

Muller's 4: 36,

Garnett's 4:

1

88,

12-13, 35, 41, 44, 69, 85; 7: 33, 41, 97, 8: (10),

Halichoerus grypus 3: 42-43 hammer stones 1: (74);

gnu

goat

38-39 34

see also deforestation habitat preservation 4: 51 habituation 4: (23)

wild 6: 62 1:

64, 68-69, 87, 93; 2:

F.

2:

Genetta G. genetta

10: 27, (77), (95)

see also wildebeest, blue

(common) 92-93

38-39 moloch 4: 36, 37

northern needle-clawed

F nigripes

88-91 88

85, 9l;

9: 33, 62, 63,

Globicephala melas 3: 55,

white-tailed 6: 82 1:

large-spotted 1:

common

46-47 9: 87

lynx canadensis 2: 40, 41 lynx lynx 2: 40, 41

F.

92-93 genet family

88,

1:

Mongolian 7: 88-89 gerenuk 6: 62, 98-99 gibbon 4: 8, 10, 12

53

Ruppell's 2: 50,

Samson

Furipteridae

2:10

F.

F.

1:

92-93

88,

European (common)

88,

59, (66); 7: 25, 63, 107;

104-105

Glyptodon 9: 66 G. panochthus 9: 64 gnawing 7: 10, 16-17, 64

88, 91 1:

gerbil 7: 13, 15 bushveld 7: 13

F

farmland, expanding Felis 2: 10

1:

65, 91; 6: 20, 47, 49, 51,

see dormouse, edible

glutton see wolverine

98

50,

1: 9; 2:

70

fur trade 1: 22, (23), 51, 54,

Madagascan 1:91,97

aquatic

62

18,

red

2: (72); 8:

facial expression 4: (56), 100 falanouc 1: 88, 97 fanaloka 1: 88

6:

Georychus capensis

fur farming 1: 22, 35, 54, 55,

of rodents 7: 72 extermination 6: 64-66

F.

68-69 (59), 64-67

35, 87, 96, 101; 5: 26, 63,

102

66-67

50, 52,

10

1:

F.

50

Funambulus pennantu

9: 10, 40,

10: 36, (52)

of

65 74-75

2: 35,

Glirulus japonicus 7: Glisglis 7: 102,

4: 40, 42, 43,

kit (swift) 2:

40

Eurotamandua 1: 9 Euryzygomatomys splnosus 8: 30

gemsbok genet 1:

1:

53

island gray 2:

06

1

gelada baboon 62-63

44, 79, 3: 83; 4: 24,

Glironia venusta 10: 14

glis

small-spotted

53 50, 53

2: 50,

Indian 2: 50, 1

.

Tibetan 6: 67, 62

common

fox

63

Eubalaena E.

Fossa fossa 1 88 fossorial animals 7: 18, 65 Fouchia 5: 28 fovea 4: 1 07 :

Eschrichtius robustus 3: 55,

62

6: 67,

110 :

habitat destruction 1: 62, 64;

gliding marsupials 10: (76)

94-95 97,

6:

H

76, (76),

.

slender-horned

Thomson's

26

gorillas in 4:

62

6: 67,

goitered 6: 67, 62

gallery forests 4: 101

1

sugar 10: 74, 86-87 88

47

5:

dama

90-91 elephants in 5: 22

9: 12,

E.

gazelle

forests in Brazil 4:

1

Heliophobius 7: 9 H. argenteocinereus 8: 56 Flelogale parvula 1 98, :

106-107 Flemibelideus lemuroides 10:

74

Hemicentetes semispinosus 9:

24

Hemiechinus H. aethiopicus 9:

H. auritus 9: 12, H. collaris 9:

1

12-13

1

121


SET INDEX

Hemigalus derbyanus 1: 88 Hemitragus jemlahicus 6: 62 herding, predator avoidance 5: 10 Herpestes H. edwardsii

Hydrochaeridae 7 12 Hydrochaeris hydrochaeris

108-109 1 ichneumon 1: 98 naso 1: 98 Herpestidae 1: 88 :

Hydromys chrysogaster 1

02

H antmeae 8: 03 H brucei 8: 03 1

1

03

Heteromyidae 7: 12 Hexaprotodon libenensis 5: 66 hibernation 2: 87-88, 93, 97, 100, 7: 19, 51, 54-55, 107. 9: 13, 18, 27, 82, 1 1, 10: 111

104, 105,

1

Himalaya Mountains 1: 30 hippo see hippopotamus Hippocamelus

H H

antisensis 6:

38-39 Hyemoschus aquaticus 6: hyena brown 2: 102, 103, 103 hyena family 2: 102-103 spotted striped

H H

hippopotamus family 5:

63

giant forest

5: 74, 75,

74 red river 5: 74 see also warthog homeothermy 1: 9-10, 14

Hommidae

Homo

8: 103,

Cape

horse

1: 15, 5: 8, 9,

11, 12,

Camargue 5:

domestic

1

rock (rock) 8: 103,

tree 8:

1

8

:

8:

42

17

12, 8:

58 feral see mustang horse family 5: 4J 4$ Mongolian wild 5: 42,

H H H

12,

(8),

:

6: 62, 1.Q6

wild) 5: 42, 45, 54-55 racehorses 5: (43)

nubian

'Mongolian wild'

42, 45, 54-55 wild (mustang) see 4

h

mustang

h

102

10: 48, 51, 52,

107

K

tonyz striatus

1:

32

Cuban

8: 30, 52.

5J Desmarest's Cuban (Cuban)

K

2: 82. (86), 93, 3:

K

8: 30,

i''''

4:

22, hi.

29, 33,

'id.

93;

13, 66. 70, 8: 57, 59,

eared 8: 30

10 H 8:

:

101

A

8

West Indian

see also reproduction imprintinu 6: 83 1: (59), 2:

1

Indricothenum 5: Indri mdri 4: 96

8.

8:

30

28

(10), 77. (77),

92-93

92 93

7:

62 K leche 6: 92 K vardonu 6: 62 Kogia K breviceps 3: 55 K s imus 3: 55 korrigum 6: 88 Kruger National Park 2: 31 kudu greater 6: 78 19 kulan see ass, Asian wild kultarr 10: 25. 21

L.

conspicillatus 10:

L.

hirsutus 10:

langur

arcticus 8:

L.

californicus 8: 64,

L.

europaeus

48

12-13, 4: 40,

36

14

65

collared 7: 9

Norway 7: 66, 90-91 lemmus lemmus 7: 90-91 96-97 97 96, 96

96; 4:

1:

black 4:

4: 96,

105 97 4:

96

96 97

flying 8:

4:

96

96 96

bamboo 4:

4:

96

4:

greater dwarf 4:

96

96

hairy-eared dwarf 4:

96

97

Milne-Edwards's sportive

97 mongoose 4: 96, 96 mouse 4: 97, 97 pygmy mouse 4: 8, 96, 97 4: 96,

red ruffed 4: 104, 105

96, 97,

9,

ruffed 4: 96, 104-105 sportive 4:

97

weasel sportive 4: 96 western woolly 4: 96

Lemur

catta 4: 96,

98-101

Leontopithecus L

6:

caissara 4:

88

/

chrysomelas 4: 88 hrysopygus 4: 86. 88

L

rosalia 4: 86,

L

i

leopard

88-91 30-33

2: 9. 10. 13,

black 2: 30, 31 clouded 2: 9, 10 2: 10, 13.

34-35

Lepilemur i

40

106-107 4:

106

slender 4: 106, 106

108-109 1 06

4: 106,

love charms Loxodonta

61

3:

L.

africana 5: 14, 16-21

L.

cyclotis 5:

charms

14

8: 67, 9:

79

Lutra

58-63

L.

lutra 1: 32,

L

mac uIicollis

L

sumatrana

1:

1:

32 32

Lutreolina crassicaudata 10: 14 Lutrogale perspicillata

1 32 Lycaon pictus 2: 50, 78-79 Lyncodon patagonicus 1: 32 lynx 2: 10, 12, 38, 40^1 :

:

8:

76-77

Canadian Eurasian 2

2: 40,

40

41

2: 41

Iberian 2: 40, 41

M Macaca

M M M M

luscata 4: 40, 48-49 nigra 4: 40. 52-53 \ilenus 4:

40

sylvanus 4: 40, 50-51 macaque 4: 8, 40, 42, 43, 66 Barbary 4: 40, 42, 5 0-51

black 4: 40, 52-53 Celebes (black) 4: 40, 52-53 4: -10, 47,

lion-tailed 4: 40,

48 49

41

Sulawesi crested (black) 4: 40,

52-53

Macrogahdia musschenbroekii

96 mustelinus 4: 96

edwardsi

albigena 4: 40

L

Japanese

melanistK 2: 30. 31

snow

9: 8,

68

logging 1: 51, 77, 2: 21; 6: 20 Lonchothrix emiliae 8: 30 Lontra L canadensis 1: 32, 64-67 L. felina 1: 32 Lophiomys imhausi 7: 66

lucky

96

bamboo

indri 4: 96,

12. 22,

4:

giant sloth 4:

mouse

92, 93, 93,

5: 12,

slow

dwarf 108

fork-marked

greater

Caribbean ground 109

Loris tardigradus 4:

eastern woolly 4: 96

golden

98-99

pygmy

4: 97,

fat-tailed

55, (61)

Litocranius walleri 6: 62,

loris 4:

Coquerel’s dwarf 4: 96 4:

18-19

puma

vexillifer 3:

L 'aterrimus 4:

brown 4: brown mouse crowned

6: 82; 8:

Lophocebus

black-and-white ruffed 4: 104,

:

locomotion, plantigrade

2: 72, 7: 13, 17,

bamboo

88 88

Asian 2: (15) Barbary 2: 17 mountain see

92

6:

lemming

dwarf

1:

104-107 Lobodon carcinophagus 3: 9, 36-37 lobtailing 3: 96, 109

Laurasia 9: 10

lemur

1:

llama

10: 74, (101)

lassa fever 7:

banded spotted

1:

10: 74, (101)

krefftii

1: 98 whale 3: 57, 92, 102, 108 Limnogale mergulus 9: 24 lice,

lizard,

Lasiorhinus

L

kuhm

Lipotyphla 9: 10

64-67 Malabar 4: 40

L. latifrons

80-81

86-89

8: 64,

88

leverets 8: Liberiictis

Upotes

48

1:

74-79

8: 64,

82-85

Lestodelphys hath 10: 14 Lestoros inca 10: 14

14-19

42

4:

Hanuman

L

122

L.

lion 1: 15: 2: 8-9, 9, 10, 13,

104-107

5: 92,

96-1Q1

ellipsiprymnus

K kob

82-83

guanicoe 5: 92, 108-109 L. pacos 5: 92, (106) Langorchestes

ringtailed 4:

ellipsyprymnus 6: 92-93

34 6

39. 8: 109, 9: 61, 77, 83,

glama

gray

ellipsiprymnus defassa 6:

1:42, 49

inbreeding

77

ellipsiprymnus 5: 75. 6: 62,

86 &Z. 97

ing partnerships 2: (60)

hutia 7: 12. 8: 11. 31,3/ Brown's 8: 30

2:

americanus

linsang 1: 89, 90, 91 African 1: 88, 90

lagotricha 4: 72,

:

implantation/fertilization.

