�@�@ •ª—Þƒgƒbƒv
Family Cyprinidae�@ƒRƒC‰È

---

Sub-Family Rohteinae FOWLER 1958�@ƒ��[ƒeƒGˆŸ‰È



Genus Amatolacypris^♀ SKELTON, SWARTZ & VREVEN 2018�@ƒAƒ}ƒgƒ‰ƒLƒvƒŠƒX‘®

�Ÿ
�@Amatolacypris trevelyani (Günther 1877)
�@�@�@syn.Barbus brookingi Gilchrist & Thompson 1913
�@�@�@�@�@Barbus trevelyani Günther 1877
�@�@�@�@�@Pseudobarbus trevelyani (Günther 1877)
�@^*2018�wThe Identity of Barbus capensis Smith 1841 and the Generic Status of Southern African Tetraploid Cyprinids [Teleostei, Cyprinidae]�FPaul H.Skelton, Ernst R.Swartz & Emmanuel J.Vreven�x�B
�@"ƒ{�[ƒ_�[ƒoƒ‹ƒu"�BIUCN Endangered�@ B1ab(ii,iii)+2ab(ii,iii)�@ ver.3.1�B

---

Genus Barbodes^♂ (BLEEKER 1859)�@ƒoƒ‹ƒ{ƒfƒX‘®
�@�@�@�¦�{Genus Cephalakompsus^♂ HERRE 1924 (Type species : Cephalakompsus pachycheilus Herre 1924)�B
�@�@�@�¦�{Genus Mandibularca^♀ HERRE 1924 (Type species : Mandibularca resina Herre 1924)�B
�@�@�@�¦�{Genus Ospatulus^♂ HERRE 1924 (Type species : Ospatulus truncatulus Herre 1924)�B
�@�@�@�¦�{Genus Spratellicypris^♀ HERRE & MYERS 1931 (Type species : Barbodes palatus Herre 1924)�B
�@�@�@�¦��Genus Systomus^♂ MCCLELLAND 1839 Sub-Genus Barbodes^♂ BLEEKER 1859�B
�@‚Q‚R�`‚Q‚T�Ž�E‚�‚g‚U�D‚T�`‚V�D‚O�B

�Ÿ
�@Barbodes amarus Herre 1924
�@�@�@syn.Puntius amarus (Herre 1924)
�@IUCN Critically Endangered�@ A1ce�@ ver.2.3�B

�Ÿ
�@Barbodes aurotaeniatus (Tirant 1885)
�@�@�@syn.Barbus aurotaeniatus Tirant 1885
�@�@�@�@�@Barbus pessuliferus (Fowler 1937)
�@�@�@�@�@Barbus stigmatosomus (Smith 1931)
�@�@�@�@�@Puntius pessuliferus Fowler 1937
�@�@�@�@�@Puntius samatensis Smith 1945
�@�@�@�@�@Puntius stigmatosomus Smith 1931
�@�@�@�@�@Systomus aurotaeniatus (Tirant 1885)
�@�@�@�@�@Puntius aurotaeniatus (Tirant 1885)
�@^*2021�wFishes of the Indo-Chinese Mekong�FNagao Natural Environment Foundation�x�B

�Ÿ
�@Barbodes banksi (Herre 1940)
�@�@�@syn.Puntius banksi (Herre 1940)
�@�@�@�@�@Puntius binotatus banksi Herre 1940
�@�@�@�@�@Systomus banksi (Herre 1940)
�@^*2021�wTo Lump, to Split or to Maintain ?, Molecular Taxonomy of the Spotted Barb Barbodes binotatus [Cyprinidae] and Closely Related Species in Peninsular Malaysia�FNoor Zahidah Ahmad Sobri, Sebastien Lavoue, Fazimah Aziz, Siti Azizah Mohd Nor, Noor Adelyna Mohammed Akib & Muhammad Zain Khaironizam�x�B


�Ÿ
�@Barbodes bantolanensis (Day 1914)
�@�@�@syn.Barbus bantolanensis Day 1914
�@�@�@�@�@Puntius bantolanensis (Day 1914)

�Ÿ
�@Barbodes baoulan Herre 1926
�@�@�@syn.Puntius baoulan (Herre 1926)
�@IUCN Critically Endangered�@ A1ce�@ ver.2.3�B

�Ÿ
�@Barbodes belinkus
�@�@�@syn.Puntius belinkus
�@�@�@�@�@Systomus belinkus

�Ÿ
�@Barbodes bunau (Rachmatika 2005)
�@�@�@syn.Puntius bunau Rachmatika 2005

�Ÿ
�@Barbodes binotatus (Valenciennes 1842)
�@�@�@syn.Barbus binotatus Valenciennes 1842
�@�@�@�@�@Barbus blitonensis Bleeker 1852
�@�@�@�@�@Barbus goniosomus (Bleeker 1860) Day 1878
�@�@�@�@�@Barbus kusanensis Bleeker 1852
�@�@�@�@�@Barbus maculatus hagenii Popta 1911
�@�@�@�@�@Barbus oresigenes Bleeker 1849
�@�@�@�@�@Barbus polyspilos Bleeker 1857
�@�@�@�@�@Capoeta binotata (Valenciennes 1842)
�@�@�@�@�@Puntius binotatus (Valenciennes 1842)
�@�@�@�@�@Puntius sibukensis Fowler 1941
�@�@�@�@�@Systomus binotatus (Valenciennes 1842)
�@�@�@�@�@Systomus goniosomus Bleeker 1860
�@^*1857�wDescriptiones Specierum Piscium Javanensium Novarum Vel Minus Cognitarum Diagnosticae�FPieter Bleeker�x�B
�@^*1878�wA Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters India, Burma and Ceylon 2, Atlas Constaining 198 Plates�FFrancis Day�x�B
�@^*1878�wThe Fishes of India, Being a Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters of India, Burma and Ceylon 1 Extra�FFrancis Day�x�B
�@^*2021�wTo Lump, to Split or to Maintain ?, Molecular Taxonomy of the Spotted Barb Barbodes binotatus [Cyprinidae] and Closely Related Species in Peninsular Malaysia�FNoor Zahidah Ahmad Sobri, Sebastien Lavoue, Fazimah Aziz, Siti Azizah Mohd Nor, Noor Adelyna Mohammed Akib & Muhammad Zain Khaironizam�x�B
�@"ƒXƒ|ƒbƒgƒoƒ‹ƒu"�B‚Q‚S�`‚Q‚U�Ž�E‚�‚g‚U�D‚O�`‚U�D‚T�BIUCN Least Concern ver.3.1�B

Barbus goniosomus=Puntius binotatus

�Ÿ
�@Barbodes cataractae (Fowler 1934)
�@�@�@syn.Barbus cataractae Fowler 1934
�@�@�@�@�@Puntius cataractae (Fowler 1934)
�@‹D�…ˆæ�i“ü�«�BIUCN Data Deficient ver.2.3�B

�Ÿ
�@Barbodes clemensi Herre 1924
�@�@�@syn.Puntius clemensi (Herre 1924)

�Ÿ
�@Barbodes colemani (Fowler 1937)
�@�@�@syn.Barbus colemani Fowler 1937
�@�@�@�@�@Protopuntius colemani (Fowler 1937)
�@�@�@�@�@Puntius colemani (Fowler 1937)

�Ÿ
�@Barbodes disus Herre 1932
�@�@�@syn.Puntius disus (Herre 1932)
�@IUCN Critically Endangered�@ A1ce�@ ver.2.3�B

�Ÿ
�@Barbodes dorsimaculatus (Ahl 1923)
�@�@�@syn.Barbus dorsimaculatus Ahl 1923
�@�@�@�@�@Puntius dorsimaculatus (Ahl 1923)
�@"ƒuƒ‰ƒbƒNƒ‰ƒCƒ“ƒoƒ‹ƒu"�B

�Ÿ
�@Barbodes dunckeri (Ahl 1929)
�@�@�@syn.Barbus dunckeri Ahl 1929
�@�@�@�@�@Puntius dunckeri
�@"ƒrƒbƒOƒXƒ|ƒbƒgƒoƒ‹ƒu"�B


�Ÿ
�@Barbodes everetti (Boulenger 1894)
�@�@�@syn.Barbus everetti Boulenger 1894
�@�@�@�@�@Puntius everetti (Boulenger 1894)
�@"ƒNƒ‰ƒEƒ“ƒoƒ‹ƒu"�B‚Q‚S�`‚R‚O�Ž�E‚�‚g‚U�D‚O�`‚U�D‚T�B


�Ÿ
�@Barbodes flavifuscus Herre 1924
�@�@�@syn.Puntius flavifuscus (Herre 1924)
�@IUCN Critically Endangered�@ A1ce�@ ver.2.3�B


�Ÿ
�@Barbodes hemictenus Jordan & Richardson 1908
�@�@�@syn.Puntius hemictenus (Jordan & Richardson 1908)
�@^*1908�wFishes from Islands of the Philippine Archipelago 1907�FDavid Starr Jordan & Robert Earl Richardson�x�B
�@IUCN Vulnerable�@ D2�@ ver.2.3�B

�Ÿ
�@Barbodes herrei (Fowler 1934)
�@�@�@syn.Barbus herrei Fowler 1934
�@�@�@�@�@Puntius herrei (Fowler 1934)
�@IUCN Critically Endangered�@ A2ce�@ ver.3.1�B


�Ÿ
�@Barbodes ivis (Seale 1910)
�@�@�@syn.Barbus ivis Seale 1910
�@�@�@�@�@Puntius ivis (Seale 1910)
�@^*1910�wNew Species of Philippine Fishes 1909�FAlvin Seale�x�B

�Ÿ
�@Barbodes joaquinae (Wood 1968)
�@�@�@syn.Puntius joaquinae Wood 1968

�Ÿ
�@Barbodes katolo Herre 1924
�@�@�@syn.Puntius katolo (Herre 1924)
�@IUCN Critically Endangered�@ A1ce�@ ver.2.3�B

�Ÿ
�@Barbodes kuchingensis (Herre 1940)
�@�@�@syn.Puntius kuchingensis Herre 1940

�Ÿ
�@Barbodes lanaoensis Herre 1924
�@�@�@syn.Puntius lanaoensis (Herre 1924)
�@�@�@�¦IUCN Critically Endangered�@ A1ce�@ ver.2.3�B


�Ÿ
�@Barbodes lateristrigus (Valenciennes 1842)
�@�@�@syn.Barbus lateristrigus Valenciennes 1842
�@�@�@�@�@Puntius lateristrigus (Valenciennes 1842)
�@�@�@�@�@Systomus lateristrigus (Valenciennes 1842)
�@^*1904�wFische von Smatra�FWalter Volz�x�B
�@"ƒXƒpƒ“ƒi�[ƒoƒ‹ƒu"�E"�§(ƒ|ƒXƒg)ƒtƒBƒbƒVƒ…"�B‚Q‚T�`‚Q‚W�Ž�E‚�‚g‚U�D‚O�`‚U�D‚T�B


�Ÿ
�@Barbodes lindog Herre 1924
�@�@�@syn.Puntius lindog (Herre 1924)
�@IUCN Vulnerable�@ A2cde�@ ver.2.3�B

�Ÿ
�@Barbodes maculatus (Valenciennes 1842)
�@�@�@syn.Systomus maculatus Valenciennes 1842�@�¦��Systomus (Barbodes) maculatus Valenciennes 1842 non McClelland 1838 non Kuhl & van Hasselt 1823�B
�@�¨Barbodes binotatus (Valenciennes 1842)�H

�Ÿ
�@Barbodes mahakkamensis
�@�@�@syn.Barbus mahakkamensis

�Ÿ
�@Barbodes manalak Herre 1924
�@�@�@syn.Puntius manalak (Herre 1924)
�@IUCN Critically Endangered�@ A1ce�@ ver.2.3�B

�Ÿ
�@Barbodes manguaoensis (Day 1914)
�@�@�@syn.Barbus manguaoensis Day 1914
�@�@�@�@�@Puntius manguaoensis (Day 1914)
�@IUCN Vulnerable�@ D2�@ ver.2.3�B

�Ÿ
�@Barbodes microps (Günther 1868)
�@�@�@syn.Barbus microps Günther 1868
�@�@�@�@�@Puntius microps (Günther 1868)

�Ÿ
�@Barbodes montanoi (Sauvage 1881)
�@�@�@syn.Puntius montanoi Sauvage 1881
�@^*2021�wBarbodes pyrpholeos, New Species, the First Cave-Dwelling Cyprinid Fish in the Philippines, with Re-Description of B.montanoi [Teleostei, Cyprinidae]�FTan Heok Hui & Daniel Edison M.Husana�x�B

�Ÿ
�@Barbodes pachycheilus (Herre 1924)
�@�@�@syn.Cephalakompsus pachycheilus Herre 1924
�@�@�@�@�@Puntius pachycheilus (Herre 1924)
�@IUCN Critically Endangered�@ A1ce�@ ver.2.3�B

�Ÿ
�@Barbodes palaemophagus (Herre 1924)
�@�@�@syn.Ospatulus palaemophagus Herre 1924

�Ÿ
�@Barbodes palavanensis (Boulenger 1895)
�@�@�@syn.Barbus palavanensis Boulenger 1895

�Ÿ
�@Barbodes palatus Herre 1924
�@�@�@syn.Puntius palatus (Herre 1924)
�@�@�@�@�@Spratellicypris palata (Herre 1924)
�@IUCN Critically Endangered�@ A1ce�@ ver.2.3�B

�Ÿ
�@Barbodes platysomus
�@�@�@syn.Barbus platysomus
�@�@�@�@�@Puntius platysomus
�@�@�@�@�@Systomus platysomus

�Ÿ
�@Barbodes polylepis

�Ÿ
�@Barbodes pyrpholeos Hui & Husana 2021
�@^*2021�wBarbodes pyrpholeos, New Species, the First Cave-Dwelling Cyprinid Fish in the Philippines, with Re-Description of B.montanoi [Teleostei, Cyprinidae]�FTan Heok Hui & Daniel Edison M.Husana�x�B


�Ÿ
�@Barbodes quinquemaculatus (Seale & Bean 1908)
�@�@�@syn.Barbus quinquemaculatus Seale & Bean 1908
�@�@�@�@�@Puntius quinquemaculatus (Seale & Bean 1908)
�@^*1908�wOn a Collection of Fishes from the Philippine Islands, made by Maj. Edgar A.Mearns, Surgeon, US Army, with Descriptions of Seven New Species�FAlvin Seale & Barton Appler Bean�x�B

�Ÿ
�@Barbodes resinus (Herre 1924)
�@�@�@syn.Mandibularca resina Herre 1924
�@�@�@�@�@Puntius resinus (Herre 1924)
�@IUCN Critically Endangered�@ A1ce�@ ver.2.3�B

�Ÿ
�@Barbodes rhombeus (Kottelat 2000)
�@�@�@syn.Puntius rhombeus Kottelat 2000
�@^*2021�wFishes of the Indo-Chinese Mekong�FNagao Natural Environment Foundation�x�B

�Ÿ
�@Barbodes sellifer Kottelat & Lim 2021
�@^*2021�wTwo New Species of Barbodes from the Malay Peninsula and Comments on Cryptic Species in the B.binotatus Group [Teleostei, Cyprinidae]�FMaurice Kottelat & Kelvin Kok Peng Lim�x�B

�Ÿ
�@Barbodes semifasciolatus (Günther 1868)
�@�@�@syn.Barbus hainani Lohberger 1929
�@�@�@�@�@Barbus semifasciolatus Günther 1868
�@�@�@�@�@Capoeta semifasciolata (Günther 1868)
�@�@�@�@�@Puntius semifasciolatus (Günther 1868)
�@^*2020�wŒÎ“ì‹›—ÞŒ´�F�}•ˆ�Fœ@•š�‰�^—›�ƒ�^—k鑫�^åÍŠó•½�^�Ó�‘–¯�^—ÀŽu‹­�^ŒÞ‰“ˆÀ�x�B
�@^*2021�wFishes of the Indo-Chinese Mekong�FNagao Natural Environment Foundation�x�B
�@"ƒ`ƒƒƒCƒj�[ƒYƒoƒ‹ƒu"�B‚P‚W�`‚Q‚S�Ž�E‚�‚g‚U�D‚O�`‚W�D‚O�B


�Ÿ
�@Barbodes sirang Herre 1932
�@�@�@syn.Puntius sirang (Herre 1932)
�@IUCN Vulnerable�@ A2cde�@ ver.2.3�B

�Ÿ
�@Barbodes snyderi (Oshima 1919)
�@�@�@syn.Puntius snyderi Oshima 1919

�Ÿ
�@Barbodes strigatus
�@�@�@syn.Barbus strigatus
�@�@�@�@�@Puntius strigatus

�Ÿ
�@Barbodes sunieri
�@�@�@syn.Puntius sunieri

�Ÿ
�@Barbodes tras Herre 1926
�@�@�@syn.Puntius tras (Herre 1926)
�@IUCN Critically Endangered�@ A1ce�@ ver.2.3�B

�Ÿ
�@Barbodes truncatulus (Herre 1924)
�@�@�@syn.Ospatulus truncatulus Herre 1924
�@IUCN Critically Endangered�@ A1ce�@ ver.2.3�B