Chilean 6: 10 Peruvian 6: 10. 12-13 human 4: 10-11, 17

66-67

western gray 10: 60, (62) kangaroo rat 7: 13, 17 bannertail 7 18-19 Ord's 7 24-25

92-97 kob 6: 62 Kobus

106

6: 62,

63 tree (Goodfellow’s tree)

koala 10:

walia 6: 106, 106-107 hneumia albk mda 1: 98

impala

Bennett's tree 10: 48 eastern gray (gray) 10: 48,

:

106, 107

Siberian 6: 106,

5:

.

6:

red 10: 25, 27

8: 62; 10: 8, (10),

keystone species 5: 26, 6: (68); 7 32, 93 khur see ass, Asian wild kiang see ass, Asian wild kinkajou 1: 20, 21 klipspringer 5: 8, 6: 60, 62

I

.iipine 6:

little

kangaroo 48-53

20 Kerodon rupestris

16-19

cristata 8: 12,

Przewalski's (Mongolian

kaluta,

8:

1

48

Lagothrix L. flavicauda 4: 72

lechwe

keratin 5: 10, 29, (37); 6: 60,

8-11, 28, 55

afncaeaustrahs 8: brachyura 8: 1

ibex

K

Karroo desert

Hystrix

5: (43)

jaguarundi 2: 10, 12 javelina see peccary, collared jerboa 7: 12, 17 jird see gerbil

Kannabateomys amblyonyx 8: 30

8:

Hystricognathi 7

60-61

L.

Lagidium peruanum 8: 30 iagomorphs 1: 10, 8: 61-63 Lagostomus maximus 8: 30 Lagostrophus fasciatus

Lariscus insignis 7:

:

103 yellow-spotted 8: 105

10. 11,

36-37

2: 10,

red 10: 11, 48, 54-59. 60,

04

western tree

draft 5: (43),

I

104-105

Hystncidae 7

5:

103

Matadi 8: 103 rock 8: 105, 106-107 southern tree 8: 102-103, 103

11

62

ancestral 1:

dawn

60,

65

60-63 Goodfellow's tree 10: 48, 6-67 gray 10: 48, 53, 60-63

104-105, 105, 106-107 eastern tree 8: 103

sapiens 1 : 14, 4: 12

9, 56,

8:

obliguidens 3: 55, 70-71 obscurus 3: 71

41,

53,

Bruce's yellow-spotted

honeybees 7: 1 honey guide, African 1 : 83 hoofed mammals 5: 8-13 Hoplomys gymnurus 8: 30 hornbill 1: 107 horns 5: 29, 31. 36-37, (37), 40, 41, 6:

36

102-105

Ahaggar

10

4:

lepidus 7: 36

8: 68,

5:

66 80-81

black 2: 37

:

84-85

pygmy

jaguar

10 48, 72 Hypsiprymnodontidae 10: 48 Hyracoidea 1: 10, 5: (12), 14 Hyracotherium 5: 42 hyrax 1: 10, 5: (10), (12), 14,

hog

83;

8: 62, 64, 66,

5: (11),

Hyperoodon ampullatus 3: 55, 90-91 Hypsiprymnodon moschatus

62

hirola 6:

13

9: 12,

H. spadiceus 7: 34,

H equmus 6: 62 H leucophaeus 6: H niger 6: 62

antelope

black-tailed 8: 64,

H. suillus 9: 12, 12

Hippotragus

1:

62-63

jackrabbit

sinensis 9: 12, 12

H

L

L.

black-backed

2: 50,

Hylopetes

hippotigres 5: 52

cruciger 3: 71

L.

jackal

alien 8:

L.

3:71

albirostris

L.

64

L.

acutus 3: 71

L.

L.

Jacobson's organ 2: 12;

hainanensis

Lepus

Lama

jaca see guinea pig

84-85

5: 74,

L.

8

1:

leptospirosis 1: 87; 7: 14

Lagenorhynchus

10:

72

J

36

Hylomys

66-67 pygmy 5: 66, 66-67, 67 Hippopotamus amphibius 5: 66, 68-73

for the Conservation of

see also tusks poaching 5: 20

H lar 4: 36, 38-39 H moloch 4: 36 H muelleri 4: 36 H syndactylus 4: 36

66, 66, 67,

5:

68-73

macrourus 10: 27, 46-47 pagurus 8: 30 IUCN see International Union /.

Isothrix

ivory

Hylochoerus meinertzhageni

common

Union for the

leprosy 9: (76)

Lepticidium

8: (41)

see also medical research Lagenodelphis hosei 3: 71

I.

104-107

5: 8, 9, 10;

8-1

10, 9:

ivory 3: 26, 5: 14, 20, 26,

Hylobates H concolor 4: 36

1

1: 8,

67 7: 15, 66, (75), 83, 86;

Nature

102, 103,

2:

60-61

3: 55,

Conservation of Nature (IUCN) 1: 16 introductions 1:18, 22, 43 lomys horsfieldi 7 36 Isoodon auratus 10: 27

102-103,

2: 102,

H. klossh 4:

48

8:

insectivores

108-109

103,

bisculus 6: 10

hippopotamus

Ima geoffrensis

Leporidae (leporids) 8: 60, 64,

laboratory animals 4: 40, 87;

:

1

1

H. c hapim 8:

9

7:

Hydropotes inermis 6: Hydrurga leptonyx 3: 9,

58-59 1

47

3:

L

4: (10)

International

Hydrodamalis gigas

Heterocephalus glaber Heterohyrax 8:

48-51

8: 30,

infanticide 1: (13); 2: 24;

interbreeding 2: 59, 6: 75, 107, 10: 41

:

H H

8: 56,

Hyaena H brunnea 2: 102 H. hyaena 2: 102, 104-107 hybridization 2: 59

4:

1:

88


SET INDEX

Macropodidae Macropus

10: 48,

M. martes 1: 32 M. pennant/ 1 50-51

50

60-63

64-65 M. rufus 1 0: 48, 54-59 Macroscelidea 1: 70; 9: 58 Macroscelides proboscideus

59

body

M. lagotls 10: 27, 44-45 M. leucura 10: 27 Madagascar, carnivores in

Madoqua M. kirku 6: 62 M. saltlana 6: 62 Makalata armata 8: 30

mammals

diversity of 1: (1

15-16 14-15

1:

1),

1:

8

origin 1:

reproduction

mammoth 5: Mammuthus manatee

12-14

1:

8

46-47

1: 70; 3:

:

5: (10), (12)

Amazonian

47

3:

47

West African

3:

West Indian

3: 46, 47,

60-61

4: 40, 42,

Mandrillus

M. leucophaeus 4: 40 M. sphinx 4: 40, 60-61 mangabey 4: 40, 42, 43 agile 4: 40, 42 black 4: 40, 42 gray-cheeked 4: 40, 40, 42 white 4: 40, 42

mangrove swamps 4: 71 Manis gigantea 9: 65 M. temminckn 9: 65 pi. tricuspis 9: 65 mara 8: 8, 30, 32-35

M

Marmosa mexlcana 10: 14 murina 10: 14

marmoset 92-93

4: 86,

86

87 marmoset family 4: 86-87 pygmy 4: 86 Geoffroy's 4: 86,

silvery 4:

86

M. dorothea 10: 14 M. fuscatus 10: 14 7: 76, 35,

36

alpine (European) 7 34, :

52-53 European

52-53

7: 34,

Marmota 7: 34 M. marmota 7 34, 52-53 M. monax 7: 34, 50-51 :

marsupials 1: 70; 10: 8-13 Australian carnivorous 10:

24-27

other plant-eating

74-77 marten 7: 36 American 1: 48-49 American pine (American) 1: 48-49 pine 1: 32, 35 10:

yellow-throated

Martes americana

M M

1:

32,

flavigula 1:

hispidus 8:

3: 56 Messelobunodon 1: 8

mesonychids

Metachirus nudicaudatus 10: 14 Metatheria 10: (10) miacid 1: 9 Miacoidea 1: 38, 98 mice see mouse

86

1:

32

Microcebus M. coquereli see Mirza

Microchiroptera 9: 80, 86 Microgale M. melanorrachis 9: 24 M. parvula 9: 24 Microperoryctes longicauda 10: 27

M

:

48-49

32

:

Japanese 9: 54 Juliana's golden 9: 40, 43 marsupial 10: (10), 26, 27,

40-43

9:

27

43

molehills 9:

8:

rat 7: 15, 18, 65; 8: 9

56

:

100-101

11, 16,

:

Allen's

swamp

4: 40,

41

black-handed spider 4: 72, 76-21 4: 72,

74-75

1

76-77 cheek-pouch 4: 40, 42 4: 72,

common

57 lesser blind 7: 100-101 Mechow's 8: 56, 57 mole rat family 8: 56-57 8: 56,

14; 7: 13;

1: (11),

Namaqua dune

58-59 8:

56

7 9; 8 56 southern dune 8: 56 mollusks 3: 25 Molossidae 9: 87 :

Monachus

:

schauinslandi

30-31

82-83 De Brazza's

42-43

4: 40,

40 4: 40

golden leaf golden snub-nosed 4: 40 green (vervet) 4: 40, 42,

:

44-47 guenon (vervet) 44-47

Hanuman

:

4: 40, 42,

langur 6: 13

83 Humboldt's woolly 82-83 leaf 4: 40, 40-42

4: 72,

:

:

Angolan

1:

98

banded

1:

78-79, 98, 1:

98

98

bushy-tailed 1: 98, 99 Bengal (Indian

common

108-109 gray (Indian gray)

108-109

Egyptian

98, 99, 99, 1:

98,

1:

.

shaker

7: (69)

18

64

singing 7: (69) spiny 7: 65, 8: 11, 12 three-striped marsupial

10 25, 27 waltzing 7: (69) :

western harvest 7: 80-81 West Indian key 8: 1

white-footed (deer) 7: 64,

82-83

wood

7: 13,

78-79

mouse-hare see pika mouselike rodents muktuk 3: 85 mulgara 10 27

7:

Mungos

72 72, 75

spider 4: 8, squirrel 4: 72,

M. gambianus 1 98 M. mungo 1 18-19, 98, 110-111

4:

:

:

78-79

swingers 4: 72 thumbless 4: 40

vervet

4: 40, 42,

Mungotictis decemlineata 1

44-47

yellow-tailed woolly 4: 72

3: 55,

1: 70; 10:

104

8: 12

9: 9, 11, 12,

9: 12

98

13

46-47 46-47

6: 12,

Chinese

98

monotremes, spiny

:

Muntiacus M. cri nitrons 6: 10 M. muntjak 6: 10 M. reevesi 6: 46-47

muntjac

domestica 10: 14 kunsi 10 14

Dinagat

16-19

:

84-85 Monotremata

108-109

7:

Japanese southern night

moonrat 1:

7: 12,

pygmy

multituberculates 7: 10

Monodon monoceros

108-109

pocket

snow see macaque,

99

Indian (Indian gray)

Indian gray

(vervet) 4: 40, 42,

:

98

giant-striped 1:

1:

-i

Monodelphis

106-107. Ill

Gambian

4: 72, 73, 84-85 northern night 4: 1? / 84-85 Old World monkey family 4: 40-43 72 owl (northern night) 4: 72, 73, 84-85 patas 4: 40, 4 7, 42, 43 proboscis 4: 40, 41, 70-71

woolly 4: 72

Indian (Indian gray)

108-109 1:

84-85

northern (northern night)

44-47

103-104, 110-111 broad-striped

gray) 1:

72-73

night (northern night) 4: 72,

savanna

(wood)

78-79 marsupial 10 (25), 27, 40 meadow jumping 7: 16 mole 7: 65 mouse family 7: 64-67 New World 7 14, 64, 65 Nicaraguan harvest 7 80 Old World 7: 14, 65, 65 pencil-tailed tree 7: 65 pig mouse 8: 38 plains 7: 65 :

4: 10, 72,

4:

long-tailed field 7: 13,

rhesus 4: 42

98

1:

1:

long-clawed marsupial 10 27

grivet (vervet) 4: 40, 42,

73,

Cozumel Island harvest 7: 80 deer 7: 64, 82-83 desert 7: 65 desert pocket 7: 22-23 field (wood) 7: 13, 78-79 hairy harvest 7: 80 harvest 7: 64 house 7: 13, 14-15, 65-66, 68-71 jumping 7: 17, 19 kangaroo 7 17

red howler 4: 72

Alexander's

brown

78

woolly

(Humboldt's woolly) 4: 72,

family

Angolan

3: 9,

:

squirrel 4: 72,

New World monkey

silvery

:

Costa Rican harvest 7 80

cloaked see colugo

Ehrenberg's 7: 18 giant

coarse-haired pocket (desertpocket) 7 22-23

(black-handed spider)

long-haired spider 4: 72

giant 8: 57

64

birch 7: 17

leapers 4: 72

57

8: 56,

72

7: 70,

African climbing 7:

mantled howler 4: 72 mustached 4: 40, 4 7

dwarf

106-107

monte 10

del

Damara 8: 56, 57 dune 8: 57

Microtus agrestis 7 92-93 migration 3: 82, 85, 87, 93-94, 103, (105), 109; 5 18, 109; 6 : 17, 23, 37, 84-85, 96, 101, 7 91,

milu see deer, Pere David's

6:

Moschus

(16)

howler

57

common

1:

:

Moschidae 6: 10 Moschiola meminna

99

44-47

see also shrew mole

1:

102, (103),

morrillo 8: 51

dusky-leaf 4:

48-51

common

:

99

98, 101

common

42-43 mole family

common

9: (83), 1

Grant's desert golden 9:

M. murina 10: 27 Micropotamogale M. lamottei 9: 24 M. ruwenzorii 9: 24 :

moose disease 6: 1 Mormoopidae 9: 87

98,

Central American spider

10-11,

mongoose

coquereli

M. murinus 4: 96 M, myoxinus 4: 96 M. rufus 4: 96

1

40 Grant's golden 9: 56-57 hairy-tailed 9 40, 42

9: 9,

5: (10)

8: 10, 56, 57,

M. humilis 4: 86 Micoureus M. alstom 10: 14 M. constantiae 10: 14

6:

1

mouse

naked 4:

shiras 6:

brown howler

40

Cape 7: 70; 8: 56 Cape dune 8: 56 30

1:

99

98,

Bolivian squirrel 4: 72

golden

44-47

40, 42,

blind 7 : 12-13, 15, 18;

:

1

golden mole family 9: 40-43

monkey

7:

Mesocricetus auratus 7 86-87

98 99

1:

14-15, 77

6:

eastern 6:

northwestern

Pousargues

9: 56-57 European 9:

Balkan blind (lesser blind)

30

family

:

73 14-19.

21-22

1:

yellow

9: 12,

6: 10, 12,

Alaskan

98-99

1

22-23

1

M. berezovskii 6: 10 M. chrysogaster 6: 1 0, 50-51 M. fuscus 6: 10 M. moschiferus 6: 1 mosquitoes 6: (22) mouflon 6: 62, 63

African 7 : 72; 8

8:

1:

9: 12,

Mindanao

moose

narrow-striped

monito

mole

mermaids 3: 53 Mesechinus M. dauricus 9: 12 M. hughi 9: 12 Mesocapromys auritus

M/co M. argentata

10: 9: 9, (10), 11

Persian 9: 40, 43 small-toothed 9: 40, 43 star-nosed 9: 40, 42,

88-89

mongoose

coast 9: 40, 41 desert (Grant's) golden

10:

M. macroura 1: 32 M. mephitis 1: 32, 84-87 Meriones unguiculatus

lesser 9:

98 98 long-nosed 1: 98 Madagascan 1: 98 marsh 1: 98, 99 Meller's 1: 98

white-tailed 1: 98,

northern marsupial 10: 43 northwestern marsupial

Microcavia australis 8: 30

Marmosops

marmot

98, 99,

Melogale personata 1 32 Melursus ursinus 2: 82 Menotyphla 9: 10, 58 Mephitidae 1: 32, 84

Mesomys

1:

greater

Liberian 1:

Selous' 1: 98, slender 1: 98

11, 40, 41, 42, (42)

82-83

:

mole

Jackson's 1:

ringtailed

Mirza coquereli 4: 96 moldewarp see mole,

giant golden 9:

100-105 Megachiroptera 9: 80, 86 Megaderma lyra 9: 98-99 Megadermatidae 9: 87, 98 Megaptera novaeangliae 3: 55, 102-105 Megatherium 9: 64 Metes metes 1: 32, 78-81

7

common

98, 99,

Mephitis

48-51

4:

:

:

Caribbean (West Indian) 3: 46, 47, 48-51

dwarf

41; 9: 91 Chinese 5 37

Mellivora capensis

8

5:

(97), 100, 6: 107, 7: 63, 8:

99, 100-105 slender-tailed (meerkat)

70 major groups 1:11

evolution

parts for 2: 21, 33,

1: 19,

32

3:

European

6: 51; 7: (31)

gray (meerkat) 1:19, 98,

8-17

1:

changing species

M M

M. leonina

t

meerkat 1:19, 100-105

96-97

32-35

:

see also laboratory animals;

medicine

Macrostylus 9: 86 Macrotis

mandrill

:

97

9: 77,

32, 35, 35,

1:

52-55. 62 7: 99; 9: 53 European 1: 54, (55) Miopithecus talapoin 4: 40 Mirounga M. angustirostris 3: 9,

trade medical research 4: 13, 32, 63, 78, 93; 7 15, (75); 8 (41 Y

surgery

90,

mink 1: 35 American

:

M. parryi 10: 48 M. robustus 10: 48 M. rufogriseus 10: 48,

1:

ringtail

masseter muscles 7 16-17, 64; 8: 12, 72 Massoutiera mzabi 7: 108 Mazama americana 6: 10 meat trade see bushmeat

M. fuliginosus 10: (62) M. giganteus 10: 48,

9:

miner's cat see raccoon,

:

6:

hairy-fronted 6: 10 Indian 6: 10

Reeves's 6:

46-47

Murexia M. longicaudata 10 27 M. rothschildi 10 27 :

:

123


SET INDEX

Muridae Murinae

65

7: 72, 7:

muriqui 4:

Neotragus

14 72

N. netscheri 8:

N. timminsi 8:

9:

7: 71

Mustela M. erminea

40-43 36

frenata 1: 32,

lutreola 1: 54, (55) nigripes 1: 32, 46-47

36-39 pu tori us 1: 32, 44-45 vison 1: 32, 52-55 mustelids 1: 32-35 nivalis 1: 32,

M. javanensis 1: 32 M. marchei 1 32 Myocastor coypus 8: 30, 44-47 :

Myocastoridae 7: 72; 8: 31 myoglobin 3: 1 1, 87 Myoictis melas 10: 27

Myomimus M. personatus 7: 102 M. roachi 7: 102 M. setzeri 7: 102

Myomorpha Myoprocta

7: (8)

30

exilis 8:

Myosciurus pumilio

Myosorex

Mexican mouse 10: 14, 77 murine mouse 10: 14

27 yvonneae 10: 27

34

9:

Myrmecobiidae 10: 24 Myrmecobius fasciatus 10: 27, 34-35

Myrmecophaga 9: 65,

tridactyla

68-71

Mysateles prehensilis 8: 30 Mystacinidae 9: 87

54 myxomatosis 8: 72, 93 Myzopodidae 9: 87 Mysticeti 3:

N Nandinia binotata 1 88 Nannosciuris exilis 7: 36 Nannospalax leucodon :

N.

narwhal

3: 55,

108-109

noise pollution 3: 88, 95

noolbenger see possum,

Virginia 10:

honey

water 10:

43 N. typhlops 10: 27, 42-43 Notoryctidae 10: 24, 26

wood

N. caurinus 10: 27,

10: 13, 24, 25, 26,

34-35 coypu nyala, mountain 6: 62 Nyctereutes procyonoides 2: 50 Nycteridae 9: 87 nutria see

pygmaeus

1

06,

4:

77; 2: 10,

72,

44-45

98 O. himalayana 8: 94 O. princeps 8: 94, 98-101 O. pusilla 8: 94 Ochotonidae 8: 60 Octodon bridges 8: 30 Octodontidae (octodonts) 7: 72; 8: 29-31

Octodontomys 8: 30 octodonts

gliroides

7: 72; 8:

29-31 30

Octomys mimax 8: Odobenidae 3: 8 Oddbenus rosmarus 24-29

70-71

N. larvatus 4: 40,

N. narica 1:

Odocoileus

hemionus Odontoceti

6: 10,

3:

34-37

0, (36)

1

54

58-59

1:

national parks/nature reserves 2: (15), 16, 31; 5: 34,

(102),

103

Neamblysomus julianae 9: 40 nectar 4: 105

Olallamys edax 8: 30 olingo 1: 20, 21 onager see ass, Asian wild Ondatra zibethicus 7: 96-97

Onychogalea

O O

fraenata 10: 48

unguifera 10: 48

opossum

1:

87

9:

Nemorhaedus goral

28

6: 62,

Alston's

mouse 10: 14 woolly mouse

10: 14

American

108

10:

14-17

Neofelis nebulosa 2: 10

bare-tailed woolly 10: 14

Neomys

black four-eyed 10: 14 black-shouldered 10: 14,

fodiens 9: 28

Neophascogale 10: 27

lorentzi

Neophoca cinerea 3: 9 Neotoma lepida 7: 84-85

16,

62

9: 65,

6: 62,

white (Arabian)

dammah

brown four-eyed

6: 62,

62 62

Otocyon megalotis 76-77 Otolemur

10: 14

bushy-tailed 10: 14

1

4:

34-35 12, 28-33

lesser (red) 1: 20, 21,

1:

2: (99) 20, 21, 30-31 :

:

2: (99)

panda bear see panda, giant pangolin 1: 70; 9: 64-67 African 9: 66 armored 9: 64 Asian 9: 66 giant 9: 65, 66 ground 9: 64-65, 65 tree 9: 65 panther

puma

1

1: 18,

giant river

Parascalops breweri 9:

Paroodectes

patagium 86

1:

35

river

(North

American river) 1: 32, 64-67 sea 1: 19, 32, 72-75 short-clawed 1: 32, 70-71 smooth-coated 1: 32 9:

1: 1

32,

1,

35

24,

25

25

Ruwenzori 9: 24, 25 ounce see leopard, snow

7: 34 Petrodomus tetradactylus 9: 59

P persephone 10: 48 P xanthopus 10: 48, 70-71 Petromuridae 7: 72; 8: 31 Petromus typicus 8: 30, 54-55 Petropseudes dahli 10: 74 pets

1: 22, 29, 35, 45, 95, 109, 2: 75; 4: 53, 78, 81,

83, 85, 87, 88, 101, 7: 15, (69), 83, 86, 8: 39, 42, (72), 9: 13, 21 see also cat, domestic; dog,

domestic Phacochoerus aethiopicus 5: 74

P africanus 5: 74, 80-83 Phalanger 10: 82

1:

40

9

Patagonia 5: 108 Pecan 5: 88 P tajacu see Tayassu

5: 5:

brush-tailed 10: 26, red-tailed 10: 25,

88, 89, 89 88, 89, 90-91

88-89 white-lipped 5: 88, 89, 89 Pectinator spekei 7: 08 peccary family

27

27

Phascogale

P calura 10: 27 P tapoatafa 10: 27 Phascolorarctos cinereus

92-97

Phascolosorex P doriae 10: 27

P dorsalis 10: 27 pheromones 7: 66, 70

P groenlandica 3: 9, 44-45 P. sibirica see Pusa sibirica P vitulina 3: 9, 40-41 Phoca rctos hookeri 3: 9 Phocidae 3: 8

Phocoena phocoena 78-79

5:

1

Pedetes capensis 7: 20-21 Pedetidae 7: 12 Pelea capreolus 6: 62 penguin 3: 38-39 Pentalagus furnessi 8: 64

Peramelemorphia 10: 24 Perameles

3: 55,

Pholidocercus 1: 9 Pholidota 1: 70; 9: 64 Phyllostomidae 9: 87 Physeter catodon 3: 55, 86-89 pig 5: 8, 10, 12

bearded bushpig

tajacu

peccary 5:8, 12 collared

Phaner furcifer 4: 96 phascogale 10: (25)

P andersom 10: 14 P opossum 10:14 Phoca

8: 108; 10: (76), 84,

Chacoan

Pgymnotis 10: 74 Phalangeridae 10: 76

Philander

Pasteurella tularense 8: 8

68-69

North American 1: 32, 64-67

spot-necked

88,

84

parasites 9:

1:

river

58-59 98

1:

48

32 1:

:

aplicalis 10: 27 P bilarm 10: 27 parapox virus 7: 44

32 Indian smooth-coated marine 1: 32 hairy-nosed

1

106

Cape Congo clawless 1: 32 European 1: 58-63 European river 1: 60

Petauroides volans 10: 74 Petaurus P breviceps 10: 74, 86-87

10: 74, 4: 40,

:

06

1

54-57 hamadryas

P.

35, 54; 8: clawless 1: 32

36 36

Pcarmelitae 10: 74

P leo 2: 10, 14-19 P leo persica 2: (1 5) P onca 2: 0, 36-37 P. pardus 2: 0, 30-33 P tigris 2: 10, 20-25 P uncia 2: 0, 34-35 Pantholops hodgsoni 6: 62

Parantechinus

1: 19,

7:

petaurista 7:

P.

P.

black 2: 30, 31 see also leopard;

94-95 P jerdoni 1: 88 P zeylonensis 1 88

2: 50,

P elegans

Petrogale

98-101

2: 82,

Paracynictis selousi

88

:

O. crassicaudatus 4:

shrew

12,

Petauridae 10: (76) Petaurista

P gracilis 10: 74 Petinomys crinitus

88

larvata 1:

Paradoxurus P hermaphroditus

Otariidae 3: 8

otter

30

44-45

Pan P paniscus 4: P troglodytes

P.

24

giant

28, 30,

1,

pantotheres 7: 10, 1 Papio P cynocephalus 4: 40,

O. leucoryx 6: 62, 90-91 Oryzorictes tetradactylus

otter

3:

6:

O. gazella 6:

9:

1

mountain 8: 30 pacarana 7: 72, 8: 28, 31 pademelon

1

90-91 Oryx O.

paca

1

giant 9: 24,

77

8: 64,

90-91 scimitar-horned 6: 62

northern

agile gracile

Nectogale elegans

68-73 oryx Arabian

7: 72; 8:

P broad bent 10: 27 P raffrayana 10: 27 Peroryctidae 10: 24

Panthera

Orycteropus afer

O. garnettii 4:

okapi 6: 52, 52-53, 58-59 Okapia johnstoni 6: 52,

20

20, 28-29 Nasuella olivacea 1: 20 Natalidae 9: 87

nasua

6:

106-109

Osbornictis piscivora 3: 9,

22, 34, 52, 58, 85, 89, 91, 101

Nasua

orca 3: 55, (57), 62-65 Orcinus orca 3: 55, 62-65

1

Peroryctes

P

4: 8, 10, 12, 13,

Oryctolagus cuniculus

94

O. collaris 8: 94,

06 Peromyscus P leucopus 7: 83 P maniculatus 7: 82-83

7: (14)

30-31

78-79

O. alpina 8:

43 Perodicticus potto 4:

:

red

14-19 Bornean 4: 12, 14-15, 18, 78-79 Sumatran 4: 12, 14-15, 18

Ochotona

40

mouse

10: 14 woolly 10: 16

10: 1:

spirit gracile

7: (31) Perissodactyla 1: 70; 5: 10, 12,

62

oxpeckers 5: 38; 6: (73) Ozotocerus bezoarticus 6 10

giant

oribi 5:

o

33

panda

7

Oreotragus oreotragus 6: 62 7 7; 6: 67, 62 Ornithorhynchus anatinus

06

1

7

white-eared 10: 14

Oreamnos americanus 108-109

Nycticebus N. coucang 4: 0. 0. 108-109

8, 13, 14, 15,

13, 14, 16,

orangutan

6:

Paguma

18-23

16, 17,

Notoryctemorphia 10: 24 Notoryctes

musimon 7: (37); 9:

snowy

short-tailed 10: 14 :

oil 3:

N. concolor 4:

pygmy

shrew 10 16 silky shrew 10: 14

virginianus 6:

84-85

Patagonian 10: 14

Noctilionidae 9: 87

100-103

red-legged 10: 48, 57 red-necked 10: 48 Pagophilus groenlandica

14

pallid fat-tailed 10:

108

N. albiventris 9:

N.

mouse

10: 14

Noctilio

100-101

Nasalis

pale-bellied woolly

N. timealeyi 10:

ocelot

28 104-105

varius 9:

Myotis lucifugus Myoxidae 7: 12

7:

28

southern 10: 27 Ningaui

numbat

Mydaus

mouse

ningaui 10: (24) Pilbara 10: 25, 27

N. leporinus 9: 1: 32, 36,

O.

gray four-eyed 10: 14, 77 gray short-tailed 10: 14 Incan shrew 10: 14 lutrine 10: 14

7:

62

O. aries 6: 62

owl

shrew 10: 14

gray-bellied slender

Ovis O. canadensis 6:

10: 14, 75

membrane

nictitating nilgai 6:

M. porcellus 8: 38 Muscardlnus avellanarius 7: 102, 106-107 muSk 1: 85 6: 51 muskox 6: 62, 104-105 muskrat 7: 13, 65, 96-97 common 7: 96-97 mustang 5: 42, 58-61

7:

gray-bellied

10: (80)

M. musculus wagneri

mouse

10: 14 elegant fat-tailed 10: 14

Zealand, mammals introduced into 1: 43;

71

18-23

13, 14, 15, 16, 17,

54-55

P bougainville 10: 27 P nasuta 10: 27 Peramelidae 10: 24 perfume industry 1: (91);

104-105 8'

(Virginia) 10:

Dorothy's slender

New

M. musculus musculus 7:

64 64

Neurotrichus gibbsii

musculus domestlcus

7: 71

124

Chilean shrew 10: 14

common

68-71

7:

7: 71

N

62

6:

brevirostris

Ourebia ourebi 6: 62 Ovibos moschatus 6: 62,

10: 14, 77

Nesolagus

M. musculus M. musculus

M. M. M. M. M. M.

pygmaeus

N.

Mus

M‘.