�Ÿ
�@Barbodes tumbus Herre 1924
�@�@�@syn.Puntius tumbus (Herre 1924)
�@IUCN Vulnerable�@ A2cde�@ ver.2.3�B

�Ÿ
�@Barbodes umalii (Wood 1968)
�@�@�@syn.Puntius umalii Wood 1968


�Ÿ
�@Barbodes wynaadensis (Day 1873)
�@�@�@syn.Barbus wynaadensis Day 1873
�@�@�@�@�@Neolissochilus wynaadensis (Day 1873)
�@�@�@�@�@Puntius wynaadensis (Day 1873)
�@^*1878�wA Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters India, Burma and Ceylon 2, Atlas Constaining 198 Plates�FFrancis Day�x�B
�@^*1878�wThe Fishes of India, Being a Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters of India, Burma and Ceylon 1 Extra�FFrancis Day�x�B

�Ÿ
�@Barbodes xouthos (Kottelat & Hui 2011)
�@�@�@syn.Puntius xouthos (Kottelat & Hui 2011)
�@�@�@�@�@Systomus xouthos Kottelat & Hui 2011
�@^*2011�wSystomus xouthos, a New Cyprinid Fish from Borneo and Revalidation of Puntius pulcher [Teleostei, Cyprinidae]�FMaurice Kottelat & Tan-Heok Hui�x�B

�Ÿ
�@Barbodes zakariaismaili Kottelat & Lim 2021
�@^*2021�wTwo New Species of Barbodes from the Malay Peninsula and Comments on Cryptic Species in the B.binotatus Group [Teleostei, Cyprinidae]�FMaurice Kottelat & Kelvin Kok Peng Lim�x�B

---

Genus Barboides^♂ BRÜNING 1929�@ƒoƒ‹ƒ{ƒCƒfƒX‘®
�@�@�@�¦�{Genus Raddabarbus^♂ VAN DEN AUDENAERDE 1971 (Type species : Raddabarbus camerunensis van den Audenaerde 1971)�B
�@�@‚Q‚S�`‚Q‚U�Ž�E‚�‚g‚U�D‚O�`‚V�D‚O�B

�Ÿ
�@Barboides britzi

�Ÿ
�@Barboides gracilis Brüning 1929
�@�@�@syn.Barbus lorenzi Loiselle & Welcomme 1971
�@�@�@�@�@Raddabarbus camerunensis van den Audenaerde 1971
�@IUCN Vulnerable�@ B2ab(iii)�@ ver.3.1�B

---

Genus Bhava^♀ SUDASINGHE, RÜBER & MEEGASKUMBURA 2023�@ƒo�[ƒEƒ@‘®


�Ÿ
�@Bhava vittata (Day 1865) Sudasinghe, Rüber & Meegaskumbura 2023
�@�@�@syn.Barbus vittatus (Day 1865) Day 1878
�@�@�@�@�@Puntius vittatus Day 1865
�@^*1878�wA Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters India, Burma and Ceylon 2, Atlas Constaining 198 Plates�FFrancis Day�x�B
�@^*1878�wThe Fishes of India, Being a Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters of India, Burma and Ceylon 1 Extra�FFrancis Day�x�B
�@^*1928�wFauna of the Chilka Lake 1915-1924 4, Fish 1 1916�FB.L.Chaudhuri�x�B
�@^*2023�wMolecular Phylogeny and Systematics of the South Asian Fresh-Water Fish Genus Puntius [Teleostei, Cyprinidae]�FHiranya Sudasinghe, Lukas Rüber & Madhava Meegaskumbura�x�B
�@"ƒOƒŠ�[ƒ“ƒXƒgƒ‰ƒCƒvƒoƒ‹ƒu"�B‹D�…ˆæ�i“ü�«�B‚Q‚O�`‚Q‚S�Ž�E‚�‚g‚U�D‚O�`‚U�D‚T�B

---

Genus Chagunius^♂ SMITH 1938�@ƒJƒOƒjƒEƒX‘®
�@‚Q‚Q�`‚Q‚V�Ž�E‚�‚g‚W�D‚O�B•ª—ÞŠw“IˆÊ’u‚Í•sˆÀ’è�B

�Ÿ
�@Chagunius baileyi


�Ÿ
�@Chagunius chagunio (Hamilton 1822)
�@�@�@syn.Barbus beavani Günther 1868
�@�@�@�@�@Barbus chagunio (Hamilton 1822) Day 1878
�@�@�@�@�@Barbus spilopholus McClelland 1839
�@�@�@�@�@Cyprinus chagunio Hamilton 1822
�@�@�@�@�@Puntius chagunio (Hamilton 1822)
�@�@�@�@�@Rohita chagunio (Hamilton 1822)
�@^*1878�wA Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters India, Burma and Ceylon 2, Atlas Constaining 198 Plates�FFrancis Day�x�B
�@^*1878�wThe Fishes of India, Being a Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters of India, Burma and Ceylon 1 Extra�FFrancis Day�x�B
�@"ƒ`ƒƒƒOƒj"�B

�Ÿ
�@Chagunius nicholsi (Myers 1924)
�@�@�@syn.Barbus nicholsi Myers 1924 non Vinciguerra 1928
�@�@�@�@�@Chagunius chagunio prashadi Talwar & Das 1986

---

Genus Cheilobarbus^♂ (SMITH 1841)�@ƒPƒCƒ�ƒoƒ‹ƒuƒX‘®
�@�@�@�¦��Genus Barbus^♂ CUVIER & CLOQUET 1816 Sub-Genus Cheilobarbus^♂ SMITH 1841�B

�Ÿ
�@Cheilobarbus capensis (Smith 1841)
�@�@�@syn.Barbus andrewi Barnard 1937
�@�@�@�@�@Barbus capensis Smith 1841�@�¦��Barbus (Cheilobarbus) capensis Smith 1841�B
�@�@�@�@�@Labeobarbus capensis (Smith 1841)
�@�@�@�@�@Pseudobarbus capensis (Smith 1841)�@�¦ƒtƒBƒbƒVƒ…ƒx�[ƒX‚Ì—LŒø–¼�Ì�B
�@"ƒNƒ‰ƒ“ƒEƒBƒŠƒAƒ€ƒCƒGƒ��[ƒtƒBƒbƒVƒ…"�E"ƒP�[ƒvƒzƒ�ƒCƒgƒtƒBƒbƒVƒ…"�BIUCN Vulnerable�@ B2ab(iii,v)�@ ver.3.1�B

---

Genus Clypeobarbus^♂ (FOWLER 1936)�@ƒNƒŠƒyƒIƒoƒ‹ƒuƒX‘®
�@�@�@�¦��Genus Barbus^♂ CUVIER & CLOQUET 1816 Sub-Genus Clypeobarbus^♂ FOWLER 1936�B
�@‚Q‚S�`‚Q‚V�Ž�B

�Ÿ
�@Clypeobarbus bellcrossi
�@�@�@syn.Barbus bellcrossi
�@"ƒS�[ƒWƒƒƒXƒoƒ‹ƒu"�BIUCN Data Deficient ver.3.1�B

�Ÿ
�@Clypeobarbus bomokandi
�@�@�@syn.Barbus bomokandi
�@�@�@�@�@? Barbus rubripinnis
�@IUCN Data Deficient ver.3.1�B

�Ÿ
�@Clypeobarbus breviclipeus Stiassny & Sakharova 2016

�Ÿ
�@Clypeobarbus congicus
�@�@�@syn.Barbus congicus
�@"ƒRƒ“ƒSƒoƒ‹ƒu"�BIUCN Least Concern ver.3.1�B


�Ÿ
�@Clypeobarbus hypsolepis
�@�@�@syn.Barbus hypsolepis
�@IUCN Least Concern ver.3.1�B

�Ÿ
�@Clypeobarbus matthesi (Poll & Gosse 1963)
�@�@�@syn.Barbus matthesi Poll & Gosse 1963
�@�@�@�@�@Enteromius matthesi (Poll & Gosse 1963)
�@IUCN Least Concern ver.3.1�B

�Ÿ
�@Clypeobarbus pleuropholis (Boulenger 1899)
�@�@�@syn.Barbus gribinguensis Pellégrin 1919
�@�@�@�@�@Barbus kemoensis Fowler 1936�@�¦��Barbus (Clypeobarbus) kemoensis Fowler 1936�B
�@�@�@�@�@�@�@�¦–ÍŽ®Ží�A’A‚µ�æ‚É‹L�Ú‚³‚ꂽ Barbus pleuropholis Boulenger 1899 ‚É“��‡�B
�@�@�@�@�@Barbus pleuropholis Boulenger 1899
�@�@�@�@�@Barbus uellensis Boulenger 1919
�@IUCN Least Concern ver.3.1�B

�Ÿ
�@Clypeobarbus pseudognathodon
�@�@�@syn.Barbus pseudognathodon
�@IUCN Near Threatened ver.3.1�B

�Ÿ
�@Clypeobarbus schoutedeni
�@�@�@syn.Barbus schoutedeni
�@IUCN Data Deficient ver.3.1�B

---

Genus Dawkinsia^♀ PETHIYAGODA, MEEGASKUMBURA & MADUWAGE 2012�@ƒ_ƒEƒLƒ“ƒVƒA‘®
�@ƒvƒ“ƒeƒBƒEƒX‘®‚æ‚蕪—£“Æ—§�B

�Ÿ
�@Dawkinsia apsara Katwate, Knight, Anoop, Raghavan & Dahanukar 2020
�@^*2020�wThree New Species of Filament Barbs of the Genus Dawkinsia [Teleostei, Cyprinidae] from the Western Ghats of India�FUnmesh Katwate, John Daniel Marcus Knight, V.K.Anoop, Rajeev Raghavan & Neelesh Dahanukar�x�B


�Ÿ
�@Dawkinsia arulia (Jerdon 1849)
�@�@�@syn.Barbus arulius (Jerdon 1849) Day 1878
�@�@�@�@�@Puntius arulius (Jerdon 1849)
�@�@�@�@�@Systomus arulius Jerdon 1849
�@^*1878�wA Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters India, Burma and Ceylon 2, Atlas Constaining 198 Plates�FFrancis Day�x�B
�@^*1878�wThe Fishes of India, Being a Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters of India, Burma and Ceylon 1 Extra�FFrancis Day�x�B
�@"ƒI�[ƒŠƒEƒXƒoƒ‹ƒu"�B‚P‚X�`‚Q‚T�Ž�E‚�‚g‚U�D‚O�`‚U�D‚T�B

�Ÿ
�@Dawkinsia assimilis (Jerdon 1849)
�@�@�@syn.Puntius assimilis (Jerdon 1849)
�@�@�@�@�@Puntius lepidus Day 1868
�@�@�@�@�@Systomus assimilis Jerdon 1849


�Ÿ
�@Dawkinsia austella Katwate, Knight, Anoop, Raghavan & Dahanukar 2020
�@^*2020�wThree New Species of Filament Barbs of the Genus Dawkinsia [Teleostei, Cyprinidae] from the Western Ghats of India�FUnmesh Katwate, John Daniel Marcus Knight, V.K.Anoop, Rajeev Raghavan & Neelesh Dahanukar�x�B

�Ÿ
�@Dawkinsia crassa Katwate, Knight, Anoop, Raghavan & Dahanukar 2020
�@^*2020�wThree New Species of Filament Barbs of the Genus Dawkinsia [Teleostei, Cyprinidae] from the Western Ghats of India�FUnmesh Katwate, John Daniel Marcus Knight, V.K.Anoop, Rajeev Raghavan & Neelesh Dahanukar�x�B

�Ÿ
�@Dawkinsia exclamatio (Pethiyagoda & Kottelat 2005)
�@�@�@syn.Puntius exclamatio Pethiyagoda & Kottelat 2005


�Ÿ
�@Dawkinsia filamentosa (Valenciennes 1844) Pethiyagoda, Meegaskumbura & Maduwage 2012
�@�@�@syn.Barbus filamentosus (Valenciennes 1844) Day 1878
�@�@�@�@�@Leuciscus filamentosus Valenciennes 1844
�@�@�@�@�@Puntius filamentosus (Valenciennes 1844)
�@�@�@�@�@Systomus filamentosus (Valenciennes 1844)
�@�@�@�@�@Systomus maderaspatensis Jerdon 1849
�@�@�@�@�@Dawkinsia singhala (Duncker 1912)
�@�@�@�@�@Barbus singhalus Duncker 1912
�@�@�@�@�@Puntius singhalus (Duncker 1912)
�@^*1878�wThe Fishes of India, Being a Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters of India, Burma and Ceylon 1 Extra�FFrancis Day�x�B
�@^*2012�wA Synopsis of the South Asian Fishes Referred to Puntius [Pisces, Cyprinidae]�FRohan Pethiyagoda, Madhava Meegaskumbura & Kalana Maduwage�x�B
�@^*2012�wDawkinsia, New Genus�FRohan Pethiyagoda, Madhava Meegaskumbura & Kalana Maduwage�x�B

�@"ƒuƒ‰ƒbƒNƒXƒ|ƒbƒgƒoƒ‹ƒu"�B‹D�…ˆæ�i“ü�«�B‚Q‚O�`‚Q‚S�Ž�E‚�‚g‚U�D‚O�`‚U�D‚T�B

�Ÿ
�@Dawkinsia rohani (Rema-Devi, Indra & Knight 2010)
�@�@�@syn.Puntius rohani Rema-Devi, Indra & Knight 2010


�Ÿ
�@Dawkinsia rubrotincta (Jerdon 1849)
�@�@�@syn.Puntius rubrotinctus (Jerdon 1849)
�@�@�@�@�@Systomus rubrotinctus Jerdon 1849

�Ÿ
�@Dawkinsia srilankensis
�@�@�@syn.Barbus srilankensis
�@�@�@�@�@Puntius srilankensis
�@"ƒXƒŠƒ‰ƒ“ƒJƒoƒ‹ƒu"�E"ƒuƒ�ƒbƒNƒtƒBƒ‰ƒ�ƒ“ƒgƒoƒ‹ƒu"�B

�Ÿ
�@Dawkinsia tambraparniei (Silas 1954)
�@�@�@syn.Puntius arulius tambraparniei Silas 1954
�@�@�@�@�@Puntius tambraparniei (Silas 1954)


�Ÿ
�@Dawkinsia uttara Katwate, Apte & Raghavan 2020
�@^*2020�wDawkinsia uttara, a New Species of Filament Barb [Teleostei, Cyprinidae] from the Western Ghats of India�FUnmesh Katwate, Deepak Apte & Rajeev Raghavan�x�B

---

Genus Desmopuntius^♂ KOTTELAT 2013�@ƒfƒXƒ‚ƒvƒ“ƒeƒBƒEƒX‘®

�Ÿ
�@Desmopuntius foerschi (Kottelat 1982)
�@�@�@syn.Barbus foerschi Kottelat 1982
�@�@�@�@�@Puntius foerschi (Kottelat 1982)
�@�@�@�@�@Systomus foerschi (Kottelat 1982)
�@‚Q‚S�`‚Q‚W�Ž�E‚�‚g‚U�D‚O�`‚U�D‚T�B


�Ÿ
�@Desmopuntius gemellus (Kottelat 1996)
�@�@�@syn.Puntius gemellus Kottelat 1996


�Ÿ
�@Desmopuntius hexazonus (Weber & de Beaufort 1912)
�@�@�@syn.Barbus hexazonus Weber & de Beaufort 1912
�@�@�@�@�@Puntius hexazonus (Weber & de Beaufort 1912)
�@�@�@�@�@Systomus hexazonus (Weber & de Beaufort 1912)
�@^*1931�wThe Fish-Fauna of the Rokan Mouth�FJohann Dietrich Franz Hardenberg�x�B


�Ÿ
�@Desmopuntius johorensis (Duncker 1904)
�@�@�@syn.Barbus eugrammus (Silas 1956)
�@�@�@�@�@? Barbus fasciatus Bleeker 1853 non Jerdon 1849
�@�@�@�@�@Barbus tetrazonus johorensis Duncker 1904
�@�@�@�@�@Puntius eugrammus Silas 1956
�@�@�@�@�@? Puntius fasciatus (Bleeker 1853)
�@�@�@�@�@Puntius johorensis Duncker 1904
�@�@�@�@�@Systomus johorensis (Duncker 1904)
�@"ƒXƒgƒ‰ƒCƒvƒoƒ‹ƒu"�B‚Q‚R�`‚Q‚T�Ž�E‚�‚g‚U�D‚O�`‚U�D‚T�B


�Ÿ
�@Desmopuntius pentazonus (Boulenger 1894)
�@�@�@syn.Barbus pentazonus Boulenger 1894
�@�@�@�@�@Capoeta pentazona (Boulenger 1894)
�@�@�@�@�@Puntius pentazonus (Boulenger 1894)
�@�@�@�@�@Systomus pentazonus (Boulenger 1894)
�@"ƒtƒ@ƒCƒuƒoƒ“ƒhƒoƒ‹ƒu"�B‚Q‚U�`‚Q‚X�Ž�E‚�‚g‚T�D‚O�`‚U�D‚O�B


�Ÿ
�@Desmopuntius rhomboocellatus (Koumans 1940)
�@�@�@syn.Barbus kahajani Hoedeman 1956
�@�@�@�@�@Barbus rhomboocellatus (Koumans 1940)
�@�@�@�@�@Puntius rhomboocellatus Koumans 1940
�@�@�@�@�@? Barbus tetrazonus Bleeker 1856 non Bleeker 1855
�@�@�@�@�@Systomus rhomboocellatus (Koumans 1940)
�@‚Q‚R�`‚Q‚W�Ž�E‚�‚g‚U�D‚T�`‚V�D‚T�B

�Ÿ
�@Desmopuntius trifasciatus (Kottelat 1996)
�@�@�@syn.Puntius trifasciatus Kottelat 1996

---

Genus Eechathalakenda^♀ MENON 1999�@ƒGƒGƒJƒ^ƒ‰ƒPƒ“ƒ_‘®
�@•ª—ÞŠw“IˆÊ’u‚Í•sˆÀ’è�B

�Ÿ
�@Eechathalakenda ophicephala (Raj 1941)
�@�@�@syn.Barbus ophicephalus Raj 1941�@�¦��Barbus (Puntius) Ophicephalus Raj 1941�B
�@�@�@�@�@Puntius ophicephalus (Raj 1941)
�@"ƒ`ƒƒƒ“ƒiƒoƒ‹ƒu"�B

---

Genus Enteromius^♂ COPE 1869�@ƒGƒ“ƒeƒ�ƒ~ƒEƒX‘®
�@�@�@�¦�{Genus Afropuntio^♂ KARAMAN 1971 (Type species : Barbus pobeguini Pellégrin 1911)�B
�@�@�@�¦�{Genus Agrammobarbus^♂ (PELLÉGRIN 1935) (Type species : Barbus apleurogrammus Boulenger 1911)�B
�@�@�@�¦�{Genus Barbus^♂ CUVIER & CLOQUET 1816 Sub-Genus Agrammobarbus^♂ PELLÉGRIN 1935 (Type species : Barbus apleurogrammus Boulenger 1911)�B
�@�@�@�¦�{Genus Barbus^♂ CUVIER & CLOQUET 1816 Sub-Genus Hemigrammocapoeta^♀ ESTÈVE 1952 (Type species : Barbus mirei Estève 1952)�B
�@�@�@�¦�{Genus Barbus^♂ CUVIER & CLOQUET 1816 Sub-Genus Hemigrammopuntius^♂ PELLÉGRIN 1923 (Type species : Barbus salessei Pellégrin 1908)�B
�@�@�@�¦�{Genus Barbus^♂ CUVIER & CLOQUET 1816 Sub-Genus Nicholsopuntius^♂ PELLÉGRIN 1933 (Type species : Barbus candens Nichols & Griscom 1917)�B
�@�@�@�¦�{Genus Barbus^♂ CUVIER & CLOQUET 1816 Sub-Genus Vanderbiltella^♀ FOWLER 1936 (Type species : Barbus lepidurus Fowler 1936)�B
�@�@�@�¦�{Genus Beirabarbus^♂ HERRE 1936 (Type species : Beirabarbus palustris Herre 1936)�B
�@�@�@�¦�{Genus Estevea^♀ WHITLEY 1953 (Type species : Barbus mirei Estève 1952)�B
�@�@�@�¦�{Genus Hemigrammocapoeta^♀ (ESTÈVE 1952) (Type species : Barbus mirei Estève 1952)�B
�@�@�@�¦�‚Genus Hemigrammocapoeta^♀ PELLÉGRIN 1927 [Genus Hemigrammocapoeta]�B
�@�@�@�¦�{Genus Hemigrammopuntius^♂ (PELLÉGRIN 1923) (Type species : Barbus salessei Pellégrin 1908)�B
�@�@�@�¦�{Genus Mannichthys^♂ SCHULTZ 1942 (Type species : Mannichthys lucileae Schultz 1942)�B
�@�@�@�¦�{Genus Nicholsopuntius^♂ (PELLÉGRIN 1933) (Type species : Barbus candens Nichols & Griscom 1917)�B
�@�@�@�¦�{Genus Parapuntius^♂ KARAMAN 1971 (Type species : Barbus anemus Boulenger 1903)�B
�@�@�@�¦�{Genus Vanderbiltella^♀ (FOWLER 1936) (Type species : Barbus lepidurus Fowler 1936)�B
�@ƒAƒtƒŠƒJƒ“�Eƒvƒ“ƒeƒBƒEƒX—Þ�B

�Ÿ
�@Enteromius ablabes (Bleeker 1863)
�@�@�@syn.Barbus ablabes (Bleeker 1863)
�@�@�@�@�@Barbus ablabes spurelli (Boulenger 1913)
�@�@�@�@�@Barbus spurrelli Boulenger 1913
�@�@�@�@�@Puntius ablabes Bleeker 1863

�Ÿ
�@Enteromius aboinensis (Boulenger 1911)
�@�@�@syn.Barbus aboinensis Boulenger 1911
�@IUCN Data Deficient ver.3.1�B

�Ÿ
�@Enteromius afrovernayi (Nichols & Boulton 1927)
�@�@�@syn.Barbus afrovernayi Nichols & Boulton 1927
�@"ƒXƒ|ƒbƒgƒe�[ƒ‹ƒoƒ‹ƒu"�BIUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius aliciae (Bigorne & Lávêque 1993)
�@�@�@syn.Barbus aliciae Bigorne & Lávêque 1993
�@IUCN Endangered�@ B1ab(iii)+2ab(iii)�@ ver.3.1�B

�Ÿ
�@Enteromius aloyi (Roman 1971)
�@�@�@syn.Barbus aloyi Roman 1971
�@IUCN Data Deficient ver.3.1�B

�Ÿ
�@Enteromius amanpoae (Lambert 1961)
�@�@�@syn.Barbus amanpoae Lambert 1961
�@IUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius amatolicus (Skelton 1990)
�@�@�@syn.Barbus amatolicus Skelton 1990
�@"ƒAƒ}ƒgƒ‰ƒoƒ‹ƒu"�BIUCN Vulnerable�@ B2ab(iii)�@ ver.3.1�B

�Ÿ
�@Enteromius anemus (Boulenger 1903)
�@�@�@syn.Barbus anemus Boulenger 1903
�@�@�@�@�@Labeobarbus anemus (Boulenger 1903)
�@�@�@�@�@Parapuntius anemus (Boulenger 1903)
�@^*1939�wLes Barbeaux de l'Afrique du Nord Française, Description d'Une Espèce Nouvelle�FJaques Pellégrin�x�B
�@IUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius annectens (Gilchrist & Thompson 1917)
�@�@�@syn.Barbus annectens Gilchrist & Thompson 1917
�@"ƒuƒ��[ƒhƒXƒgƒ‰ƒCƒvƒoƒ‹ƒu"�BIUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius anniae (Lávêque 1983)
�@�@�@syn.Barbus anniae Lávêque 1983
�@IUCN Vulnerable�@ D2�@ ver.3.1�B

�Ÿ
�@Enteromius anoplus (Weber 1897)
�@�@�@syn.Barbus anoplus Weber 1897
�@�@�@�@�@Barbus anoplus typicus Barnard 1943
�@�@�@�@�@Barbus karkensis Gilchrist & Thompson 1913
�@"ƒ`ƒƒƒr�[ƒwƒbƒhƒoƒ‹ƒu"�B‹D�…ˆæ�i“ü�«�BIUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius ansorgii (Boulenger 1904)
�@�@�@syn.Barbus ansorgii Boulenger 1904
�@IUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius apleurogrammus (Boulenger 1911)
�@�@�@syn.Agrammobarbus apleurogrammus (Boulenger 1911)
�@�@�@�@�@? Agrammobarbus babaulti (Pellégrin 1935)
�@�@�@�@�@Barbus amboseli Banister 1980
�@�@�@�@�@Barbus aphantogrammus Regan 1920
�@�@�@�@�@Barbus apleurogrammus Boulenger 1911
�@�@�@�@�@? Barbus babaulti Pellégrin 1935
�@�@�@�@�@Barbus lapsus Banister 1973
�@�@�@�@�@Barbus scheemanni Klausewitz 1957
�@�@�@�@�@? Barbus zanzibaricus paucior Hilgendorf 1905
�@"ƒC�[ƒXƒgƒAƒtƒŠƒJƒŒƒbƒhƒtƒBƒ“ƒoƒ‹ƒu"�BIUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius arambourgi (Pellégrin 1935)
�@�@�@syn.Barbus arambourgi Pellégrin 1935
�@�@�@�@�@Labeobarbus arambourgi (Pellégrin 1935)
�@IUCN Data Deficient ver.3.1�B

�Ÿ
�@Enteromius arcislongae (Keilhack 1908)
�@�@�@syn.Barbodes arcislongae (Keilhack 1908)
�@�@�@�@�@Barbus arcislongae (Keilhack 1908)
�@�@�@�@�@Barbus trispilopleurus arcislongae Keilhack 1908
�@‹D�…ˆæ�i“ü�«�B‚Q‚S�Ž�E‚�‚g‚W�D‚O�BIUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius argenteus (Günther 1868)
�@�@�@syn.Barbus argenteus Günther 1868
�@�@�@�@�@Barbus crocodilensis Fowler 1934�@IUCN Least Concern ver.3.1�B
�@�@�@�@�@Puntius argenteus (Günther 1868)
�@"ƒ��[ƒYƒtƒBƒ“ƒoƒ‹ƒu"�B

�Ÿ
�@Enteromius aspilus (Boulenger 1907)
�@�@�@syn.Barbus aspilus Boulenger 1907
�@IUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius atakorensis (Daget 1957)
�@�@�@syn.Barbus atakorensis Daget 1957
�@‚Q‚Q�`‚Q‚T�Ž�E‚�‚g‚T�D‚T�`‚U�D‚O�BIUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius atkinsoni (Bailey 1969)
�@�@�@syn.Barbus atkinsoni Bailey 1969
�@"ƒ_ƒbƒVƒ…ƒhƒbƒgƒoƒ‹ƒu"�BIUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius atromaculatus (Nichols & Griscom 1917)
�@�@�@syn.Barbus atromaculatus Nichols & Griscom 1917
�@�@�@�@�@Barbus lepidurus Fowler 1936
�@�@�@�@�@Puntius atromaculatus (Nichols & Griscom 1917)
�@�@�@�@�@Vanderbiltella lepidura (Fowler 1936)
�@IUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius bagbwensis (Norman 1932)

�Ÿ
�@Enteromius baleensis Prokofiev, Levin & Golubtsov 2021
�@^*2021�wA New Species of Enteromius from the Bale Mountain Region, South-Eastern Ethiopia [Teleostei, Cyprinidae]�FArtém Mikhailovich Prokofiev, Boris Alexandrovich Levin & Alexander Serafimovich Golubtsov�x�B
�@IUCN Vulnerable�@ B1ab(iii)+2ab(iii);D2�@ ver.3.1�B

�Ÿ
�@Enteromius barnardi (Jubb 1965)
�@�@�@syn.Barbus barnardi Jubb 1965
�@"ƒuƒ‰ƒbƒNƒoƒbƒNƒoƒ‹ƒu"�BIUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius barotseensis (Pellégrin 1920)
�@�@�@syn.Barbus barotseensis (Pellégrin 1920)
�@�@�@�@�@Barbus radiatus barotseensis Pellégrin 1920
�@IUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius baudoni (Boulenger 1918)
�@�@�@syn.Barbus baudoni Boulenger 1918
�@�@�@�@�@Barbus baudoni ubanguensis Pellégrin 1920
�@�@�@�@�@Barbus svenssoni Johnels 1954
�@�@�@�@�@Barbus voltae Hopson 1965
�@IUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius bawkuensis (Hopson 1965)
�@�@�@syn.Barbus bawkuensis Hopson 1965
�@IUCN Endangered�@ B1ab(iii)+2ab(iii)�@ ver.3.1�B

�Ÿ
�@Enteromius bifrenatus (Fowler 1935)
�@�@�@syn.Barbus bifrenatus
�@"ƒnƒCƒtƒBƒ“ƒoƒ‹ƒu"�BIUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius bigornei (Lávêque, Teugels & van den Audenaerde 1988)
�@�@�@syn.Barbus bigornei Lávêque, Teugels & van den Audenaerde 1988
�@IUCN Near Threatened ver.3.1�B

�Ÿ
�@Enteromius boboi (Schultz 1942)
�@�@�@syn.Barbus boboi Schultz 1942

�Ÿ
�@Enteromius bourdariei (Pellégrin 1928)
�@�@�@syn.Barbus bourdariei Pellégrin 1928
�@IUCN Endangered�@ B1ab(iii)+2ab(iii)�@ ver.3.1�B

�Ÿ
�@Enteromius brachygrammus (Boulenger 1915)
�@�@�@syn.Barbus brachygrammus Boulenger 1915
�@"ƒ‰ƒCƒ“ƒXƒ|ƒbƒgƒoƒ‹ƒu(ƒEƒtƒBƒp�j"�BIUCN Data Deficient ver.3.1�B

�Ÿ
�@Enteromius brazzai (Pellégrin 1901)
�@�@�@syn.Barbus alvarezi Roman 1971
�@�@�@�@�@Barbus brazzai Pellégrin 1901
�@�@�@�@�@Barbus tshopoensis de Vos 1991
�@IUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius breviceps (Trewavas 1936)
�@�@�@syn.Barbus breviceps Trewavas 1936
�@"ƒVƒ‡�[ƒgƒwƒbƒhƒoƒ‹ƒu"�BIUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius brevidorsalis (Boulenger 1915)
�@�@�@syn.Barbus brevidorsalis Boulenger 1915
�@�@�@�@�@Barbus puellus Nichols & Boulton 1927
�@"ƒhƒ��[ƒtƒoƒ‹ƒu"�BIUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius brevilateralis (Poll 1967)
�@�@�@syn.Barbus brevilateralis Poll 1967
�@IUCN Data Deficient ver.3.1�B

�Ÿ
�@Enteromius brevipinnis (Jubb 1966)
�@�@�@syn.Barbus brevipinnis Jubb 1966
�@"ƒVƒ‡�[ƒgƒtƒBƒ“ƒoƒ‹ƒu"�BIUCN Near Threatened ver.3.1�B

�Ÿ
�@Enteromius brichardi (Poll & Lambert 1959)
�@�@�@syn.Barbus brichardi Poll & Lambert 1959
�@IUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius cadenati (Daget 1962)
�@�@�@syn.Barbus cadenati Daget 1962
�@IUCN Vulnerable�@ D2�@ ver.3.1�B

�Ÿ
�@Enteromius callipterus (Boulenger 1907)
�@�@�@syn.Barbodes callipterus (Boulenger 1907)
�@�@�@�@�@Barbus callipterus Boulenger 1907
�@�@�@�@�@Puntius callipterus (Boulenger 1907)
�@"ƒNƒŠƒbƒp�[ƒoƒ‹ƒu"�BIUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius camptacanthus camptacanthus (Bleeker 1863)
�@�@�@syn.Barbodes camptacanthus (Bleeker 1863)
�@�@�@�@�@Barbus camptacanthus Bleeker 1863
�@�@�@�@�@Puntius camptacanthus (Bleeker 1863)
�@�@�@�@�@Puntius tholonianus Thominot 1886
�@"ƒAƒtƒŠƒJƒ“ƒŒƒbƒhƒtƒBƒ“ƒoƒ‹ƒu"�B‚Q‚Q�`‚Q‚W�Ž�E‚�‚g‚U�D‚W�`‚V�D‚W�BIUCN Least Concern ver.3.1�B


�Ÿ
�@Enteromius camptacanthus cottesi (Pellégrin 1907)
�@�@�@syn.Barbus camptacanthus cottesi Pellégrin 1907
�@��Enteromius holotaenius (Boulenger 1904)�H��Enteromius rubrostigmus (Poll & Lambert 1964)�H

�Ÿ
�@Enteromius candens (Nichols & Griscom 1917)
�@�@�@syn.Barbus candens Nichols & Griscom 1917
�@�@�@�@�@Nicholsopuntius candens (Nichols & Griscom 1917)
�@"ƒoƒ^ƒtƒ‰ƒCƒoƒ‹ƒu(ƒAƒvƒŠƒRƒbƒg�j"�B‚Q‚S�`‚Q‚U�Ž�E‚�‚g‚T�D‚O�`‚U�D‚O�BIUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius carcharhinoides (Stiassny 1991)
�@�@�@syn.Barbus carcharhinoides Stiassny 1991
�@IUCN Critically Endangered�@ B1ab(iii)+2ab(iii)�@ ver.3.1�B

�Ÿ
�@Enteromius carens (Boulenger 1912)
�@�@�@syn.Barbus carens�@Boulenger 1912
�@IUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius castrasibutus (Fowler 1936)
�@�@�@syn.Barbus castrasibutus Fowler 1936
�@IUCN Data Deficient ver.3.1�B

�Ÿ
�@Enteromius catenarius (Poll & Lambert 1959)
�@�@�@syn.Barbus catenarius Poll & Lambert 1959
�@IUCN Data Deficient ver.3.1�B

�Ÿ
�@Enteromius caudosignatus (Poll 1967)
�@�@�@syn.Barbus caudosignatus Poll 1967
�@IUCN Data Deficient ver.3.1�B