Central American woolly

62

N. batesi 6:

75 75

5: 74, 5: 74,

lard pig 5: (78)

pig family pig fish 3:

pig

mouse

pika

5:

74-75

78 38 62

8:

8: 60,

Alpine 8: 94

American

8: 67, 94, 96,

98-101 Asian 8:

97

collared 8: 94, 97,

common

98

(American)

96, 98-101

8: 94,


SET INDEX

North American

Eurasian 8: 98, 100

Himalayan 8: 94, 96 pika family 8: 94-97 Rocky Mountain (American)

Old World 12-15

8: 94, 96, 98-101 Russian steppe 8: 94, 97

tree

8-13

mammals

possum 10: 8, 8:

30

10: (10), 75-77 Arnh5mland 10: 80

brush-tipped ringtail 10: 74 common brushtail 10: 70, 74, 76, 78-81

common copper

27

P maculata 10: 27 P novaeguineae 10: 27 Platanista

green

P gangetica 3: 55, (61) P minor 3: 55, (61) platypus 1:14

honey

106-109 1

1

10-1

1

Pliohippus 5: 42

74

ringtail 10:

39,

5:

10: 74, 76,

ringtail 10: (77)

74 74 76

scaly-tailed 10:

P aureospinula 9: P truei 9: 12, 13 :

striped 10: 74,

1

Weyland

32 1

:

32

ringtail 10:

:

Potamochoerus P larvatus 5: 74 P porcus 5: 74 Potamogale velox

9:

pollution

61, 62, 64, 69,

Potamogalidae

1

71, 75; 2: (88); 3: 12, 41,

59, 61, 65, 67, 74, 79,

83

95

Pongo P abelii 4: 12, 14-19 P pygmaeus 4: 12, 14-19 Pontoporia blainvillei 3:55, (61)

pony Falabella 5: (43) feral

see mustang

42-43 wild see mustang porcupine 1: 50-51; 8: 10, 12-15 Welsh

5:

African 14,

8: 10, 12,

12-13,

16-19

American American

8:

10

tree 8: 9

bicolored tree 8: 27

13 brown hairy dwarf 8: 12 brush-tailed 8: 14, 16 Cape 8: 12, 75 Brazilian tree 8: 12,

crested (African) 8: 10, 12,

12-13, 14, 16-19 Europe 8: (19) hairy dwarf 8: 12 in

long-tailed 8: 12, 14, 16

Malayan 8: 12, 72, 14 Mexican tree (tree) 8: 12,

26-27

New World

7: 72;

12-15

North African (African) 8: 10, 12, 12-13, 14,

6-19

72-73

10: 48, 50, 51,

Potorous

P longipes 10: 48 P tridactyl us 10: 48, 72-73 Potos flavus 20 potto 4: 106, 106-107 golden 4: 106 Martin's false 4: 106 pouches 10 (10) prairie dog 1: 46-47; 7: 13, 15, 18, 36 black-tailed 7: 34, 56-59

56-59 Utah

7:

58

white-tailed 7: 58 Presbytis 4: 41

higher 4:

1

lower

1

4:

1

,

1

06-1 07 9: 65

Prionodon P linsang 1: 88 P pardicolor 1 88 :

Proboscidea 1: 10, 5: 14 Procapra picticaudata 6: 62 Procavia capensis 8: 103,

106-107 Procaviidae 8: 102

Procolobus 4: 41-42 P badius 4: 40 Procyon

P cancrivorus P gloverellani

1: 1:

20 26

rock see pika

pseudantechinus Woolley's 10: 27

Pseudantechinus P macdonnellensis 10: 27 P woolleyae 10: 27 Pseudocheiridae 10: (76) Pseudocheirus peregrinus

74

P archeri 10: 74 P cupreus 1 0: 74, 90-91 Pseudochirulus caroli 10:

74

P cinereus 10: 74 Pseudois nayaur 6: 62 Pseudopotto martini 4: 1 06 pseudoruminants 5: 70 Pseudoryx nghetinhensis 6: 62 Pteronura brasiliensis 1: 32,

vlei 7:

Pudu P mephistophiles

6:

1

1

42-43

2: 10, 13,

punare 8: 30 Pusa sibirica 3: 9 Pygathrix roxellana 4:

Q

quagga

5:

45,

40

48

see porcupine, North

American quills 8: 12, (14),

16-17, (21)

see also spines 10: 48, 52,

68-69

(25), 27 Guinea 10: 25, 27 northern 10: 27, 32-33

quoll 10:

New

raccoon Barbados

1:

26

common

1:

20, 21,

22-27 26

Island 1:

20 1:

Rangifer tarandus 6:

desert 10:

musky

1

R.

1:

Amami

43; 8: 61, 62 8: 62,

64

American swamp 8: 66 annamite 8: 64 brush 8: 64 bunyoro 8: 63, 64 bush see hare, snowshoe chocolate Dutch 8: 72-73 cottontail 1: 87 desert 8: 66

76-77

rattus 7:

giant R. affinisi 7:

0,

Raphicerus campestris 6: 62 rat 1: 15; 7: 12 African marsh 7:

65

Amazon bamboo

8:

36 36

.

7:

30

33

red ape see orangutan

Red

30 Asian climbing 7: 64 Atlantic bamboo 8: 30 Australasian water 7: 65

Lists of Threatened Species (IUCN) 1: 16

Redunca R. arundinum R.

Australian water 7: 9, 17,

65 8: 31

black (ship) 7: 11, 13,

14-15, 72, 75, 76-77

30 broad-headed spiny

bicolor 7:

R indica 7: 34, 62-63 recolonization 3: 14, 35,

8:

62

6:

fulvorufula 6: 62

R. redunca 6: 62 reedbuck bohor 6: 62 mountain 6: 62 southern 6: 62

reestablishment 3: 31, 93

30 14-15, 72-75 8:

.

77; 8: 10; 10:

72

rufous 10: 48, 57 see also potoroo Rattus R. norvegicus 7: 72-75

R.

7:

48

10: 48,

Ratufa

20-25

brown

74

see badger, honey rat-kangaroo 10: 51, 52 ratel

20-21

20, 20, 21, 21

armored 8: 30 armored spiny

rat;

ratufa see squirrel, Indian

refection 9: 29, 29 reforestation 4: 91

89 20-25 22-23

regeneration, of land 10:

1

brush-furred 7: 65 cane 7: 72; 8: 31

reindeer

Central American climbing

reintroduction 1: 46; 2: 41,

7:

reindeer

64

7:

64

72-75. 77; 8: 10; 10: 11 dassie 7: 72; 8: 31. 54-55 desert wood 7: 84-85 golden-backed tree 7: 67 greater cane 8: 30 greedy olalla 8: 30 (ship) 7:

7

7,

13,

14-15, 72, 75, 76-77 Indonesian key-footed 7: 64

Kenyan crested 7: 66 lesser cane 8: 30 Natal

multimammate

New World

7:

7:

65

64

Norway (brown) 7: 14-15, 72-75 77; 8: 10; 10: Old World 7: 14, 65 1

Owl's spiny 8: 30

6:

R.

hirsutus 7: 81

R megalotis 7: 80-81 R. paradoxus 7: 81 R. rodriguez 7: 81 R. spectabilis 7:

religious

81

ceremonies

5:

relocation 5:

34

repopulation reproduction

1:

61, 75

1:

12-14

25

see also embryonic diapause; implantation/fertilization;

breeding rhebok, gray 6: 62 Rheithrosciurus macrotis 7: 36 rhinoceros 5: 8, 8, 9, 11, 12 black 5: 28, 33, (34), 36-39 giraffe 5: 8

greater Indian (Indian) 5: 28,

painted tree 8: 30 Panama spiny 8: 30, 31

plague (ship) 7: 7 7, 13, 14-15, 72, 75, 76-77 plain brush-tailed 8:

plains viscacha 8: 7:

7: 29, 107 Reithrodon tomys

30-35

84-85

pouched

90-91,

45, (66), 91,

grass (white) 5: 28, 29,

pack (desert wood) 7:

moss

5: 34; 6:

Chilean rock 8: 30 chinchilla 7: 12 common (brown) 7: 14-15,

house

6: 10, 12,

(56), 69; 4: 87, 88,

.

R rabbit

103

rat-bite fever 7:

Central American vesper

P pudu 6: puku 6: 62

puma

79; 7: 14, 74; 9: 84, 97,

raccoon family

rat;

mole

rat;

viscacha

bristle-spined 8:

Pteropodidae 9: 86 Pteropus 9: 86 P giganteus 9: 88-91 Ptilocercus lowu 9: 59 pudu 6: 10, 12 northern 6: 10 southern 6: 10, 12

water

vole,

kangaroo

8:

1:

30

65

see also chinchilla

rabbit

bamboo

wood)

84-85

viscacha rat 8: 29, 30

62

ringtail 1:

64

water spe

crab-eating

13, 14-15, 72,

7,

tuft-tailed spiny tree 8:

volcano 8: 63, 64 see also cottontail

Cozumel

27

7

76-77

smooth-tailed giant 7: 65 South American climbing

7:

64-66 8:

7:

75,

hare,

rabies 1: 26, 87, 2: (67), 77,

110-111

ship

trade (desert

snowshoe see snowshoe

10: 28, 32, 37, 43, 71, (95) Proteles cristata 2: 102,

quokka

Priodontes maximus

64

14-15,

spiny 7: 72; 8: 31 spiny tree 8: 30

hemorrhagic disease 72 rabbit warrens 8: 70-71

quillpig

P comata 4: 40 P femoralis 4: 40 primates 1: 10, 4: 8-1

riverine 8: 62,

13,

7 7,

72, 75, 76-77 rufous tree 8: 30

7:

64-67

8:

64-67

29

rock 8:

8: 64, (93)

rabbit family

100; 3: 12, 19, 59, 74, 107, 5: 41, 60; 9: 85,

68-69

long-footed 10: 48 long-nosed (potoroo)

60-61 62, 64,

8:

66 marsh 8: 65 Mexican volcano 8: 64 Old World (European) 8: 60-61, 62, 64, 68-73

swamp

35, 39, 44, 49, 55, (97),

73

plains (black-tailed) 7: 34,

Asian brush-tailed 8: 12 bahia hairy dwarf 8: 12

13, 14,

9:

48, 50, 51,

:

7: 8;

African brush-tailed 8: 12

8:

potoroo 10: 72-73

24

roof (ship) 7:

8:

62, 69, 74; 2: 21, 28, 33,

P

Virginia

Gilbert's 10:

noise pollution 3: 88,

74

European 68-73

Sumatran 8: 63, 64 Sumatran short-eared

Pseudochirops

see also opossum,

Poelagus marjorita 8: 64 Poiana richardsom 1 88 polecat 1: 44-45 European 1: 32, 34, 35, 35 marbled 1 32, 34, 34 :

8

P diadema 4: 96 P tattersalli 4: 96 P verreauxi 4: 96

10:

western pygmy 10: 74

Poecilogale albinucha

1:

fat-tailed 10: 25,

mountain brushtail 10: 74 mountain pygmy 10: 74,

rat family 7:

pygmy

Prototheria 10: 104

rock ringtail 10:

(102)

Podogymnura

:

Propaleotherium Propithecus

75-76,

10: 8, 74,

76, 77, (77)

poaching 4: 24, 27,

63

6:

P crassicaudatus 8: 64 P randensis 8: 64 P rupestris 8: 64

ringtail 10: 74,

marsupial gliding 7: (37)

Plecotus auritus 9:

1

110-111 American 2: 26, pranking 6: 97

74, 76

ringtail 10:

102-103 Lead beater's 77, 88-89

duck-billed 10: 105,

6: 60, 62, (63),

protection, legal 1: 35, 47, 51,

Daintree River ringtail 10: 74 eastern pygmy 10: 74 feathertail 10: 74

Planigale

Poecilictis libyca 1

pronghorn

90-91

27

platyrrhines 4:

78-79

55,

porpoising 3: 69

13 Plagiodontia aedium plague 7: 14, 76-77 planigale 10: (24) common 10: 25,

(harbor) 3: 55,

domestic (European) 8: 60-61, 62, 64, 68-73

forest 8:

Prolagus 8: (96)

Pronolagus

78-79 harbor 3:

1,

10:

78

3:

30

8:

26-27

8-1

common

Papuan

19

8:

8: 12, 13, 14,

porpoise

P albicans 4: 72 P irrorata 4: 72 P monachus 4: 72 P pithecia 4: 72

P lotor 1: 20, 22-27 P pygmaeus 1: 26 Procyonidae 1: 20-21

ÂŤ

Proechimys semispinosus

Porcus piscus

87

Pithecia

9-1

8 );

7: 72, 8:

South African

8:

pinto 5: 59

placental

(

20-25

upper Amazon 8: 12 porcupinelike rodents

Sardinian 8: (96) pingers 3: 79 Pinnipedia 1: 70; 3: pipistrelle 9:

7:

8: 10, 12, 13, 14,

18

30

30

40-41 Indian 5: 28, 40-41 Javan 5: 28 northern white 5: 34 rhinoceros family 5:

28-29

southern white

5: 34,

35

125


SET INDEX

square-lipped (white) 5: 28,

30-35 Sumatran 5: 28, 29 white 5: 28, 29, 30-35 woolly 5: 28 29,

Rhinoceros R. sondaicus R.

28

5:

40-41

unicornis 5: 28,

106-107

Rhinopithecus roxellana see Pygathrix roxellana

Rhinopomatidae Rhynchocyon

9: 59,

59 R. petersi 9: 59 Rhynchogale melleri 1 98 Rhyncholestes raphanurus :

vulgaris 7: 34,

7:

46-47 42-45

1-12 mouselike 7: 16-19 porcupinelike 8: 8-1 squirrel-like 7: 16-19 Romerolagus diazi 8: 64 rorquals 3: 55, 106, 107 1

2, 6:

1

pseudoruminants Rupicapra R. pyrenaica

6:

70

5:

R. rupicapra 6: 62, Rupicaprini 6: 108

9-10,

10;

86

3 8-13 64 :

94

oedipus 4: 86 saiga 6: 62 Saiga tartarica 6: 62

.

4:

6:

1

08 6: 6:

sewellel 7: (29)

4: 72,

sheep

73

:

21 (32), rat,

naked saola 6: 62 28-31

1

:

70;

Scapanus orarius

9 58 :

9:

40

4:

96

Sirenia 1: 10; 3: 47, 5: (12)

patches 4: 39, 52, 58

Island

Soay

6:

shrew

1:

American water

9:

36-37

62,

87, (88), 91

pygmy

7:

black-striped 7: colonial

7:

S.

hippurus 7: 36

5.

lowii 7:

5.

tenuis 7:

ground

Supercohort 41

brown-throated three-toed (three-toed) 9: 65, giant ground 9: 64

72-73

banded

65

34

36

38

testing surgical techniques 4: 13

38-41

antelope

36

62-63 Mindanao

flying 7:

8: 64, (93)

5.

bachmam

8:

S.

dicei 8: (93)

8: (93)

64

S.

floridanus 8: 64,

S.

insonus 8: (93)

S.

palustris

hefnen

5. transitionalis 8:

90-93 8:

64

(93)

symbiosis 6: (73) 6: 62, 70-73 syndactyly 10: 75, (94)

Syncerus caffer

Tachyglossus aculeatus 10:

9:

110-111

100-103

Himalayan 6: 62 takh see horse, Mongolian

red 7: 41

takin 6: 62,

wild

flying 7: 34,

36

36

S.

virginiae 10:

9

audubonn

tahr,

South African ground

9:

aquaticus

S.

7: 34,

36 36 Prevost's 7: 36 pygmy 7: 36

murina 10:

Solenodontidae sonar 3: 83

S.

plantain 7:

gilberti 10:

Solenodon S. Cuban us 9: 9 S. paradoxus 9: 9

6: 62,

80-81

pale giant 7:

scaly-tailed 7: 12, 17, (19)

36 7: 10,

36-37 southern flying 60-61

54-55 three-striped ground 7: 36 tree 7: 17, 34, 35, 36 tufted ground 7: 36 squirrel-like rodents 7: 16-19 stances 5: 9-10

108

43

talapoin 4: 40, 47, Talpa T.

europaea

T.

street 9:

9: 40,

44-47

40

Talpidae 9: 9

tamandua 7: 34,

spotted giant flying 7: 36 squirrel family 7: 34-3 7 thirteen-lined ground 7: 34,

74

3:

grimmia

Tadarida brasiliensis

S.

9

Sylvicapra

34

northern flying 7: 61 northern striped palm

S.

9:

.

Indian giant 7: 34, 62-63 Javanese flying 7: 36 Low's 7: 36 Malabar (Indian giant) 7: 34,

slender 7:

Cuban

ground

8-9

7:

dolichura 10:

1

74

squirrel, thirteen-lined

(37); 10: (76)

crassicaudata 10: 27

8: (78)

5:

scrota 5: 74, 76-79 suslik, thirteen-lined see

Sylvilagus

S.

9: 8, 9, (10),

barbatus

S.

flying 7: 8, 13, 17, 34, 37,

red giant flying 7:

(102), 109; 9: 16

meerkat

swimming therapy

7: 34, 36,

98,

:

S.

36

7:

S.

1:

1

100-105 suricate see

36 36-37

aitkeni 10:

27

Suricata suricatta

Sus

S.

38 38 38-39

human

eastern fox 7: 34, 46-47 Eurasian red 7: 42-45 European red 7: 34

red-cheeked

Sminthopsis

40 40

4:

surgery animal parts for surgery 5: 79

35 9:

10

1:

grizzled 4:

60-61

flying) 7: 34,

36 36

sureli 4:

eastern flying (southern

horse-tailed 7: 1:

Hispaniola 9: 9

30-33

8: 12 8-9, 61

Cape ground

38-39

9: 28,

Sundasciuris

black flying 7: 34, black giant 7: 36

Harris'

)

solenodon

10; 9: 8, 9, (10), 11

6:

5: (12)

bottom" see whale,

blue

43-44, 47 gray-cheeked flying 7: 36 ground 7: 18, 19, 34, 36

32 striped 1: 32, 84-87 western hog-nosed 1: 32 western spotted 1: 32 sloth 9: 64-65, 65-66, 66, 66

snowshoes

African forest 9: 28 American short-tailed 9:

skunk 1: 34 Andes 1: 32 hooded 1: 32 pygmy spotted

snares 4: 24, 53

63

pelengensis 10: 74

Suncus etruscus

springhare 7: 12, 18, 20-21 spy-hopping 3: 71, 96, 107

gray

see also fur trade

snakes

barbary 6: 62 blue 6: 62 desert bighorn 6: 100 domestic 6: 62, 63 wild 6: 63

10: 27,

laniarius ( harrisii) 10: 27, 28-31

Scandentia

39

Orkney

Sarcophilus S. harrisii

11;

6: 60-63 American bighorn 6: 100-103

monk 4: 72 white-nosed 4: 72 Salanoia concolor 1 98 poisonous 9: 8, 11, sand puppy see mole

1: 15; 5: 11,

celebensis 10: 74

S.

.

three-toed 9: 65, 72-73 Smilodon fatalis 2: 1

68-69

72

32 16-17

American gray (gray) 7: 34, 36, 38-41 43-44, 47 American red 7: 42 Arctic ground 2: 65

three-toed 9: 65 southern two-toed 9: 65

24

1:

mammals

African

maned

46

S.

"sulfur

squirrel 7: 11, 12, 13, 15 African ground 1: 101

Simias concolor see Nasalis concolor

(

pygmaea

3:

27

styloglossus 5: (12)

32

spines 9: 13, 15, see also quills

squid

Hoffmann's two-toed

62 62

Setonix brachyurus 10: 48,

buffy 4: 72

saliva 9: 17,

40 64-67

hypoleucos 4: 40 S. obscurus see Trachypithecus obscurus sengi 9: 58

Setifer setosus 9:

Guianan

18-19

S.

servalines 2:

bald-faced 4: 72

40

Sigmoceros lichtensteinii see Alcelaphus lichtensteinii Sign Language, American ~ 4: 13, (16), 27 simakobu 4: 40

12

9,

serval 2: 10, 46-47

78-79

saki

4:

inquisitive 9:

shrew mouse 8: 102 branch 8: 102 variable 8: 102 siamang 4: 8, 36, 37 Siebenschlafer 7: 104

3: (59),

Strigocuscus

subungulates

springbok 96-97

32

on shore 66, 67, 96

strandings,

lower

springhaas see springhare

54-55

111. 3: 12, 15; 5:91; 6: 66; 10: 56, 91

geei see Trachypithecus geei

Japanese mainland

72

42

Gibb's (American) 9:

2: 33, 36, 37, 39, 47, 100,

entellus 4: 40,

serow

Saimiri

54-55

9:

the stomach 3: (27), 91

stridulation 9:

1

kraemeri 10: 74 maculatus 10: 82-83 papuensis 10: 74

spiny

1: (74);

31-32, (31)

stotting 6: 35, 97; 8:

1

Spilogale

5.

skin trade 1: (23), 69, 72, 74;

3: 9, 13, 18,

Serengeti Plains 6: 84, (85)

S.

28-29

9:

Tibetan water 9: 28 see also elephant shrew; otter shrew; tree shrew shrew mole 9: 40, 41

sitting

S.

94

9: 8,

California

102 Semnopithecus

5 imperator subgrisescens

bearded

32

7:

imperator 4: 86, 94-95 imperator imperator

boliviensis 4:

3:

fur) 3: 8,

vestitus 8:

S. gracilis 1:

diademed 4: 96 golden-crowned Verreaux's 4: 96

3: 9, 12,

5.