�Ÿ
�@Enteromius cercops (Whitehead 1960)
�@�@�@syn.Barbus cercops Whitehead 1960
�@"ƒ‹ƒAƒ“ƒuƒ�ƒoƒ‹ƒu"�B

�Ÿ
�@Enteromius cernuus (Barnard 1938)
�@�@�@syn.Barbus anoplus cernuus (Barnard 1938)
�@�@�@�@�@Barbus cernuus Barnard 1938
�@^*2021�wAllopatric Differentiation in the Enteromius anoplus Complex in South Africa, with the Revalidation of E.cernuus and E.oraniensis and Description of a New Species, E.mandelai [Teleostei, Cyprinidae]�FManda Juliet Kambikambi, Wilbert T.Kadye & Albert Chakona�x�B

�Ÿ
�@Enteromius chicapaensis (Poll 1967)
�@�@�@syn.Barbus chicapaensis Poll 1967
�@IUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius chiumbeensis (Pellégrin 1936)
�@�@�@syn.Barbus chiumbeensis
�@IUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius chlorotaenius (Boulenger 1911)
�@�@�@syn.Barbus chlorotaenius Boulenger 1911
�@IUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius choloensis (Norman 1925)
�@�@�@syn.Barbus choloensis Norman 1925
�@IUCN Vulnerable�@ D2�@ ver.3.1�B

�Ÿ
�@Enteromius citrinus (Boulenger 1920)
�@�@�@syn.Barbus citrinus Boulenger 1920
�@IUCN Data Deficient ver.3.1�B

�Ÿ
�@Enteromius clauseni (van den Audenaerde 1976)
�@�@�@syn.Barbus clauseni van den Audenaerde 1976
�@IUCN Data Deficient ver.3.1�B

�Ÿ
�@Enteromius collarti (Poll 1945)
�@�@�@syn.Barbus collarti Poll 1945
�@IUCN Vulnerable�@ D2�@ ver.3.1�B

�Ÿ
�@Enteromius condei (Mahnert & Géry 1982)
�@�@�@syn.Barbus condei Mahnert & Géry 1982

�Ÿ
�@Enteromius deguidei (Matthes 1964)
�@�@�@syn.Barbus deguidei Matthes 1964
�@IUCN Data Deficient ver.3.1�B

�Ÿ
�@Enteromius deserti (Pellégrin 1909)
�@�@�@syn.Barbus deserti Pellégrin 1909
�@^*1939�wLes Barbeaux de l'Afrique du Nord Française, Description d'Une Espèce Nouvelle�FJaques Pellégrin�x�B
�@"ƒTƒnƒ‰ƒoƒ‹ƒu"�BIUCN Near Threatened ver.3.1�B

�Ÿ
�@Enteromius devosi (Banyankimbona, Vreven & Snoeks 2012)
�@�@�@syn.Barbus devosi Banyankimbona, Vreven & Snoeks 2012

�Ÿ
�@Enteromius dialonensis (Daget 1962)
�@�@�@syn.Barbus dialonensis Daget 1962
�@IUCN Vulnerable�@ B1ab(iii)+2ab(iii);D2�@ ver.3.1�B

�Ÿ
�@Enteromius diamouanganai (Teugels & Mamonekene 1992)
�@�@�@syn.Barbus diamouanganai Teugels & Mamonekene 1992
�@IUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius ditinensis (Daget 1962)
�@�@�@syn.Barbus ditinensis Daget 1962
�@IUCN Vulnerable�@ B1ab(iii)+2ab(iii);D2�@ ver.3.1�B

�Ÿ
�@Enteromius dorsolineatus (Trewavas 1936)
�@�@�@syn.Barbus dorsolineatus Trewavas 1936
�@"ƒNƒlƒlƒoƒ‹ƒu"�BIUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius eburneensis (Poll 1941)
�@�@�@syn.Barbus eburneensis Poll 1941
�@�@�@�@�@Barbus flomoi Schultz 1942
�@‚Q‚Q�`‚Q‚T�Ž�E‚�‚g‚T�D‚T�BIUCN Vulnerable�@ B1ab(iii)+2ab(iii);D2�@ ver.3.1�B

�Ÿ
�@Enteromius erythrozonus (Poll & Lambert 1959)
�@�@�@syn.Barbus erythrozonus Poll & Lambert 1959
�@IUCN Data Deficient ver.3.1�B

�Ÿ
�@Enteromius eutaenius (Boulenger 1904)
�@�@�@syn.Barbus eutaenius Boulenger 1904
�@"ƒIƒŒƒ“ƒWƒtƒBƒ“ƒoƒ‹ƒu"�B‚Q‚O�`‚Q‚U�Ž�E‚�‚g‚U�D‚T�`‚V�D‚T�BIUCN Data Deficient ver.3.1�B

�Ÿ
�@Enteromius evansi (Fowler 1930)
�@�@�@syn.Barbus evansi Fowler 1930
�@IUCN Least Concern ver.3.1�B


�Ÿ
�@Enteromius fasciolatus (Günther 1868)
�@�@�@syn.Barbodes barilioides Boulenger 1914
�@�@�@�@�@Barbodes fasciolatus (Günther 1868)
�@�@�@�@�@Barbus barilioides Boulenger 1914
�@�@�@�@�@Barbus fasciolatus Günther 1868
�@�@�@�@�@Puntius fasciolatus (Günther 1868)
�@"ƒAƒtƒŠƒJƒ“(ƒoƒ“ƒh�jƒoƒ‹ƒu"�B‚Q‚O�`‚Q‚U�Ž�E‚�‚g‚U�D‚O�BIUCN Least Concern ver.3.1�B


�Ÿ
�@Enteromius foutensis (Lávêque, Teugels & van den Audenaerde 1988)
�@�@�@syn.Barbus foutensis Lávêque, Teugels & van den Audenaerde 1988
�@IUCN Vulnerable�@ B2ab(iii);D2�@ ver.3.1�B

�Ÿ
�@Enteromius gibbosus (Peters 1852)
�@�@�@syn.Barbus gibbosus Peters 1852 non Valenciennes 1842
�@��Enteromius paludinosus (Peters 1852)�H

�Ÿ
�@Enteromius greenwoodi (Poll 1967)
�@�@�@syn.Barbus greenwoodi Poll 1967
�@IUCN Data Deficient ver.3.1�B

�Ÿ
�@Enteromius guildi (Loiselle 1973)
�@�@�@syn.Barbus guildi Loiselle 1973
�@IUCN Data Deficient ver.3.1�B

�Ÿ
�@Enteromius guineensis (Pellégrin 1913)
�@�@�@syn.Barbus guineensis Pellégrin 1913
�@IUCN Data Deficient ver.3.1�B

�Ÿ
�@Enteromius guirali (Thominot 1886)
�@�@�@syn.Barbus camptacanthus melanepipterus (Pellégrin 1924)
�@�@�@�@�@Barbus guirali Thominot 1886
�@�@�@�@�@Barbus guirali melanepipterus (Pellégrin 1924)
�@�@�@�@�@Barbus melanepipterus Pellégrin 1924
�@IUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius gurneyi (Günther 1868)
�@�@�@syn.Barbus gurneyi Günther 1868
�@"ƒŒƒbƒhƒe�[ƒ‹ƒoƒ‹ƒu"�BIUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius haasianus (David 1936)
�@�@�@syn.Barbus haasianus David 1936
�@�@�@�@�@Barbus woehlerti Trewavas 1938
�@�@�@�@�@Puntius wöhlerti (Trewavas 1938)
�@"ƒVƒbƒNƒ‹ƒoƒ‹ƒu"�BIUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius hamiltoni (Gilchrist & Thompson 1913)
�@�@�@syn.Barbus afrohamiltoni Crass 1960
�@�@�@�@�@Barbus hamiltoni Gilchrist & Thompson 1913 non Jerdon 1849
�@�@�@�@�@Enteromius afrohamiltoni (Crass 1960)�@�¦ƒtƒBƒbƒVƒ…ƒx�[ƒX‚Ì—LŒø–¼�Ì�B
�@"ƒnƒ~ƒ‹ƒgƒ“ƒoƒ‹ƒu"�BIUCN Least Concern ver.3.1�B


�Ÿ
�@Enteromius holotaenius (Boulenger 1904)
�@�@�@syn.Barbus holotaenius Boulenger 1904
�@�@�@�@�@Puntius holotaenius (Boulenger 1904)
�@"ƒXƒ|ƒbƒgƒXƒP�[ƒ‹ƒoƒ‹ƒu"�BIUCN Least Concern ver.3.1�B

�Ÿ
�@Barbus holotaenius macracanthus Pellégrin 1930
�@��Enteromius prionacanthus (Mahnert & Géry 1982)�H��Enteromius rubrostigmus (Poll & Lambert 1964)�H

�Ÿ
�@Barbus holotaenius ovomaculatus Pellégrin 1930

�Ÿ
�@Enteromius huguenyi (Bigorne & Lávêque 1993)
�@�@�@syn.Barbus huguenyi Bigorne & Lávêque 1993
�@IUCN Endangered�@ B1ab(iii)+2ab(iii)�@ ver.3.1�B

�Ÿ
�@Enteromius hulstaerti (Poll 1945)
�@�@�@syn.Barbus hulstaerti Poll 1945
�@�@�@�@�@Capoeta hulstaerti (Poll 1945)
�@�@�@�@�@Puntius hulstaerti (Poll 1945)
�@"ƒoƒ^ƒtƒ‰ƒCƒoƒ‹ƒu"�BIUCN Least Concern ver.3.1�B


�Ÿ
�@Enteromius humeralis (Boulenger 1902)
�@�@�@syn.Barbus dolichosomus Nichols & Griscom 1917
�@�@�@�@�@Barbus humeralis Boulenger 1902
�@IUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius humilis (Boulenger 1902)
�@�@�@syn.Barbus humilis Boulenger 1902

�Ÿ
�@Enteromius inaequalis (Lávêque, Teugels & van den Audenaerde 1988)
�@�@�@syn.Barbus inaequalis Lávêque, Teugels & van den Audenaerde 1988
�@IUCN Data Deficient ver.3.1�B

�Ÿ
�@Enteromius innocens (Pfeffer 1896)
�@�@�@syn.Barbus innocens Pfeffer 1896
�@"ƒCƒ“ƒRƒ“ƒXƒsƒLƒ…ƒI�[ƒXƒoƒ‹ƒu"�BIUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius jacksoni (Günther 1889)
�@�@�@syn.Barbus jacksoni Günther 1889
�@�@�@�@�@Barbus jacksonii mitior Hilgendorf 1905
�@�@�@�@�@Barbus nummifer Boulenger 1904
�@�@�@�@�@Barbus pappenheimi Boulenger 1905
�@"ƒWƒƒƒNƒ\ƒ“ƒoƒ‹ƒu"�BIUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius jae (Boulenger 1903)
�@�@�@syn.Barbus jae Boulenger 1903
�@"ƒ„ƒGƒoƒ‹ƒu"�B‚Q‚P�`‚Q‚T�Ž�E‚�‚g‚T�D‚T�`‚U�D‚O�BIUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius janssensi (Poll 1976)
�@�@�@syn.Barbus janssensi Poll 1976
�@IUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius kamolondoensis (Poll 1938)
�@�@�@syn.Barbus kamolondoensis Poll 1938
�@IUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius kerstenii (Peters 1868)
�@�@�@syn.Barbus akeleyi Hubbs 1918
�@�@�@�@�@Barbus kerstenii Peters 1868
�@�@�@�@�@Barbus lumiensis Boulenger 1903
�@�@�@�@�@Barbus minchinii Boulenger 1906
�@�@�@�@�@Barbus nigrolineus Pfeffer 1889
�@�@�@�@�@Barbus salmo Pfeffer 1896
�@�@�@�@�@Barbus tangandensis Jubb 1954
�@�@�@�@�@? Barbus zanzibaricus paucior Hilgendorf 1905
�@�@�@�@�@Labeobarbus kerstenii (Peters 1868)
�@"ƒŒƒbƒhƒXƒ|ƒbƒgƒoƒ‹ƒu"�BIUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius kessleri (Steindachner 1866)
�@�@�@syn.Barbus caudimaculus Günther 1868
�@�@�@�@�@Barbus kessleri (Steindachner 1866)
�@�@�@�@�@Puntius caudimaculus (Günther 1868)
�@�@�@�@�@Puntius kessleri Steindachner 1866
�@"ƒMƒ‹ƒo�[ƒoƒ‹ƒu"�BIUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius kissiensis (Daget 1954)
�@�@�@syn.Barbus kissiensis Daget 1954
�@IUCN Vulnerable�@ D2�@ ver.3.1�B

�Ÿ
�@Enteromius kuiluensis (Pellégrin 1930)
�@�@�@syn.Barbus kuiluensis Pellégrin 1930
�@IUCN Data Deficient ver.3.1�B

�Ÿ
�@Enteromius lamani (Lönnberg & Rendahl 1920)
�@�@�@syn.Barbus lamani Lönnberg & Rendahl 1920
�@IUCN Data Deficient ver.3.1�B

�Ÿ
�@Enteromius laticeps (Pfeffer 1889)
�@�@�@syn.Barbus laticeps Pfeffer 1889
�@IUCN Vulnerable�@ D2�@ ver.3.1�B

�Ÿ
�@Enteromius lauzannei (Lávêque & Paugy 1982)
�@�@�@syn.Barbus lauzannei Lávêque & Paugy 1982
�@IUCN Endangered�@ B1ab(iii)+2ab(iii)�@ ver.3.1�B

�Ÿ
�@Enteromius leonensis (Boulenger 1915)
�@�@�@syn.Barbus leonensis Boulenger 1915
�@IUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius liberiensis (Steindachner 1894)
�@�@�@syn.Barbus camptacanthus liberiensis Steindachner 1894
�@�@�@�@�@Barbus liberiensis (Steindachner 1894)
�@IUCN Endangered�@ B2ab(iii)�@ ver.3.1�B

�Ÿ
�@Enteromius lineomaculatus (Boulenger 1903)
�@�@�@syn.Barbus lineomaculatus Boulenger 1903
�@�@�@�@�@Puntius lineomaculatus (Boulenger 1903)
�@"ƒ‰ƒCƒ“ƒXƒ|ƒbƒgƒo�[ƒxƒ‹"�BIUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius litambus (Keilhack 1908)
�@�@�@syn.Barbus litambus Keilhack 1908
�@�@�@�@�@Barbus rhoadesii Boulenger 1908
�@�@�@�@�@Labeobarbus litambus (Keilhack 1908)
�@IUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius lornae (Ricardo-Bertram 1943)
�@�@�@syn.Barbus lornae Ricardo-Bertram 1943
�@IUCN Data Deficient ver.3.1�B

�Ÿ
�@Enteromius loveridgii (Boulenger 1916)
�@�@�@syn.Barbus loveridgii Boulenger 1916
�@IUCN Data Deficient ver.3.1�B

�Ÿ
�@Enteromius lufukiensis (Boulenger 1917)
�@�@�@syn.Barbus lufukiensis Boulenger 1917
�@IUCN Near Threatened ver.3.1�B

�Ÿ
�@Enteromius luikae (Ricardo 1939)
�@�@�@syn.Barbus luikae Ricardo 1939
�@"ƒ‹ƒCƒJƒo�[ƒxƒ‹"�BIUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius lujae (Boulenger 1913)
�@�@�@syn.Barbus lujae Boulenger 1913
�@IUCN Data Deficient ver.3.1�B

�Ÿ
�@Enteromius lukindae (Boulenger 1915)
�@�@�@syn.Barbus lukindae Boulenger 1915
�@IUCN Data Deficient ver.3.1�B

�Ÿ
�@Enteromius lukusiensis (David & Poll 1937)
�@�@�@syn.Barbus lukusiensis David & Poll 1937
�@IUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius luluae (Fowler 1930)
�@�@�@syn.Barbus luluae Fowler 1930
�@IUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius machadoi (Poll 1967)
�@�@�@syn.Barbus machadoi Poll 1967
�@IUCN Data Deficient ver.3.1�B

�Ÿ
�@Enteromius macinensis (Daget 1954)
�@�@�@syn.Barbus macinensis Daget 1954
�@IUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius macrops (Boulenger 1911)
�@�@�@syn.Barbodes macrops (Boulenger 1911)
�@�@�@�@�@Barbus francisci Boulenger 1916
�@�@�@�@�@Barbus gambiensis Svensson 1933
�@�@�@�@�@Barbus macrops Boulenger 1911
�@�@�@�@�@Barbus weidholzii Holly 1928
�@�@�@�@�@Mannichthys lucileae Schultz 1942
�@�@�@�@�@Puntius gambiensis (Svensson 1933)
�@^*2022�wAn Annotated Catalogue of the Type Specimens Described by Maximilian Holly Housed in the Natural History Museum of Vienna 1, [Chordata, Actinopterygii] and [Echinodermata, Asteroidea]�FNina Gidalevna Bogutskaya, Ernst Mikschi, Mario Dominik Riedl, Stefan Szeiler, Pedro R.Frade & Anja Palandačić�x�B
�@"ƒuƒ‰ƒbƒNƒXƒgƒ‰ƒCƒvƒoƒ‹ƒu"�BIUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius macrotaenius (Worthington 1933)
�@�@�@syn.Barbus macrotaenius Worthington 1933
�@"ƒuƒ��[ƒhƒoƒ“ƒhƒoƒ‹ƒu"�BIUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius magdalenae (Boulenger 1906)
�@�@�@syn.Barbus magdalenae Boulenger 1906
�@"ƒuƒ“ƒWƒƒƒRƒoƒ‹ƒu"�BIUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius mandelai Kambikambi, Kadye & Chakona 2021
�@^*2021�wAllopatric Differentiation in the Enteromius anoplus Complex in South Africa, with the Revalidation of E.cernuus and E.oraniensis and Description of a New Species, E.mandelai [Teleostei, Cyprinidae]�FManda Juliet Kambikambi, Wilbert T.Kadye & Albert Chakona�x�B

�Ÿ
�@Enteromius manicensis (Pellégrin 1919)
�@�@�@syn.Barbus hondeensis Määr 1962
�@�@�@�@�@Barbus manicensis Pellégrin 1919
�@"ƒCƒGƒ��[ƒoƒ‹ƒu"�BIUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius marmoratus (David & Poll 1937)
�@�@�@syn.Barbus marmoratus David & Poll 1937
�@IUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius martorelli (Roman 1971)
�@�@�@syn.Barbus martorelli Roman 1971
�@IUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius mattozi (Guimarães 1884)
�@�@�@syn.Barbus mattozi Guimarães 1884
�@�@�@�@�@Barbus rapax Steindachner 1894
�@�@�@�@�@Barbus sauvagei Pellégrin 1912
�@�@�@�@�@Barbus serrulus Gilchrist & Thompson 1913
�@"ƒy�[ƒp�[ƒ}ƒEƒX"�BIUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius mediosquamatus (Poll 1967)
�@�@�@syn.Barbus mediosquamatus Poll 1967
�@IUCN Data Deficient ver.3.1�B

�Ÿ
�@Enteromius melanotaenius (Stiassny 1991)
�@�@�@syn.Barbus melanotaenius Stiassny 1991
�@IUCN Critically Endangered�@ B1ab(iii)+2ab(iii)�@ ver.3.1�B

�Ÿ
�@Enteromius mimus (Boulenger 1912)
�@�@�@syn.Barbus mimus Boulenger 1912
�@"ƒGƒ�ƒ\ƒ“ƒMƒ�ƒoƒ‹ƒu"�BIUCN Least Concern ver.3.1�B


�Ÿ
�@Enteromius miolepis (Boulenger 1902)
�@�@�@syn.Barbus decioi Fowler 1958�@�¦Barbus nicholsi Vinciguerra 1928‚Ì•Ï–¼�B
�@�@�@�@�@Barbus miolepis Boulenger 1902
�@�@�@�@�@Barbus nicholsi Vinciguerra 1928 non Myers 1924
�@�@�@�@�@Barbus squamosissimus Steindachner 1912
�@�@�@�@�@Barbus treadwelli Pellégrin 1933
�@^*1958�wSome New Taxonomic Names of Fishlike Vertebrates�FHenry Weed Fowler�x�B
�@"ƒWƒOƒUƒOƒoƒ‹ƒu"�BIUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius mocoensis (Trewavas 1936)
�@�@�@syn.Barbus mocoensis Trewavas 1936
�@IUCN Data Deficient ver.3.1�B

�Ÿ
�@Enteromius mohasicus (Pappenheim 1914)
�@�@�@syn.Barbus mohasicus Pappenheim 1914
�@�@�@�@�@Barbus mohasicus paucisquamatus Pellégrin 1933

�Ÿ
�@Enteromius motebensis (Steindachner 1894)
�@�@�@syn.Barbus kamaiae (David & Poll 1937)
�@�@�@�@�@Barbus motebensis Steindachner 1894
�@�@�@�@�@Barbus motebensis kamaiae David & Poll 1937
�@"ƒ}ƒŠƒRƒoƒ‹ƒu"�BIUCN Vulnerable�@ B1ab(i,ii,iii,iv,v)+2ab(i,ii,iii,iv,v)�@ ver.3.1�B

�Ÿ
�@Enteromius multilineatus (Worthington 1933)
�@�@�@syn.Barbus carpenteri (Fowler 1949)
�@�@�@�@�@Barbus multilineatus Worthington 1933
�@�@�@�@�@Puntius carpenteri Fowler 1949
�@"ƒJƒbƒp�[ƒXƒgƒ‰ƒCƒvƒoƒ‹ƒu"�BIUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius musumbi (Boulenger 1910)
�@�@�@syn.Barbus musumbi Boulenger 1910
�@IUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius neefi (Greenwood 1962)
�@�@�@syn.Barbus neefi Greenwood 1962
�@"ƒTƒCƒhƒXƒ|ƒbƒgƒoƒ‹ƒu"�BIUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius neglectus (Boulenger 1903)
�@�@�@syn.Barbus neglectus Boulenger 1903
�@�@�@�@�@Labeobarbus neglectus (Boulenger 1903)
�@IUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius neumayeri (Fischer 1884)
�@�@�@syn.Barbus carpio Pfeffer 1896
�@�@�@�@�@Barbus kerstenii luhondo (Pappenheim 1914)
�@�@�@�@�@Barbus luazomelus Lönnberg 1911
�@�@�@�@�@Barbus luhondo Pappenheim 1914
�@�@�@�@�@Barbus nairobiensis Boulenger 1911
�@�@�@�@�@Barbus neumayeri Fischer 1884
�@�@�@�@�@Barbus percivali Boulenger 1903
�@�@�@�@�@Barbus percivali kitalensis Pellégrin 1935
�@�@�@�@�@Barbus portali Boulenger 1906
�@�@�@�@�@Barbus serrifer Boulenger 1900
�@"ƒmƒCƒ}ƒCƒ„�[ƒoƒ‹ƒu"�BIUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius nigeriensis (Boulenger 1903)
�@�@�@syn.Barbus nigeriensis Boulenger 1903
�@�@�@�@�@Puntius nigeriensis (Boulenger 1903)
�@IUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius nigrifilis (Nichols 1928)
�@�@�@syn.Barbus nigrifilis Nichols 1928
�@IUCN Data Deficient ver.3.1�B

�Ÿ
�@Enteromius nigroluteus (Pellégrin 1930)
�@�@�@syn.Barbus nigroluteus Pellégrin 1930
�@IUCN Endangered�@ B1ab(iii)+2ab(iii)�@ ver.3.1�B

�Ÿ
�@Enteromius niokoloensis (Daget 1959)
�@�@�@syn.Barbus niokoloensis Daget 1959
�@IUCN Vulnerable�@ D2�@ ver.3.1�B

�Ÿ
�@Enteromius nounensis (van den Bergh & Teugels 1998)
�@�@�@syn.Barbus nounensis van den Bergh & Teugels 1998
�@IUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius nyanzae (Whitehead 1960)
�@�@�@syn.Barbus nyanzae Whitehead 1960
�@"ƒjƒƒƒ“ƒUƒoƒ‹ƒu"�B‚P‚W�`‚Q‚W�Ž�E‚�‚g‚U�D‚T�`‚V�D‚T�BIUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius okae (Fowler 1949)
�@�@�@syn.Barbus okae (Fowler 1949)
�@�@�@�@�@Puntius okae Fowler 1949
�@IUCN Data Deficient ver.3.1�B

�Ÿ
�@Enteromius oligogrammus (David 1937)
�@�@�@syn.Barbus oligogrammus David 1937
�@ŒÎŽY�B‚Q‚Q�`‚Q‚V�Ž�E‚�‚g‚V�D‚Q�`‚W�D‚O�BIUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius olivaceus (Seegers 1996)
�@�@�@syn.Barbus olivaceus Seegers 1996
�@"ƒIƒŠ�[ƒuƒOƒŠ�[ƒ“ƒoƒ‹ƒu(ƒEƒtƒBƒp�j"�BIUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius oraniensis (Barnard 1943)
�@�@�@syn.Barbus anoplus oraniensis Barnard 1943
�@^*2021�wAllopatric Differentiation in the Enteromius anoplus Complex in South Africa, with the Revalidation of E.cernuus and E.oraniensis and Description of a New Species, E.mandelai [Teleostei, Cyprinidae]�FManda Juliet Kambikambi, Wilbert T.Kadye & Albert Chakona�x�B

�Ÿ
�@Enteromius owenae (Ricardo-Bertram 1943)
�@�@�@syn.Barbus owenae Ricardo-Bertram 1943
�@IUCN Data Deficient ver.3.1�B

�Ÿ
�@Enteromius pallidus (Smith 1841)
�@�@�@syn.Barbus hemipleurogrammus Boulenger 1911
�@�@�@�@�@Barbus pallidus Smith 1841
�@"ƒS�[ƒ‹ƒfƒB�[ƒoƒ‹ƒu"�BIUCN Least Concern ver.3.1�B


�Ÿ
�@Enteromius paludinosus (Peters 1852)
�@�@�@syn.Barbus akakianus Boulenger 1911
�@�@�@�@�@Barbus amphigrammus Boulenger 1903
�@�@�@�@�@Barbus helleri Hubbs 1918
�@�@�@�@�@Barbus ivongoensis Fowler 1935
�@�@�@�@�@Barbus longicaudus Boulenger 1905
�@�@�@�@�@Barbus macropristis Boulenger 1903
�@�@�@�@�@Barbus macropristis meruensis Lönnberg 1907
�@�@�@�@�@Barbus meruensis (Lönnberg 1907)
�@�@�@�@�@Barbus paludinosus Peters 1852
�@�@�@�@�@Barbus taitensis Günther 1868
�@�@�@�@�@Barbus thikensis Boulenger 1905
�@�@�@�@�@Barbus tsotsorogensis Fowler 1935
�@�@�@�@�@Barbus vinciguerrai Pfeffer 1896
�@�@�@�@�@Barbus welwitschii Günther 1868
�@�@�@�@�@Puntius paludinosus (Peters 1852)
�@"ƒXƒgƒŒ�[ƒgƒtƒBƒ“ƒoƒ‹ƒu"�B Barbus gibbosus Peters 1852 ‚̓”ƒ@ƒ‰ƒ“ƒVƒGƒ“ƒk‚ª�æŽæ‚è‚Å–³Œø�BIUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius papilio (Banister & Bailey 1979)
�@�@�@syn.Barbus papilio Banister & Bailey 1979
�@IUCN Data Deficient ver.3.1�B

�Ÿ
�@Enteromius parablabes (Daget 1957)
�@�@�@syn.Barbus parabablabes (Daget 1957)
�@�@�@�@�@Barbus parablabes Daget 1957
�@IUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius paradoxus (Poll 1938)
�@�@�@syn.Barbus paradoxus Poll 1938
�@��Enteromius radiatus (Peters 1853)�H

�Ÿ
�@Enteromius parajae (van den Bergh & Teugels 1998)
�@�@�@syn.Barbus parajae van den Bergh & Teugels 1998
�@IUCN Least Concern ver.3.1�B


�Ÿ
�@Enteromius pellegrini (Poll 1939)
�@�@�@syn.Barbus pellegrini Poll 1939
�@�@�@�@�@Barbus serrifer trimaculatus Pellégrin 1935
�@"ƒyƒŒƒOƒŠƒ“ƒoƒ‹ƒu"�BIUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius perince (Rüppell 1835)
�@�@�@syn.Barbus donaldson-smithi Fowler 1958
�@�@�@�@�@Barbus lawrae Hopson 1965
�@�@�@�@�@Barbus lepidus Pfaff 1933
�@�@�@�@�@Barbus perince Rüppell 1835
�@^*1958�wSome New Taxonomic Names of Fishlike Vertebrates�FHenry Weed Fowler�x�B
�@IUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius petchkovskyi (Poll 1967)
�@�@�@syn.Barbus petchkovskyi Poll 1967
�@IUCN Data Deficient ver.3.1�B

�Ÿ
�@Enteromius pinnimaculatus Mipounga, Cutler, Beh, Adam & Sidlauskas 2019
�@^*2019�wEnteromius pinnimaculatus sp.nov. [Cypriniformes, Cyprinidae] from Southern Gabon�FH.K.Mipounga, J.Cutler, J.H.Mve Beh, B.Adam & B.L.Sidlauskas�x�B

�Ÿ
�@Enteromius pleurogrammus (Boulenger 1902)
�@�@�@syn.Barbus pleurogrammus Boulenger 1902
�@IUCN Data Deficient ver.3.1�B

�Ÿ
�@Enteromius pobeguini (Pellégrin 1911)
�@�@�@syn.Afropuntio pobeguini (Pellégrin 1911)
�@�@�@�@�@Barbus mirei Estève 1952�@�¦��Barbus (Hemigrammocapoeta) mirei Estève 1952�B
�@�@�@�@�@Barbus pobeguini Pellégrin 1911
�@�@�@�@�@Barbus pobeguini var.mauritanicus Pellégrin 1937
�@�@�@�@�@Barbus pobeguini mauritanicus (Pellégrin 1937)
�@�@�@�@�@Estevea mirei (Estève 1952) Whitley 1953
�@�@�@�@�@Hemigrammocapoeta mirei (Estève 1952)
�@^*1939�wLes Barbeaux de l'Afrique du Nord Française, Description d'Une Espèce Nouvelle�FJaques Pellégrin�x�B
�@^*1953�wStudies in Ichthyology 16�FGilbert Percy Whitley�x�B
�@IUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius poechii (Steindachner 1911)
�@�@�@syn.Barbus bernardcarpi Jubb 1958
�@�@�@�@�@Barbus poechii (Steindachner 1911)
�@�@�@�@�@Barbus pöchii Steindachner 1911
�@"ƒ_ƒbƒVƒ…ƒe�[ƒ‹ƒoƒ‹ƒu"�BIUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius potamogalis Cope 1869
�@^*1869�wSynopsis of the Cyprinidæ of Pennsylvania�FEdward Drinker Cope�x�B
�@IUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius prionacanthus (Mahnert & Géry 1982)
�@�@�@syn.Barbus prionacanthus Mahnert & Géry 1982
�@IUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius profundus (Greenwood 1970)
�@�@�@syn.Barbus profundus (Greenwood 1970)
�@�@�@�@�@Barbus radiatus profundus Greenwood 1970
�@IUCN Data Deficient ver.3.1�B

�Ÿ
�@Enteromius pseudotoppini (Seegers 1996)
�@�@�@syn.Barbus pseudotoppini Seegers 1996
�@IUCN Vulnerable�@ D2�@ ver.3.1�B

�Ÿ
�@Enteromius pumilus (Boulenger 1901)
�@�@�@syn.Barbus pumilus Boulenger 1901
�@IUCN Data Deficient ver.3.1�B

�Ÿ
�@Enteromius punctitaeniatus (Daget 1954)
�@�@�@syn.Barbus punctitaeniatus Daget 1954
�@IUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius pygmaeus (Poll & Gosse 1963)
�@�@�@syn.Barbus pygmaeus Poll & Gosse 1963
�@IUCN Data Deficient ver.3.1�B

�Ÿ
�@Enteromius quadrilineatus (David 1937)
�@�@�@syn.Barbus lineomaculatus quadrilineatus David 1937
�@�@�@�@�@Barbus quadrilineatus (David 1937)
�@IUCN Endangered�@ B1ab(i,ii,iii,iv,v)+2ab(i,ii,iii,iv,v)�@ ver.3.1�B

�Ÿ
�@Enteromius quadripunctatus (Pfeffer 1896)
�@�@�@syn.Barbus quadripunctatus Pfeffer 1896
�@"ƒtƒH�[ƒXƒ|ƒbƒgƒoƒ‹ƒu"�B‚Q‚R�`‚Q‚V�Ž�E‚�‚g‚U�D‚W�`‚V�D‚T�BIUCN Data Deficient ver.3.1�B

�Ÿ
�@Enteromius radari Kisekelwa, Snoeks, Decru, Schedel, Isumbisho & Vreven 2022
�@^*2022�wA Mismatch Between Morphological and Molecular Data in Lineages of Enteromius [Cypriniformes, Cyprinidae] from the Lowa Basin, East Democratic Republic of the Congo DRC, with the Description of a New Species�FTchalondawa Kisekelwa, Jos Snoeks, Eva Decru, Frederic Dieter Benedikt Schedel, Mwapu Isumbisho & Emmanuel Vreven�x�B

�Ÿ
�@'Enteromius radiatus (Günther 1868)'
�@�@�@syn.Barbus radiatus Günther 1868 non Peters 1853
�@—vŽí�¬–¼•Ï�X�B��Enteromius radiatus (Peters 1853)�H