5.

pygmy white-toothed

Australian 3: 9, 11

S.

sc iu re us 4: 72,

piebald 9: 28

3: (27), 91

strepsirhines see primates,

insidiosus 8:

S.

hammers

in

3: 86, 87, 89, 91

S.

5.

sifaka

Selvinia betpakdalaensis

Saguinus

S,

9,

sea leopard see seal, leopard 1:

30-33

tailed 9: 28,

40-43 stomach stones

4:

Spilocuscus

northern (American) short-

20-23

American 1: 48, 50 sacred animals 4: 59, (66); 9: 91

S.

13

16-17

Steller's 3: 9, 12,

4:

.

1

Galapagos 3: 9 New Zealand 3:

sable,

28

shrew family

9

86-87

108

s

5. fusciollis 4:

least 9:

Asiatic 9:

3:

28

Indian house 9:

76-77 stoat 1:19, 32, 34, 36,

54-55

7: 34,

62

6: 67,

Stenella longirostris 3: 55,

as

Sphiggurus

8-9

gray 3: 42-43 harbor 3: 8, 9, 40-41 harp 3: 9, 12, 44-45 Hawaiian monk 3: 9, 30-31 leopard 3: 9, 11, 37, 38-39

sea lion

62

9:

American

44-45 southern elephant

8-9

28

greater white-toothed

36-37

3: 9, 11,

ringed 2:

spermaceti

steenbok

stones

Spermophilus tridecemlineatus

forest 9: ~

36-37

palustris 9:

Soricidae 9: 9, 58 Spalacopus cyanus 8: 30 Speothos venaticus 2: 50

9: 28,

29

fur 1: (11)

9, ,

34-35

pygmy

songs gibbons 4: 36, 39 whale songs 3: 99-100, 105 Sorex S. araneus 9: 34-35 S. minutus 9: 28 5.

28

southern fur (Cape 1 1

common

29

46, 47, 47, 53

saddleback (harp)

92-93

38-39

Eurasian water 9: 28,

(harbor) 3: 8, 9,

3:

29

10

9:

Eurasian 9:

3: 8, 9, 9, 10, 11,

phocid

Rousettus aegyptiacus

4:

1: (11), 9:

Eurasian

northern elephant 3: 12, 32-35 northern fur 3: 9, 12, 14-15

1

5:

38-41

scorpion 1: (102) Scutisorex somerem 9: 28 sea canary see beluga sea cow

monk

evolution 7: 12

126

niger 7: 34,

S.

eared

Rocky Mountain tick fever 7: 14 rodents 1: 10; 7: 8-1 cavylike 8: 8-1

S,

S.

40-41 crabeater 38

raccoon rockhare see hare, greater red rockhare

S.

carolinensis 7: 34,

common

10: 14

rumen 6: 8 ruminants

elephant Etruscan

circus tricks 3: (23)

Rhynchomeles prattorum 10: 27 rinderpest 6: 73, 77, 85 ringing of trees 7: 37 ringtail see under possum;

S.

5.

:

R. cirnei 9:

9:

29

28 Etruscan white-toothed

see also dugong sea grass 3: 46, 52 seal 1: 10; 3 8-13. 64; 7: 12 Antarctic fur 3: 9, 12 Baikal 3: 9, 10-11 Cape fur 3: 8, 9, 16-17 Caribbean monk 3: 9

62-63

extinct 7:

9: 28,

desert 9:

9: 2ÂŁ,

Steller's 3:

86

9:

R.'chrysopygus

expansion

armored

Sciurus

Rhinocerotidae 5: 28 Rhinolophidae 9: 87 Rhinolophus hipposideros 9:

Scelidotherium 9: 64 Sciuridae 7: 12 Sciurognathi (sciurognaths) 7: (8), 12, 16-19; 8: 8 Sciuromorpha 7: 8

9:

southern

9:

68 65

Tamandua tetradactyla tamaraw 6: 62

9:

65

tamarin

4: 8 bearded emperor

4:

94

black-chinned emperor 4:

94

black-faced

88 88

lion 4:

black lion 4: 86,

cotton-top 4: 86,

87

94-95 golden-headed lion 4: 88

emperor

4: 86,


SET INDEX

golden

thylacine

lion 4: 86, 87,

saddleback 4: 86, 95 tamarin family 4: 86-87 Tamias sibiricus 7:

T.

striatus 7: 34,

48-49

tapetum lucidum 2 10-12; 4: 85, 106, 107 :

tapir

12

5: 8, 11,

62 63

Baird's 5: 62,

Brazilian

T.

64-65 63

Malayan 5: 62, 63, mountain 5: 62, 63 South American (Brazilian) 5: 62, 63, 64-65 tapir family 5: 62-63

Tapirus

bairdu 5: 62

T.

indicus 5: 62

T.

pinchaque

T.

terrestris 5: 62,

107 106, 107

tarsier 4: 11, 106,

4:

western

107

4: 106, 107,

102-103 Tarsius T.

bancanus

4:

T pumilus 4: 1 T. spectrum 4:

06 06 06 1

1

Tasmanian devil

10: (24), 27,

T.

T.

(16),

derbianus gigas oryx 6: 62, 76

Taxidea taxus

1:

32,

76

6:

76-77

pecan

5:

88

I tajacu

5:

88, 90-91

T.

Tayassuidae

5:

tayra 1: 32; 4: teeth aardvark 9: 65, (66), bats 9: (85), (95)

selenodont 1: 32

6:

78-79

tenrec

8: 12; 9: 8, 9, 11

aquatic 9: 24, 25, 25 common 9: 24, 26-27 four-toed rice 9: 24 greater (common) 9: 24, 9: 24,

large-eared 9: 24,

25

hedgehog

rice 9:

9:

9: 10,

25

25

24-25 tailless (common) 9: 24, 26-27 tenrec family 9: 24-25 streaked 9: 24,

T.

T.

T.

ecaudatus

26-27 24

Tenrecidae 9: 9, 1 1, termite mounds, as vantage points 6: 88, 88-89 termites 2: 50, 76, 1 10-1 11, 10:

34

62 Theropithecus gelada

4: 40,

Thomomys 7:

talpoides

26-27

Thrichomys apereoides

Thryonomyidae Thryonomys

8:

7: ?2; 8:

T gregorianus 8: 30 I swinderianus 8: 30

30 31

variegata 4: 96,

8:

79 Trypanosoma cruzi tsessebe 6: 88

8:

plains 8:

1:

70; 5: (12);

88 V' tangalunga 1: 88 \/ zibetha 1: 88 Viverricula indica 1 88 Viverridae 1:18, 88, 98 vole 7: 1 3, 7, 65 bank 7: 74-75 field 7: 92-93 1:

9-10,

tularemia 8: 81, 93 3: 69, 77 tunnels, foraging 7:

tuna

100-101 see also burrows

1

5,

1

1,

34

36-39

north European water (water) 7: 9, 98-99

92-93 southern red-backed 7: 94-95 waterd: 54; 7: 9, 98-99 Vombatiformes 10: 75 Vombatus ursinus 10: 74, 98-101 vomeronasal organ 5: 62, 65 Vormela peregusna

1:

32

X 1:

10; 9:

64-66

Y

36 1:

32,

yak

6:

74-75

wild 6: 75 1:

34

32,

weasel family

webbed

1:

yapok see opossum, water

wetlands

whale

Yellowstone National Park

32-35

fingers 4: 71

Weil's disease 7: 14,

74

10,

54-59:

(1 1),

14;

5: (10),

1: (11); 3:

bowhead

20-23 zebra 2: 8-9, 6: (85), 95

66,

54, 55,

110-111 dwarf sperm 3: 55 gray 3: 55, 57, (57), 59, 92-97 3: 55, (57),

killer 3: 37, 39, 55,

62-65

.

83, 94,

96

12-13;

46-51

common

3: 55, 108,

102-105

5: 12,

3: 9,

Burchell's (plains) 5: 42,

98-101

humpback

76

Zalophus californianus

7: 12

(57), 58,

67

z

24

1:

1:

6: 66,

Yersinia pestis 7:

baleen 3: 54-55, 56, 58

blue

104, 105,

9: (46)

Xenarthra

32, 34, 35,

short-tailed see stoat

3:

5:

Wyulda squamicaudata 10: 74

34

short-tailed (field) 7:

7: 12, 8:

worms

(least)

North African banded Patagonian

V megaspila

87, 7: 34,

Conservation of Nature

long-tailed 1: 32,

30

1:

World Conservation Union see International Union for the

36-39 30

woodchuck

(106), 108, 111

96

5:

common

1: 19,

northern hairy-nosed 10: 74, (101) southern hairy-nosed 10: 74, 101, (101)

50-51 wool production

in

62,

32, 34, 35,

1: 19,

least

1

65

desert 5: 74 washing bears 1: (24) Washington Convention see Convention on

5: 75; 6:

18, 32, 35,

98-101

80-83

74

5:

European

:

27

75,

5:

African striped 1: 32,

28

2: 50, 53,

.

weasel

1

timber (gray)

wombat 10: (10), 77, 93 common 10: 74, 75,

defassa 6: 93 water reabsorption

5: 92,

Mexican 2: 54 red 2: 59 Tasmanian see thylacine

wapiti see elk warfarin 7: 75

waterbuck 92-93

04

53

54-57 59 wolverine 1: 56-57

Endangered Species of Wild Fauna and Flora

plains 8: 28, 30 viscacha rat 8: 29,

truffles 5:

50

variegata rubra 4: 104 variegata variegata 1

2: 50,

54-57.

marsupial see thylacine

International Trade

northern 8: 30

1

28-29, 31 forest 8: 30 strong 8: 30

62-63

86

2: 50, 53,

maned 60

10: 48, 50,

common

95

(53)

104-105

mountain

triok, Tate's 10: 74, (77)

tuco-tuco

9:

6:

walrus 3: 8, 9, 10, 11, 24-29 want see mole, European

warthog

Viverra

Trichys fasciculata 8:

9:

:

viscacha 7: 12, 8: 28

3:

Tubulidentata

6:

2 82

vicuna 5: 92, 93, 110-111

10: (80)

Tetracerus quadricornis

94-97 maritimus 2: 82, 84-89 melanoleuca 2: 98

U. arctos 2: 82, 92,

whiteness in mammals 8: (84) wildcat 2: 10, 13, 48-49 African 2: 72 European 2: 72

gray 59

yellow-footed rock 10: 48, 70-71 wallaroo 10: 60-61 10: 48,

3: 58, 89, 91,

93, 101, 105, 107, 108 whistlepig see woodchuck

Falkland Island 2: 50, 53,

wallowing 5: 38, 69, 71, 77, 76-77, 83

2: 82,

58 59

tuberculosis, bovine 1: 81;

6: 87, 9: 64, 66, 69;

40

90-93

1

I caninus 10: 74 I vulpecula 10: 74, 78-81

9: 24,

americanus

U.