�Ÿ
�@Enteromius radiatus (Peters 1853)
�@�@�@syn.Barbus aurantiacus Boulenger 1910
�@�@�@�@�@Barbus bangwelensis Boulenger 1905
�@�@�@�@�@Barbus doggetti Boulenger 1904
�@�@�@�@�@Barbus myersi Poll 1939
�@�@�@�@�@Barbus okavangoensis (Barnard 1941)
�@�@�@�@�@Barbus palustris (Herre 1936)
�@�@�@�@�@Barbus radiatus Peters 1853 non Günther 1868
�@�@�@�@�@Barbus radiatus aurantiacus (Boulenger 1910)
�@�@�@�@�@Barbus rogersi Boulenger 1911
�@�@�@�@�@Barbus rubellus Crass 1960
�@�@�@�@�@Beirabarbus aurantiacus (Boulenger 1910)
�@�@�@�@�@Beirabarbus okavangoensis Barnard 1941
�@�@�@�@�@Beirabarbus palustris Herre 1936
�@"ƒŒƒbƒhƒAƒCƒoƒ‹ƒu"�BIUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius raimbaulti (Daget 1962)
�@�@�@syn.Barbus raimbaulti Daget 1962
�@IUCN Vulnerable�@ B1ab(iii)+2ab(iii)�@ ver.3.1�B

�Ÿ
�@Enteromius rohani (Pellégrin 1921)
�@�@�@syn.Barbus rohani Pellégrin 1921

�Ÿ
�@Enteromius roussellei (Ladiges & Voelker 1961)
�@�@�@syn.Barbus roussellei Ladiges & Voelker 1961
�@IUCN Data Deficient ver.3.1�B

�Ÿ
�@Enteromius rouxi (Daget 1962)
�@�@�@syn.Barbus rouxi Daget 1962
�@IUCN Data Deficient ver.3.1�B

�Ÿ
�@Enteromius rubrostigmus (Poll & Lambert 1964)
�@�@�@syn.Barbus miolepis rubrostigmus Poll & Lambert 1964
�@�@�@�@�@Barbus rubrostigmus (Poll & Lambert 1964)
�@IUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius salessei (Pellégrin 1908)
�@�@�@syn.Barbus apogonostomatus Pellégrin 1913
�@�@�@�@�@Barbus salessei Pellégrin 1908
�@�@�@�@�@Hemigrammopuntius salessei (Pellégrin 1908)
�@IUCN Vulnerable�@ B1ab(ii,iii)+2ab(ii,iii);D2�@ ver.3.1�B

�Ÿ
�@Enteromius sensitivus (Roberts 2010)
�@�@�@syn.Barbus sensitivus Roberts 2010

�Ÿ
�@Enteromius serengetiensis (Farm 2000)
�@�@�@syn.Barbus serengetiensis Farm 2000
�@IUCN Vulnerable�@ D2�@ ver.3.1�B

�Ÿ
�@Enteromius sexradiatus (Boulenger 1911)
�@�@�@syn.Barbus sexradiatus Boulenger 1911
�@"ƒJƒrƒ�ƒ“ƒhƒoƒ‹ƒu"�BIUCN Data Deficient ver.3.1�B

�Ÿ
�@Enteromius seymouri (Tweddle & Skelton 2008)
�@�@�@syn.Barbus seymouri Tweddle & Skelton 2008

�Ÿ
�@Enteromius stanleyi (Poll & Gosse 1974)
�@�@�@syn.Barbus gobioides Poll & Gosse 1963 non Bleeker 1852
�@�@�@�@�@Barbus stanleyi Poll & Gosse 1974
�@IUCN Least Concern ver.3.1�BBarbus gobioides Poll & Gosse 1963 ‚̓uƒŒ�[ƒJ�[‚ª�æŽæ‚è�A–³Œø–¼�Ì�B

�Ÿ
�@Enteromius stauchi (Daget 1967)
�@�@�@syn.Barbus stauchi Daget 1967
�@IUCN Endangered�@ B1ab(iii)+2ab(iii)�@ ver.3.1�B

�Ÿ
�@Enteromius stigmasemion (Fowler 1936)
�@�@�@syn.Barbus stigmasemion Fowler 1936

�Ÿ
�@Enteromius stigmatopygus (Boulenger 1903)
�@�@�@syn.Barbus alberti Poll 1939
�@�@�@�@�@Barbus gourmansis Pellégrin 1934
�@�@�@�@�@Barbus karoualensis Blache & Miton 1960
�@�@�@�@�@Barbus stigmatopygus Boulenger 1903
�@�@�@�@�@Barbus werneri Boulenger 1905
�@�@�@�@�@Labeobarbus stigmatopygus (Boulenger 1903)
�@�@�@�@�@Labeobarbus werneri (Boulenger 1905)
�@IUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius subinensis (Hopson 1965)
�@�@�@syn.Barbus subinensis Hopson 1965
�@IUCN Endangered�@ B2ab(iii,v)�@ ver.3.1�B

�Ÿ
�@Enteromius sublineatus (Daget 1954)
�@�@�@syn.Barbus sublineatus Daget 1954
�@‚Q‚Q�`‚Q‚V�Ž�E‚�‚g‚U�D‚W�`‚V�D‚W�BIUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius sylvaticus (Loiselle & Welcomme 1971)
�@�@�@syn.Barbus sylvaticus Loiselle & Welcomme 1971
�@IUCN Endangered�@ B1ab(iii)+2ab(iii)�@ ver.3.1�B

�Ÿ
�@Enteromius syntrechalepis (Fowler 1949)
�@�@�@syn.Barbus syntrechalepis (Fowler 1949)
�@�@�@�@�@Puntius syntrechalepis Fowler 1949
�@IUCN Data Deficient ver.3.1�B

�Ÿ
�@Enteromius taeniopleurus (Boulenger 1917)
�@�@�@syn.Barbus taeniopleurus Boulenger 1917
�@IUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius taeniurus (Boulenger 1903)
�@�@�@syn.Barbus taeniurus Boulenger 1903
�@IUCN Vulnerable�@ B2ab(iii)�@ ver.3.1�B

�Ÿ
�@Enteromius tanapelagius (Graaf, Dejen, Sibbing & Osse 2000)
�@�@�@syn.Barbus tanapelagius Graaf, Dejen, Sibbing & Osse 2000
�@IUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius tegulifer (Fowler 1936)
�@�@�@syn.Barbus tegulifer Fowler 1936
�@IUCN Data Deficient ver.3.1�B

�Ÿ
�@Enteromius tetraspilus (Pfeffer 1896)
�@�@�@syn.Barbus tetraspilus Pfeffer 1896
�@IUCN Data Deficient ver.3.1�B

�Ÿ
�@Enteromius tetrastigmus (Boulenger 1913)
�@�@�@syn.Barbus tetrastigmus Boulenger 1913
�@IUCN Data Deficient ver.3.1�B

�Ÿ
�@Enteromius teugelsi (Bamba, Vreven & Snoeks 2011)
�@�@�@syn.Barbus teugelsi Bamba, Vreven & Snoeks 2011

�Ÿ
�@Enteromius thamalakanensis (Fowler 1935)
�@�@�@syn.Barbus fitzsimonsi Fowler 1935
�@�@�@�@�@Barbus thamalakanensis Fowler 1935
�@"ƒ^ƒ}ƒ‰ƒJƒ“ƒoƒ‹ƒu"�BIUCN Least Concern ver.3.1

�Ÿ
�@Enteromius thespesios Manda, Snoeks, Decru, Bills & Vreven 2019
�@^*2019�wEnteromius thespesios [Teleostei, Cyprinidae], a New Minnow Species with a Remarkable Sexual Dimorphism from the South-Eastern Part of the Upper Congo River�FB.Katemo Manda, J.Snoeks, E.Decru, R.Bills & E.Vreven�x�B

�Ÿ
�@Enteromius thysi (Trewavas 1974)
�@�@�@syn.Barbus thysi Trewavas 1974
�@IUCN Endangered�@ B1ab(iii)+2ab(iii)�@ ver.3.1�B

�Ÿ
�@Enteromius tiekoroi (Lávêque, Teugels & van den Audenaerde 1987)
�@�@�@syn.Barbus tiekoroi Lávêque, Teugels & van den Audenaerde 1987
�@IUCN Data Deficient ver.3.1�B

�Ÿ
�@Enteromius tomiensis (Fowler 1936)
�@�@�@syn.Barbus tomiensis Fowler 1936
�@IUCN Data Deficient ver.3.1�B

�Ÿ
�@Enteromius tongaensis (Rendahl 1935)
�@�@�@syn.Barbus tongaensis Rendahl 1935
�@IUCN Data Deficient ver.3.1�B

�Ÿ
�@Enteromius toppini (Boulenger 1916)
�@�@�@syn.Barbus toppini Boulenger 1916
�@�@�@�@�@Barbus umbeluziensis Groenewald 1958
�@"ƒC�[ƒXƒgƒR�[ƒXƒgƒoƒ‹ƒu"�BIUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius traorei (Lávêque, Teugels & van den Audenaerde 1987)
�@�@�@syn.Barbus traorei Lávêque, Teugels & van den Audenaerde 1987
�@IUCN Endangered�@ B1ab(iii)+2ab(iii)�@ ver.3.1�B

�Ÿ
�@Enteromius treurensis (Groenewald 1958)
�@�@�@syn.Barbus treurensis Groenewald 1958
�@"ƒgƒDƒ‹ƒA�[ƒoƒ‹ƒu"�BIUCN Endangered�@ B1ab(i,ii,iv,v)+2ab(i,ii,iv,v)�@ ver.3.1�B

�Ÿ
�@Enteromius trimaculatus (Peters 1852)
�@�@�@syn.Barbus breijeri Weber 1897
�@�@�@�@�@Barbus decipiens Boulenger 1907
�@�@�@�@�@Barbus katangae Boulenger 1900
�@�@�@�@�@Barbus kurumani de Castelnau 1861
�@�@�@�@�@Barbus serrifer trimaculatus (Peters 1852)
�@�@�@�@�@Barbus trimaculatus Peters 1852
�@"ƒXƒŠ�[ƒXƒ|ƒbƒgƒoƒ‹ƒu"�BIUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius trinotatus (Fowler 1936)
�@�@�@syn.Barbus trinotatus Fowler 1936
�@IUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius trispiloides (Lávêque, Teugels & van den Audenaerde 1987)
�@�@�@syn.Barbus trispiloides Lávêque, Teugels & van den Audenaerde 1987
�@‚Q‚Q�`‚Q‚W�Ž�E‚�‚g‚U�D‚T�`‚W�D‚O�BIUCN Data Deficient ver.3.1�B

�Ÿ
�@Enteromius trispilomimus (Boulenger 1907)
�@�@�@syn.Barbus trispilomimus Boulenger 1907
�@IUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius trispilopleurus (Boulenger 1902)
�@�@�@syn.Barbus trispilopleurus Boulenger 1902
�@IUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius trispilos (Bleeker 1863)
�@�@�@syn.Barbus trispilos (Bleeker 1863)
�@�@�@�@�@Barbus trispilos quinquepunctatus Pellégrin 1911
�@�@�@�@�@�@�@�¦��Enteromius tiekoroi (Lávêque, Teugels & van den Audenaerde 1987)�B
�@�@�@�@�@Puntius trispilos Bleeker 1863
�@IUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius turkanae (Hopson & Hopson 1982)
�@�@�@syn.Barbus turkanae
�@"ƒcƒ‹ƒJƒiƒoƒ‹ƒu"�BIUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius unitaeniatus (Günther 1867)
�@�@�@syn.Barbodes unitaeniatus (Günther 1867)
�@�@�@�@�@Barbus inermoides Nichols & Boulton 1927
�@�@�@�@�@Barbus labialis Gilchrist & Thompson 1913
�@�@�@�@�@Barbus macrurus Gilchrist & Thompson 1913
�@�@�@�@�@Barbus unitaeniatus Günther 1867
�@�@�@�@�@Puntius unitaeniatus (Günther 1867)
�@"ƒXƒŒƒ“ƒ_�[ƒoƒ‹ƒu"�BIUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius urostigmus (Boulenger 1917)
�@�@�@syn.Barbus urostigmus Boulenger 1917
�@IUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius usambarae (Lönnberg 1907)
�@�@�@syn.Barbus usambarae Lönnberg 1907
�@�@�@�@�@Puntius usambarae (Lönnberg 1907)
�@IUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius validus (Stiassny, Liyandja & Monsembula-Iyaba 2016)
�@�@�@syn.Barbus validus Stiassny, Liyandja & Monsembula-Iyaba 2016

�Ÿ
�@Enteromius vanderysti (Poll 1945)
�@�@�@syn.Barbus vanderysti Poll 1945
�@�@�@�@�@Labeobarbus vanderysti (Poll 1945)
�@IUCN Data Deficient ver.3.1�B

�Ÿ
�@Enteromius vandewallei Lederoun & Vreven 2016

�Ÿ
�@Enteromius venustus (Bailey 1980)
�@�@�@syn.Barbus venustus Bailey 1980
�@"ƒpƒ“ƒK“ñƒŒƒbƒhƒoƒ‹ƒu"�BIUCN Data Deficient ver.3.1�B

�Ÿ
�@Enteromius vernayi (Nichols & Boulton 1927)
�@�@�@syn.Barbus vernayi Nichols & Boulton 1927
�@��Enteromius afrovernayi (Nichols & Boulton 1927)�H

�Ÿ
�@Enteromius viktorianus (Lohberger 1929)
�@�@�@syn.Barbus viktorianus Lohberger 1929
�@"ƒrƒNƒgƒŠƒAƒoƒ‹ƒu"�BIUCN Data Deficient ver.3.1�B

�Ÿ
�@Enteromius viviparus (Weber 1897)
�@�@�@syn.Barbus viviparus Weber 1897
�@�@�@�@�@Puntius viviparus (Weber 1897)
�@"ƒ{�[ƒXƒgƒ‰ƒCƒvƒoƒ‹ƒu"�BŽí�¬–¼‚É‘Ù�¶‚Æ‚ ‚邪�AŽÀ�Û‚Í’P‚È‚é—‘ŽY‚Ý‚Á•ú‚µ�BIUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius walkeri (Boulenger 1904)
�@�@�@syn.Barbus walkeri Boulenger 1904
�@IUCN Vulnerable�@ B1ab(iii)+2ab(iii);D2�@ ver.3.1�B

�Ÿ
�@Enteromius walshae Mamonekene, Zamba & Stiassny 2018
�@^*2018�wA New Small Barb [Cyprininae, Smiliogastrini] from the Louesse, Lekoumou, Upper Niari Basin and Djoulou, Upper Ogowe Basin Rivers in the Republic of Congo, West-Central Africa�FVictor Mamonekene, Armel Ibala Zamba & Melanie L.J.Stiassny�x�B

�Ÿ
�@Enteromius wellmani (Boulenger 1911)
�@�@�@syn.Barbus wellmani Boulenger 1911
�@IUCN Data Deficient ver.3.1�B

�Ÿ
�@Enteromius yardiensis Englmaier, Tesfaye & Bogutskaya 2020
�@^*2020�wA New Species of Enteromius [Actinopterygii, Cyprinidae, Smiliogastrinae] from the Awash River, Ethiopia and the Re-Establishment of E.akakianus�FGernot K.Englmaier, Genanaw Tesfaye & Nina G.Bogutskaya�x�B

�Ÿ
�@Enteromius yeiensis (Johnsen 1926)
�@�@�@syn.Barbus yeiensis Johnsen 1926
�@IUCN Data Deficient ver.3.1�B

�Ÿ
�@Enteromius yongei (Whitehead 1960)
�@�@�@syn.Barbus yongei Whitehead 1960
�@"ƒ“ƒ]ƒCƒAƒoƒ‹ƒu"�BIUCN Least Concern ver.3.1�B

�Ÿ
�@Enteromius zalbiensis (Blache & Miton 1960)
�@�@�@syn.Barbus zalbiensis Blache & Miton 1960
�@IUCN Vulnerable�@ B1ab(iii)+2ab(iii);D2�@ ver.3.1�B

�Ÿ
�@Enteromius zanzibaricus (Peters 1868)
�@�@�@syn.? Barbus altus Pfeffer 1896
�@�@�@�@�@Barbus argyrotaenius Boulenger 1912
�@�@�@�@�@Barbus kiperegensis Steindachner 1914
�@�@�@�@�@Barbus pfefferi Boulenger 1905
�@�@�@�@�@Barbus zanzibaricus Peters 1868
�@"ƒUƒ“ƒWƒoƒ‹ƒoƒ‹ƒu"�B‚Q‚Q�`‚Q‚W�Ž�E‚�‚g‚V�D‚Q�`‚V�D‚W�BIUCN Least Concern ver.3.1�B

---

Genus Haludaria^♀ PETHIYAGODA 2013�@ƒnƒ‹ƒ_ƒŠƒA‘®
�@�@�@�¦��Genus Dravidia^♀ PETHIYAGODA, MEEGASKUMBURA & MADUWAGE 2012�B
�@�@�@�¦�‚Genus Dravidia^♀ LEHRER 2010 [Sarcophagidae, Diptera, Insecta]�B

�Ÿ
�@Haludaria afasciata (Jayaram 1990)
�@�@�@syn.Puntius afasciatus Jayaram 1990

�Ÿ
�@Haludaria fasciatus (Jerdon 1849)
�@�@�@syn.Barbus fasciatus (Jerdon 1849)
�@�@�@�@�@Cirrhinus fasciatus Jerdon 1849 non Bleeker 1853
�@�@�@�@�@Dravidia fasciatus (Jerdon 1849) Pethiyagoda, Meegaskumbura & Maduwage 2012
�@�@�@�@�@Puntius fasciatus (Jerdon 1849)
�@�@�@�@�@Puntius fasciatus pradhani
�@^*2012�wA Synopsis of the South Asian Fishes Referred to Puntius [Pisces, Cyprinidae]�FRohan Pethiyagoda, Madhava Meegaskumbura & Kalana Maduwage�x�B
�@"ƒ�ƒ�ƒ“ƒoƒ‹ƒu"�B‚Q‚Q�`‚Q‚U�Ž�E‚�‚g‚U�D‚O�`‚U�D‚T�B


�Ÿ
�@Haludaria kannikattiensis (Arunachalam & Johnson 2003)
�@�@�@syn.Puntius kannikattiensis Arunachalam & Johnson 2003


�Ÿ
�@Haludaria melanampyx (Day 1865)
�@�@�@syn.Barbus melanampyx (Day 1865) Day 1878
�@�@�@�@�@Labeo melanampyx Day 1865
�@�@�@�@�@Puntius melanampyx (Day 1865)
�@^*1878�wA Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters India, Burma and Ceylon 2, Atlas Constaining 198 Plates�FFrancis Day�x�B
�@^*1878�wThe Fishes of India, Being a Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters of India, Burma and Ceylon 1 Extra�FFrancis Day�x�B

�Ÿ
�@Haludaria pradhani (Tilak 1973)
�@�@�@syn.Puntius melanampyx pradhani Tilak 1973

---

Genus Hampala^♀ KUHL & VAN HASSELT ex VAN HASSELT 1823�@ƒnƒ€ƒpƒ‰‘®
�@�@�@�¦�{Genus Hampala^♀ BLEEKER 1860 (Type species : Capoeta macrolepidota Valenciennes 1842)�B
�@�@�@�¦�{Genus Heteroleuciscus^♂ SAUVAGE 1874 (Type species : Heteroleuciscus jullieni Sauvage 1874)�B
�@‚Q‚U�Ž�E‚�‚g‚V�D‚O�B

�Ÿ
�@Hampala ampalong
�@�@�@syn.Barbus ampalong
�@�@�@�@�@Capoeta ampalong

�Ÿ
�@Hampala bimaculata
�@�@�@syn.Barbus hampal bimaculatus


�Ÿ
�@Hampala dispar Smith 1934
�@^*2021�wFishes of the Indo-Chinese Mekong�FNagao Natural Environment Foundation�x�B

�Ÿ
�@Hampala lopezi
�@IUCN Critically Endangered�@ A1a, B1+2a�@ ver.2.3�B


�Ÿ
�@Hampala macrolepidota Kuhl & van Hasselt 1823
�@�@�@syn.Barbus hampal Günther 1868
�@�@�@�@�@Barbus macrolepidotus (Kuhl & van Hasselt 1823) Day 1878
�@�@�@�@�@Capoeta macrolepidota Valenciennes 1842 non Kuhl & van Hasselt 1823
�@�@�@�@�@Heterolenciscus jullieni (Sauvage 1874)�@�¦ƒXƒyƒ‹ƒ~ƒX�B
�@�@�@�@�@Heteroleuciscus jullieni Sauvage 1874
�@^*1850�wCatalogue of Malayan Fishes 1849�FTheodore Cantor�x�B
�@^*1878�wA Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters India, Burma and Ceylon 2, Atlas Constaining 198 Plates�FFrancis Day�x�B
�@^*1878�wThe Fishes of India, Being a Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters of India, Burma and Ceylon 1 Extra�FFrancis Day�x�B
�@^*2021�wFishes of the Indo-Chinese Mekong�FNagao Natural Environment Foundation�x�B
�@"ƒnƒ“ƒpƒ‰ƒoƒ‹ƒu"�B


�Ÿ
�@Hampala sabana
�@�@�@syn.Hampala macrolepidota sabana

�Ÿ
�@Hampala salweenensis

---

Genus Namaquacypris^♀ SKELTON, SWARTZ & VREVEN 2018�@ƒiƒ}ƒNƒAƒLƒvƒŠƒX‘®

�Ÿ
�@Namaquacypris hospes (Barnard 1938)
�@�@�@syn.Barbus hospes Barnard 1938
�@�@�@�@�@Pseudobarbus hospes (Barnard 1938)
�@^*2018�wThe Identity of Barbus capensis Smith 1841 and the Generic Status of Southern African Tetraploid Cyprinids [Teleostei, Cyprinidae]�FPaul H.Skelton, Ernst R.Swartz & Emmanuel J.Vreven�x�B
�@"ƒiƒ}ƒNƒAƒoƒ‹ƒu"�BIUCN Least Concern ver.3.1�B

---

Genus Oliotius^♂ KOTTELAT 2013�@ƒIƒŠƒIƒeƒBƒEƒX‘®

�Ÿ
�@Oliotius oligolepis (Bleeker 1853)
�@�@�@syn.Barbus oligolepis (Bleeker 1853)
�@�@�@�@�@Capoeta oligolepis Bleeker 1853
�@�@�@�@�@Puntius oligolepis (Bleeker 1853)
�@"ƒ`ƒFƒbƒJ�[ƒoƒ‹ƒu"�B‚Q‚O�`‚Q‚S�Ž�E‚�‚g‚U�D‚O�`‚U�D‚T�B

---

Genus Oreichthys^♂ SMITH 1933�@ƒIƒŒƒCƒNƒeƒBƒX‘®
�@ƒIƒXŒÂ‘Ì‚Ì”w•h�L’£�B

�Ÿ
�@Oreichthys andrewi Knight 2014

�Ÿ
�@Oreichthys coorgensis (Jayaram 1982)
�@�@�@syn.Puntius coorgensis Jayaram 1982


�Ÿ
�@Oreichthys cosuatis (Hamilton 1822)
�@�@�@syn.Barbus cosuatis (Hamilton 1822) Day 1876
�@�@�@�@�@Cyprinus cosuatis Hamilton 1822
�@�@�@�@�@Leuciscus cosuatis (Hamilton 1822)
�@�@�@�@�@Puntius cosuatis (Hamilton 1822)
�@�@�@�@�@Rohtee pangut Sykes 1839
�@�@�@�@�@Systomus malacopterus McClelland 1839
�@^*1876�wOn Some of the Fishes of the Deccan�FFrancis Day�x�B
�@^*1878�wA Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters India, Burma and Ceylon 2, Atlas Constaining 198 Plates�FFrancis Day�x�B
�@^*1878�wThe Fishes of India, Being a Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters of India, Burma and Ceylon 1 Extra�FFrancis Day�x�B


�Ÿ
�@Oreichthys crenuchoides
�@�@�@syn.Oreichthys umangii
�@"ƒhƒŒ�[ƒvƒtƒBƒ“ƒoƒ‹ƒu"�B


�Ÿ
�@Oreichthys duospilus Knight & Kumar 2015

�Ÿ
�@Oreichthys incognito Knight & Kumar 2015

�Ÿ
�@Oreichthys parvus Smith 1933
�@^*2021�wFishes of the Indo-Chinese Mekong�FNagao Natural Environment Foundation�x�B

---

Genus Osteobrama^♀ HECKEL 1843�@ƒIƒXƒeƒIƒuƒ‰ƒ}‘®
�@�@�@�¦�{Genus Smiliogaster^♀ BLEEKER 1860 (Type species : Leuciscus belangeri Valenciennes 1844)�B��Smiliogaster�B
�@�@�@�¦�© [Osteobramae RAINBOTH 1991]�B
�@�@�@�¦�© [Osteobramina MENON 1999]�B
�@�@�@�¦�© [Oxygastri BLEEKER 1860]�B�‚Genus Oxygaster^♀ VAN HASSELT 1823�B
�@�@�@�¦�© [Smiliogastrini BLEEKER 1864]�B
�@‚Q‚Q�`‚Q‚T�Ž�B


�Ÿ
�@Osteobrama alfrediana (Valenciennes 1844)
�@�@�@syn.Leuciscus alfredianus Valenciennes 1844
�@�@�@�@�@Rohtee alfrediana (Valenciennes 1844) Day 1876
�@^*1876�wOn Some of the Fishes of the Deccan�FFrancis Day�x�B
�@^*1878�wA Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters India, Burma and Ceylon 2, Atlas Constaining 198 Plates�FFrancis Day�x�B
�@^*1878�wThe Fishes of India, Being a Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters of India, Burma and Ceylon 1 Extra�FFrancis Day�x�B


�Ÿ
�@Osteobrama bakeri (Day 1873)
�@�@�@syn.Rohtee bakeri Day 1873
�@^*1878�wA Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters India, Burma and Ceylon 2, Atlas Constaining 198 Plates�FFrancis Day�x�B
�@^*1878�wThe Fishes of India, Being a Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters of India, Burma and Ceylon 1 Extra�FFrancis Day�x�B


�Ÿ
�@Osteobrama belangeri (Valenciennes 1844)
�@�@�@syn.Leuciscus belangeri Valenciennes 1844
�@�@�@�@�@Osteobrama microlepis (Blyth 1858)
�@�@�@�@�@Rohtee belangeri (Valenciennes 1844) Day 1878
�@�@�@�@�@Rohtee blythi Bleeker 1860
�@�@�@�@�@Rohtee microlepis (Blyth 1858)
�@�@�@�@�@Smiliogaster belangeri (Valenciennes 1844)
�@�@�@�@�@Systomus microlepis Blyth 1858 non Bleeker 1850
�@^*1878�wA Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters India, Burma and Ceylon 2, Atlas Constaining 198 Plates�FFrancis Day�x�B
�@^*1878�wThe Fishes of India, Being a Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters of India, Burma and Ceylon 1 Extra�FFrancis Day�x�B
�@^*1989�wDescription of a New Cyprinid, Osteobrama brevipectoralis sp.nov. from Manipur, India with Remarks on the Systematic Position of the Genus Osteobrama Heckel and Allied Genera�FRaj Tilak & Akhlaq Husain�x�B

�Ÿ
�@Osteobrama bhimensis

�Ÿ
�@Osteobrama brevipectoralis Tilak & Husain 1989
�@��Osteobrama belangeri (Valenciennes 1844)�H


�Ÿ
�@Osteobrama cotio (Hamilton 1822)
�@�@�@syn.Abramis cotio (Hamilton 1822)
�@�@�@�@�@Abramis gangetica (Swainson 1839)
�@�@�@�@�@Cyprinus cotio Hamilton 1822�@�¦��cotis�B
�@�@�@�@�@Leuciscus cotio (Hamilton 1822)
�@�@�@�@�@Leuciscus gangeticus Swainson 1839
�@�@�@�@�@Osteobrama cotio (Hamilton 1822)
�@�@�@�@�@Rohtee cotio (Hamilton 1822) Day 1878
�@^*1878�wA Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters India, Burma and Ceylon 2, Atlas Constaining 198 Plates�FFrancis Day�x�B
�@^*1878�wThe Fishes of India, Being a Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters of India, Burma and Ceylon 1 Extra�FFrancis Day�x�B

�Ÿ
�@Osteobrama cunma (Day 1888)
�@�@�@�@syn.Osteobrama cotio cunmum (Day 1888)
�@�@�@�@�@�@Rohtee cotio cunma (Day 1888)
�@�@�@�@�@�@Rohtee cunma Day 1888
�@�@�@�@�@�@Rohtee roeboides

�Ÿ
�@Osteobrama feae
�@�@�@syn.Rohtee feae


�Ÿ
�@Osteobrama neilli (Day 1873)
�@�@�@syn.Rohtee neilli Day 1873
�@^*1878�wA Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters India, Burma and Ceylon 2, Atlas Constaining 198 Plates�FFrancis Day�x�B
�@^*1878�wThe Fishes of India, Being a Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters of India, Burma and Ceylon 1 Extra�FFrancis Day�x�B

�Ÿ
�@Osteobrama peninsularis (Silas 1952)
�@�@�@�@syn.Osteobrama cotio peninsularis Silas 1952

�Ÿ
�@Osteobrama serrata Singh, Verma, Yumnam & Vishwanath 2017
�@^*2017�wMolecular Phylogenetic Analysis of Genus Osteobrama Heckel 1843 and Discovery of Osteobrama serrata sp.nov. from North East India�FMohinder Singh, Rashmi Verma, Rameshori Yumnam & Waikhom Vishwanath�x�B

�Ÿ
�@Osteobrama tikarpadaensis Shangningam, Rath, Tudu & Kosygin 2020
�@^*2020�wA New Species of Osteobrama [Teleostei, Cyprinidae] from the Mahanadi River, India with a Note on the Validity of O.dayi�FBungdon Shangningam, Shibananda Rath, Asha Kiran Tudu & Laishram Kosygin�x�B
�@^*2021�wFirst Record of Osteobrama tikarpadaensis [Teleostei, Cyprinidae] from Maharashtra India�FShibananda Rath, Asha Kiran Tudu & Bungdon Shangningam�x�B


�Ÿ
�@Osteobrama vigorsii (Sykes 1839)
�@�@�@syn.Abramis vigorsii (Sykes 1839)
�@�@�@�@�@Leuciscus duvaucelii (Valenciennes 1842)
�@�@�@�@�@Osteobrama dayi (Hora & Misra 1940)
�@�@�@�@�@Osteobrama rapax Günther 1868
�@�@�@�@�@Rohita duvaucelii Valenciennes 1842
�@�@�@�@�@Rohtee dayi Hora & Misra 1940
�@�@�@�@�@Rohtee vigorsii Sykes 1839
�@�@�@�@�@Systomus vigorsii (Sykes 1839)
�@^*1876�wOn Some of the Fishes of the Deccan�FFrancis Day�x�B
�@^*1878�wA Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters India, Burma and Ceylon 2, Atlas Constaining 198 Plates�FFrancis Day�x�B
�@^*1878�wThe Fishes of India, Being a Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters of India, Burma and Ceylon 1 Extra�FFrancis Day�x�B
�@‹D�…ˆæ�i“ü�«�BIUCN Least Concern ver.3.1�B

---

Genus Pethia^♀ PETHIYAGODA, MEEGASKUMBURA & MADUWAGE 2012�@ƒyƒeƒBƒA‘®
�@ƒvƒ“ƒeƒBƒEƒX‘®‚æ‚蕪—£“Æ—§�B

�Ÿ
�@Pethia arunachalensis Shangningam, Kosygin & Chowdhury 2020
�@^*2020�wPethia arunachalensis, a New Species of Small Barb [Teleostei, Cyprinidae] from Arunachal Pradesh, India�FBungdon Shangningam, Laishram Kosygin & Basudhara Roy Chowdhury�x�B
�@^*2021�wMolecular Phylogeny and Phylogeography of the Fresh-Water Fish Genus Pethia [Teleostei, Cyprinidae] in Sri Lanka�FHiranya Sudasinghe, Tharindu Ranasinghe, Jayampathi Herath, Kumudu Wijesooriya, Rohan Pethiyagoda, Lukas Ruber & Madhava Meegaskumbura�x�B

�Ÿ
�@Pethia ater (Linthoingambi & Vishwanath 2007)
�@�@�@syn.Puntius ater Linthoingambi & Vishwanath 2007
�@�@�@�@�@Puntius atrus Sudasinghe, Ranasinghe, Herath, Wijesooriya, Pethiyagoda, Rüber & Meegaskumbura 2021�@�¦ƒXƒyƒ‹ƒ~ƒX�B
�@^*2021�wMolecular Phylogeny and Phylogeography of the Fresh-Water Fish Genus Pethia [Teleostei, Cyprinidae] in Sri Lanka�FHiranya Sudasinghe, Tharindu Ranasinghe, Jayampathi Herath, Kumudu Wijesooriya, Rohan Pethiyagoda, Lukas Rüber & Madhava Meegaskumbura�x�B

�Ÿ
�@Pethia aurea Knight 2013
�@^*2013�wPethia aurea [Teleostei, Cyprinidae], a New Species of Barb from West Bengal, India, with Re-Description of P.gelius and P.canius�FJohn Daniel Marcus Knight�x�B
�@^*2021�wMolecular Phylogeny and Phylogeography of the Fresh-Water Fish Genus Pethia [Teleostei, Cyprinidae] in Sri Lanka�FHiranya Sudasinghe, Tharindu Ranasinghe, Jayampathi Herath, Kumudu Wijesooriya, Rohan Pethiyagoda, Lukas Rüber & Madhava Meegaskumbura�x�B

�Ÿ
�@Pethia bandula (Kottelat & Pethiyagoda 1991)
�@�@�@syn.Puntius bandulus Kottelat & Pethiyagoda 1991
�@^*2021�wMolecular Phylogeny and Phylogeography of the Fresh-Water Fish Genus Pethia [Teleostei, Cyprinidae] in Sri Lanka�FHiranya Sudasinghe, Tharindu Ranasinghe, Jayampathi Herath, Kumudu Wijesooriya, Rohan Pethiyagoda, Lukas Rüber & Madhava Meegaskumbura�x�B
�@"ƒoƒ“ƒhƒ‰ƒoƒ‹ƒu"�BIUCN Critically Endangered�@ B1+2c, C1�@ ver.2.3�B

�Ÿ
�@Pethia cania (Hamilton 1822)
�@�@�@syn.Cyprinus canius Hamilton 1822
�@�@�@�@�@Puntius cania (Hamilton 1822)
�@�@�@�@�@Systomus canius (Hamilton 1822)
�@^*2021�wMolecular Phylogeny and Phylogeography of the Fresh-Water Fish Genus Pethia [Teleostei, Cyprinidae] in Sri Lanka�FHiranya Sudasinghe, Tharindu Ranasinghe, Jayampathi Herath, Kumudu Wijesooriya, Rohan Pethiyagoda, Lukas Rüber & Madhava Meegaskumbura�x�B