56 Vicugna vicugna 10-1

Trichosurus

Tenrec 9: 9

59

vibrissae 8:

48-51 senegalensis 3: 47

manatus

9:

60-61 58, 58, 59

tribal

25

spiny 9:

50

49

whiptail 10: 48, 57

hill

4:

Tremarctos ornatus 2: 82 warfare 4: 30 Trichechus T. inunguis 3: 47

25 24

50

Vespertiliomdae 9: 87

9: 59,

9: 58,

2:

common

V.

58-59

pen-tailed 9:

pygmy

cinereoargenteus

Ursus

1/

10, 5: (10);

1:

Philippine 9:

9: 24,

25

24,

shrew

tree

rock 10: 51

10: 48,

whale meat 3: 59, 67 whale songs 3: 99-100, 105 whale watching 3: (57), 71, 95, 104

wildebeest 5: 12, 47; black 6: 62 blue 6: 62, 82-85 wolf 2: 9 Ethiopean 2: 50, 52

70-71

Urotrichus 9: 54

y

48 48-49

6: 10,

10: 48,

ringtailed rock (yellow-

swamp

Varecia

tree dwelling 2: 31

long-tailed 9: 24,

pygmy shrew

napu

common

26-27 greater hedgehog lesser

T.

red-necked

50 64-65

toolache 10: 53

2: (67)

Tragulus 6: 48 T. javanicus 6: 10,

Telicomys 7: 11-12

48

forest 10:

prettyfacc 10: 48, 57 Proserpine rock 10: 48,

European Urocyon

vaccination, against rabies

78-79

Papuan

66

55, 57, 58,

whaling industry

northern nail-tailed 10: 48

rufous hare 10: 48, 57 spectacled hare 10: 48

U.

50

48

footed rock) 10: 48,

Vampyrum spectrum

T strepsiceros 6: Tragulidae 6: 10

9

teledu

lesser forest 10:

3:

3: 55,

white see beluga

bridled nail-tailed 10: 48,

V

62

sperm 3: 86-89

48

0:

wallaby 10: 48-53 banded hare 10: 48, 57

8-13

U- thibetanus

scriptus 6:

1

urchin see hedgehog,

U.

T obscurus 4: 40 Tragelaphus T. buxtom 6: 62 T.

Wallabia bicolor

43

U.

Trachypithecus T geei 4: 40

88 89

southern right 108-109

w

even-toed 1: 10; 5: 10, 88 odd-toed 1: 70; 5: 10, 7 7,

Uropsilus investigator 9:

3: (57),

2: 50,

right 3: 55 short-finned pilot

64-67 74-75

Bennett's (red-necked) 10: 48, 64-65

Urogale everetti

1,

31-32, (31)

whale watching 71, 95, 104

Tayassu

zerda

U. littoralis 2:

topi 6: 62, (85), 88-89 torpor 7: 80; 9: 20-21, 29, 38, 82, 102; 10: 40, 85, 103 tourism 3: 51, 71, (94), 95, 5: 39, 9: 53, 101

76

1/

5:

pygmy right 3: 54, 55upygmy sperm 3: 55

50

87

titi

tool users 1: (74), 4: 10-1

Taurotragus T. derbianus 6: 62, 76-77 T. derbianus derbianus

70-73

2: 50,

vulpes 2: 50,

black-headed 4: 80 red 4: 72, 80-81 white (red) 4: 72, 80-81 Umfolozi park 5: 34 Uncia uncia 2: 34-35 ungulates (hoofed mammals)

Thomson's

ruppelli 2:

2:

V.

17,

bald (red) 4: 72, 80-81 black 4: 9, 72

dusky 4: 72 masked 4: 72 yellow-handed 4: 72 toddy 1: 95 toddy cat see civet, common palm tommie see gazelle,

lagopus

3: 55, 106-107 northern bottlenose 3: 55, 90-91 northern right 3: 109

minke

2:

V.

5: 14,

uakari

2: 23, (23)

V.

1/

chama

26, 66, 72, 74, 80-81, 86,

u

23

V-

cana

V.

3: 55,

66-67

velox 2: 50, 68-69 velox herbes 2: 69

Tympanoctomys barrerae 8: 30 typhus 7: 14, 74

14

long-finned pilot 50

2:

50 50 corsac 2: 50

V.

V.

see also ivory

tiger 2: 10, 20-25 Bali 2: 21 Bengal 2: 21, 27, 22, Caspian 2: 21 Chinese 2: 20, 21

28-31

6:

87

9:

106, 107

6:

tusks 3: 26, 84-85,

tiger-horses 5: 52

Tarsipes rostratus 10: 74,

9:

72-7 5

88

ticks 6: 18; 7:

white

107

spectral 4: 106,

minor

Tursiops truncatus 3: 55,

Siberian 2: 20, 21,

64-65

T.

east Caucasian 6: 106, 107

22-23 Sumatran 2: 21 Tasmanian see thylacine

62

5:

59,

west Caucasian

Indochinese 2: 21 Javan 2: 21

T.

pygmy

tiang 6:

glis 9:

48

Thyropteridae

,

62, 63,

5:

thetis 10:

60-61 59

T.

tur

10: 27, 36-37 Thylamis T. elegans 10: 14 T. pailidior 10: 14 Thylogale T. stigmatica 10: 48

48

T.

Vulpes V. bengalensis

Jupaia

2: 80; 10: 24, 26,

28, 36-37 Thylacinidae 10: 24, 26 Thylacinus cynocephalus

88-91 lion 4: 86

58,

(plains) 5: 42,

46-51 Grevy's 5: 42, (44), 52-53 mountain 5: 42 plains 5: 42, 46-51 zebra family 5: 42-45 zokor

7:

zorilla 1:

1

32, 34,

34

127


SMALL CARNIVORES

Picture Credits Abbreviations Frank Lane Picture Agency

FLPA

OSF

Oxford

Scientific Films

Ulrich/OSF;

= top; b = bottom;

c

=

center;

I

=

left;

r

=

T.

Leeson/Sunset/FLPA; 44-45 Mike

Birkhead/OSF; 46-47 Alan

Tom t

42-43

& Sandy

Carey/OSF; 48-49

50-51 Alan & Sandy Carey/OSF;

Ulrich/OSF;

52-53 Marianne Wilding/Survival Anglia/OSF; 54 Dean

right

Conger/Corbis; 55 Richard Packwood/OSF; 56-57, 57 Daniel

Jacket tl

caracal, Pete Oxford/naturepl.com; tr

gorilla,

bl

Cox/OSF; 58-59

Gordon/OSF; 61

group of

dolphins, Robert Harding Picture Library;

J.

lowland

Niall

Benvie/OSF; 60-61 Nick

Benvie/OSF; 62-63 Bridget

Niall

Wheeler/Survival Anglia/OSF;

64-65 Joe

McDonald/Animals Animals/OSF; 66 Alan & Sandy

Martin Rugner/Naturphotographie; br

Carey/OSF; 66-67 Daniel

Rothchild's giraffe, Gerard Lacz/FLPA

J.

Cox/OSF; 68-69 Nick

Gordon/OSF; 70-71 Michael Leach/OSF; 72-73 Stan 9 Stephen

J.

Krasemann/Bruce Coleman Collection;

Osolinski/OSF; 73

10-11 Peter Davey/FLPA; 12-13 K.G.

Preston-Mafham/Premaphotos

Howard

Hall/OSF;

74

Jeff

Foott/Okapia/OSF; 75 Claude Steelman/Survival

15

Wildlife;

Anglia/OSF; 76-77 Bob Bennett/OSF; 78-79 Neil

Rafi

Ben-Shahar/OSF; 18-19 Steve Turner/OSF; 21 Tom

Latham/OSF; 79 David Fox/OSF; 80 Richard

Vezo/naturepl.com; 22-23 Daniel

Packwood/OSF; 80-81 Robin Redfern/OSF; 82-83

Bettmann/Corbis; 24, 24-25

Tom

J.

Cox/OSF; 23

Ulrich/OSF; 26t Ralph

Anthony Bannister/ABPL/OSF; 84-85 Richard Day/OSF;

Reinhold/Animals Animals/OSF; 26b Richard Day/OSF;

86 TC Nature/OSF; 86-87

Zig Leszczynski/Animals

27 Wendy Shattil/Bob Rozinski/OSF; 28-29 Konrad

Animals/OSF; 87 Daniel

Cox/OSF; 89

Wothe/OSF; 30-31 Stan Osolinski/OSF; 33 Mark

Shah/naturepl.com; 92-93 Gerard Lacz/FLPA; 94-95

Hamblin/OSF; 35 Mark Deeble & Victoria Stone/OSF;

Werner Pfunder/OSF; 96-97, 97 Doug Allan/OSF;

36-37 Jorge

100-101, 102, 102-103, 103, 104-105 David

Sierra/OSF;

38 Press-Tige Pictures/OSF;

Anup

Macdonald/OSF; 106-107 Adrian Bailey/OSF; 108-109

38-39 David Thompson/OSF; 40-41 Mark Hamblin/OSF; 41 Robin Redfern/OSF; 42

J.

Tom

Belinda Wright/OSF;

110-111 Steve Turner/OSF

Artists Denys Ovenden,

While every effort has been

made

Priscilla

Barrett with Michael Long,

to trace the copyright holders rectify

128

of

illustrations

any omissions or

Graham

Allen,

reproduced

inaccuracies.

in this

Malcolm McGregor

book, the publishers

will

be pleased

to



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SMALL CARNIVORES Raccoons, Weasels, Otters, Skunks

LARGE CARNIVORES Big Cats, Dogs, Bears,

SEA Seals,

Hyenas

...

MAMMALS

Sea Lions, Whales, Dolphins, Manatees

PRIMATES Apes, Monkeys, Marmosets, Lemurs

LARGE HERBIVORES Elephants, Rhinos, Horses, Pies

RUMINANT (HORNED) HERBIVORES Deer, Cattle, Antelope, Goats,

RODENTS Squirrels, Rats,

RODENTS

2

Sheep

1

Mice

AND LAGOMORPHS

Porcupines, Cavies, Rabbits

...

INSECTIVORES AND BATS Hedgehogs, Moles, Anteaters, Bats

...

MARSUPIALS Kangaroos, Possums, Koala

I

90 Sherman Rjrnpike Danbury, CT 06816

SET ISBN 0 7172 5742 8

VOLUME

ISBN

0 7172 574.3 6


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