�Ÿ
�@Pethia castor Conway, Pinion & Kottelat 2021
�@^*2021�wMolecular Phylogeny and Phylogeography of the Fresh-Water Fish Genus Pethia [Teleostei, Cyprinidae] in Sri Lanka�FHiranya Sudasinghe, Tharindu Ranasinghe, Jayampathi Herath, Kumudu Wijesooriya, Rohan Pethiyagoda, Lukas Rüber & Madhava Meegaskumbura�x�B
�@^*2021�wTwo New Species of Pethia [Teleostei, Cyprinidae], Representing a Sympatric Species Pair, from the Ayeyarwady Drainage, Myanmar�FKevin W.Conway, Amanda K.Pinion & Maurice Kottelat�x�B

�Ÿ
�@Pethia chakpiensis Shangningam & Kosygin 2023
�@^*2023�wPethia chakpiensis [Cyprinidae], a New Species of Cyprinid Fish from the Chindwin-Irrawaddy Drainage of Manipur, India�FBungdon Shangningam & Laishram Kosygin�x�B


�Ÿ
�@Pethia conchonia (Hamilton 1822)
�@�@�@syn.Barbus conchonius (Hamilton 1822) Day 1878
�@�@�@�@�@Cyprinus conchonius Hamilton 1822
�@�@�@�@�@Puntius conchonius (Hamilton 1822)
�@�@�@�@�@Puntius conchonius khagariansis Srivastava & Munshi 1988
�@�@�@�@�@Puntius pyropterus (McClelland 1839)
�@�@�@�@�@Systomus conchonius (Hamilton 1822)
�@�@�@�@�@Systomus pyropterus McClelland 1839
�@^*1878�wA Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters India, Burma and Ceylon 2, Atlas Constaining 198 Plates�FFrancis Day�x�B
�@^*1878�wThe Fishes of India, Being a Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters of India, Burma and Ceylon 1 Extra�FFrancis Day�x�B
�@^*2021�wMolecular Phylogeny and Phylogeography of the Fresh-Water Fish Genus Pethia [Teleostei, Cyprinidae] in Sri Lanka�FHiranya Sudasinghe, Tharindu Ranasinghe, Jayampathi Herath, Kumudu Wijesooriya, Rohan Pethiyagoda, Lukas Rüber & Madhava Meegaskumbura�x�B
�@"ƒ��[ƒWƒBƒoƒ‹ƒu"�B‚P‚W�`‚Q‚Q�Ž�E‚�‚g‚U�D‚O�`‚W�D‚O�B


�Ÿ
�@Pethia cumingii (Günther 1868)
�@�@�@syn.Barbus cumingii Günther 1868
�@�@�@�@�@Puntius cumingii (Günther 1868)
�@^*1878�wThe Fishes of India, Being a Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters of India, Burma and Ceylon 1 Extra�FFrancis Day�x�B
�@^*2021�wMolecular Phylogeny and Phylogeography of the Fresh-Water Fish Genus Pethia [Teleostei, Cyprinidae] in Sri Lanka�FHiranya Sudasinghe, Tharindu Ranasinghe, Jayampathi Herath, Kumudu Wijesooriya, Rohan Pethiyagoda, Lukas Rüber & Madhava Meegaskumbura�x�B
�@"ƒgƒD�[ƒXƒ|ƒbƒgƒoƒ‹ƒu"�B‚Q‚Q�`‚Q‚V�Ž�E‚�‚g‚T�D‚T�`‚U�D‚T�BIUCN Lower Risk/conservation dependent ver.2.3�B

�Ÿ
�@Pethia didi (Kullander & Fang 2005)
�@�@�@syn.Puntius didi Kullander & Fang 2005
�@^*2021�wMolecular Phylogeny and Phylogeography of the Fresh-Water Fish Genus Pethia [Teleostei, Cyprinidae] in Sri Lanka�FHiranya Sudasinghe, Tharindu Ranasinghe, Jayampathi Herath, Kumudu Wijesooriya, Rohan Pethiyagoda, Lukas Rüber & Madhava Meegaskumbura�x�B
�@‚Q‚R�Ž�E‚�‚g‚U�D‚T�B

�Ÿ
�@Pethia dikhuensis Praveenraj, Limaakum, Knight, Moulitharan & Imchen 2022
�@^*2022�wPethia dikhuensis (Teleostei: Cyprinidae), a new species from Nagaland, Northeast, India�FJayasimhan Praveenraj, Limaakum Limaakum, John Daniel Marcus Knight, Nallathambi Moulitharan & Nungsangtemjen Imchen�x�B

�Ÿ
�@Pethia erythromycter (Kullander 2008)
�@�@�@syn.Puntius erythromycter Kullander 2008
�@^*2021�wMolecular Phylogeny and Phylogeography of the Fresh-Water Fish Genus Pethia [Teleostei, Cyprinidae] in Sri Lanka�FHiranya Sudasinghe, Tharindu Ranasinghe, Jayampathi Herath, Kumudu Wijesooriya, Rohan Pethiyagoda, Lukas Rüber & Madhava Meegaskumbura�x�B
�@"ƒŒƒbƒhƒŠƒbƒvƒoƒ‹ƒu"�B

�Ÿ
�@Pethia expletiforis Dishma & Vishwanath 2013
�@^*2021�wMolecular Phylogeny and Phylogeography of the Fresh-Water Fish Genus Pethia [Teleostei, Cyprinidae] in Sri Lanka�FHiranya Sudasinghe, Tharindu Ranasinghe, Jayampathi Herath, Kumudu Wijesooriya, Rohan Pethiyagoda, Lukas Rüber & Madhava Meegaskumbura�x�B


�Ÿ
�@Pethia gelia (Hamilton 1822)
�@�@�@syn.Barbus gelius (Hamilton 1822) Day 1878
�@�@�@�@�@Cyprinus gelius Hamilton 1822
�@�@�@�@�@Puntius gelius (Hamilton 1822)
�@�@�@�@�@Systomus gelius (Hamilton 1822)
�@^*1878�wA Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters India, Burma and Ceylon 2, Atlas Constaining 198 Plates�FFrancis Day�x�B
�@^*1878�wThe Fishes of India, Being a Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters of India, Burma and Ceylon 1 Extra�FFrancis Day�x�B
�@^*2021�wMolecular Phylogeny and Phylogeography of the Fresh-Water Fish Genus Pethia [Teleostei, Cyprinidae] in Sri Lanka�FHiranya Sudasinghe, Tharindu Ranasinghe, Jayampathi Herath, Kumudu Wijesooriya, Rohan Pethiyagoda, Lukas Rüber & Madhava Meegaskumbura�x�B
�@"ƒS�[ƒ‹ƒfƒ“ƒoƒ‹ƒu"(ŠCŠO)�B‚P‚W�`‚Q‚Q�Ž�E‚�‚g‚U�D‚O�`‚V�D‚O�B


�Ÿ
�@Pethia guganio (Hamilton 1822)
�@�@�@syn.Barbus carletoni Fowler 1924
�@�@�@�@�@Barbus guganio (Hamilton 1822) Day 1878
�@�@�@�@�@Cyprinus guganio Hamilton 1822
�@�@�@�@�@Leuciscus guganio (Hamilton 1822)
�@�@�@�@�@Puntius carletoni (Fowler 1924)
�@�@�@�@�@Puntius guganio (Hamilton 1822)
�@^*1878�wA Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters India, Burma and Ceylon 2, Atlas Constaining 198 Plates�FFrancis Day�x�B
�@^*1878�wThe Fishes of India, Being a Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters of India, Burma and Ceylon 1 Extra�FFrancis Day�x�B
�@^*2012�wA Synopsis of the South Asian Fishes Referred to Puntius [Pisces, Cyprinidae]�FRohan Pethiyagoda, Madhava Meegaskumbura & Kalana Maduwage�x�B
�@"ƒOƒ‰ƒXƒoƒ‹ƒu"�B

�Ÿ
�@Pethia khugae (Linthoingambi & Vishwanath 2007)
�@�@�@syn.Puntius khugae Linthoingambi & Vishwanath 2007
�@^*2021�wMolecular Phylogeny and Phylogeography of the Fresh-Water Fish Genus Pethia [Teleostei, Cyprinidae] in Sri Lanka�FHiranya Sudasinghe, Tharindu Ranasinghe, Jayampathi Herath, Kumudu Wijesooriya, Rohan Pethiyagoda, Lukas Rüber & Madhava Meegaskumbura�x�B

�Ÿ
�@Pethia longicauda Katwate, Paingankar, Raghavan & Dahanukar 2014
�@^*2014�wPethia longicauda, a New Species of Barb [Teleostei, Cyprinidae] from the Northern Western Ghats, India�FUnmesh Katwate, Mandar S.Paingankar, Rajeev Raghavan & Neelesh Dahanukar�x�B
�@^*2021�wMolecular Phylogeny and Phylogeography of the Fresh-Water Fish Genus Pethia [Teleostei, Cyprinidae] in Sri Lanka�FHiranya Sudasinghe, Tharindu Ranasinghe, Jayampathi Herath, Kumudu Wijesooriya, Rohan Pethiyagoda, Lukas Rüber & Madhava Meegaskumbura�x�B

�Ÿ
�@Pethia lutea Katwate, Katwate, Raghavan, Paingankar & Dahanukar 2014
�@^*2021�wMolecular Phylogeny and Phylogeography of the Fresh-Water Fish Genus Pethia [Teleostei, Cyprinidae] in Sri Lanka�FHiranya Sudasinghe, Tharindu Ranasinghe, Jayampathi Herath, Kumudu Wijesooriya, Rohan Pethiyagoda, Lukas Rüber & Madhava Meegaskumbura�x�B

�Ÿ
�@Pethia macrogramma (Kullander 2008)
�@�@�@syn.Puntius macrogrammus Kullander 2008
�@^*2021�wMolecular Phylogeny and Phylogeography of the Fresh-Water Fish Genus Pethia [Teleostei, Cyprinidae] in Sri Lanka�FHiranya Sudasinghe, Tharindu Ranasinghe, Jayampathi Herath, Kumudu Wijesooriya, Rohan Pethiyagoda, Lukas Rüber & Madhava Meegaskumbura�x�B

�Ÿ
�@Pethia manipurensis Menon, Devi & Vishwanath 2000
�@�@�@syn.Puntius manipurensis (Menon, Devi & Vishwanath 2000)
�@^*2021�wMolecular Phylogeny and Phylogeography of the Fresh-Water Fish Genus Pethia [Teleostei, Cyprinidae] in Sri Lanka�FHiranya Sudasinghe, Tharindu Ranasinghe, Jayampathi Herath, Kumudu Wijesooriya, Rohan Pethiyagoda, Lukas Rüber & Madhava Meegaskumbura�x�B

�Ÿ
�@Pethia meingangbii (Arunkumar & Singh 2003)
�@�@�@syn.Puntius bizonatus Vishwanath & Laisram 2004
�@�@�@�@�@Puntius meingangbii Arunkumar & Singh 2003
�@^*2021�wMolecular Phylogeny and Phylogeography of the Fresh-Water Fish Genus Pethia [Teleostei, Cyprinidae] in Sri Lanka�FHiranya Sudasinghe, Tharindu Ranasinghe, Jayampathi Herath, Kumudu Wijesooriya, Rohan Pethiyagoda, Lukas Rüber & Madhava Meegaskumbura�x�B

�Ÿ
�@Pethia melanomaculata (Deraniyagala 1956)
�@�@�@syn.Puntius melanomaculatus Deraniyagala 1956
�@�@�@�@�@Puntius ticto melanomaculatus Deraniyagala 1956
�@^*2021�wMolecular Phylogeny and Phylogeography of the Fresh-Water Fish Genus Pethia [Teleostei, Cyprinidae] in Sri Lanka�FHiranya Sudasinghe, Tharindu Ranasinghe, Jayampathi Herath, Kumudu Wijesooriya, Rohan Pethiyagoda, Lukas Rüber & Madhava Meegaskumbura�x�B

�Ÿ
�@Pethia muvattupuzhaensis (Beevi & Ramachandran 2005)
�@�@�@syn.Puntius muvattupuzhaensis Beevi & Ramachandran 2005

�Ÿ
�@Pethia nankyweensis (Kullander 2008)
�@�@�@syn.Puntius nankyweensis Kullander 2008
�@^*2021�wMolecular Phylogeny and Phylogeography of the Fresh-Water Fish Genus Pethia [Teleostei, Cyprinidae] in Sri Lanka�FHiranya Sudasinghe, Tharindu Ranasinghe, Jayampathi Herath, Kumudu Wijesooriya, Rohan Pethiyagoda, Lukas Rüber & Madhava Meegaskumbura�x�B

�Ÿ
�@Pethia narayani (Hora 1937)
�@�@�@syn.Barbus narayani Hora 1937
�@�@�@�@�@Puntius narayani (Hora 1937)
�@^*2021�wMolecular Phylogeny and Phylogeography of the Fresh-Water Fish Genus Pethia [Teleostei, Cyprinidae] in Sri Lanka�FHiranya Sudasinghe, Tharindu Ranasinghe, Jayampathi Herath, Kumudu Wijesooriya, Rohan Pethiyagoda, Lukas Rüber & Madhava Meegaskumbura�x�B
�@‚Q‚S�`‚Q‚U�Ž�B

�Ÿ
�@Pethia nigripinnis (Knight, Devi, Indra & Arunachalam 2012) Pethiyagoda, Meegaskumbura & Maduwage 2012
�@�@�@syn.Puntius nigripinnis Knight, Devi, Indra & Arunachalam 2012
�@^*2012�wA Synopsis of the South Asian Fishes Referred to Puntius [Pisces, Cyprinidae]�FRohan Pethiyagoda, Madhava Meegaskumbura & Kalana Maduwage�x�B
�@^*2021�wMolecular Phylogeny and Phylogeography of the Fresh-Water Fish Genus Pethia [Teleostei, Cyprinidae] in Sri Lanka�FHiranya Sudasinghe, Tharindu Ranasinghe, Jayampathi Herath, Kumudu Wijesooriya, Rohan Pethiyagoda, Lukas Rüber & Madhava Meegaskumbura�x�B


�Ÿ
�@Pethia nigrofasciata (Günther 1868)
�@�@�@syn.Barbus nigrofasciatus Günther 1868
�@�@�@�@�@Puntius nigrofasciatus (Günther 1868)
�@^*1878�wA Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters India, Burma and Ceylon 2, Atlas Constaining 198 Plates�FFrancis Day�x�B
�@^*1878�wThe Fishes of India, Being a Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters of India, Burma and Ceylon 1 Extra�FFrancis Day�x�B
�@^*2012�wA Synopsis of the South Asian Fishes Referred to Puntius [Pisces, Cyprinidae]�FRohan Pethiyagoda, Madhava Meegaskumbura & Kalana Maduwage�x�B
�@^*2021�wMolecular Phylogeny and Phylogeography of the Fresh-Water Fish Genus Pethia [Teleostei, Cyprinidae] in Sri Lanka�FHiranya Sudasinghe, Tharindu Ranasinghe, Jayampathi Herath, Kumudu Wijesooriya, Rohan Pethiyagoda, Lukas Rüber & Madhava Meegaskumbura�x�B
�@"ƒuƒ‰ƒbƒNƒ‹ƒr�["�B‚Q‚Q�`‚Q‚U�Ž�E‚�‚g‚U�D‚O�`‚U�D‚T�BIUCN Lower Risk/conservation dependent ver.2.3�B


�Ÿ
�@Pethia ornata
�@�@�@syn.Puntius ornatus

�Ÿ
�@Pethia padamya (Kullander & Britz 2008)
�@�@�@syn.Puntius padamyus (Kullander & Britz 2008)
�@^*2021�wMolecular Phylogeny and Phylogeography of the Fresh-Water Fish Genus Pethia [Teleostei, Cyprinidae] in Sri Lanka�FHiranya Sudasinghe, Tharindu Ranasinghe, Jayampathi Herath, Kumudu Wijesooriya, Rohan Pethiyagoda, Lukas Rüber & Madhava Meegaskumbura�x�B
�@"ƒIƒfƒbƒTƒoƒ‹ƒu"�B



�Ÿ
�@Pethia phutunio (Hamilton 1822)
�@�@�@syn.Barbus phutunio (Hamilton 1822) Day 1878
�@�@�@�@�@Cyprinus phutunio Hamilton 1822 non Hamilton 1822 (�‚puntio)
�@�@�@�@�@Puntius phutunio (Hamilton 1822)
�@�@�@�@�@Systomus leptosomus McClelland 1839
�@�@�@�@�@Systomus phutunio (Hamilton 1822)
�@^*1878�wA Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters India, Burma and Ceylon 2, Atlas Constaining 198 Plates�FFrancis Day�x�B
�@^*1878�wThe Fishes of India, Being a Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters of India, Burma and Ceylon 1 Extra�FFrancis Day�x�B
�@^*2021�wMolecular Phylogeny and Phylogeography of the Fresh-Water Fish Genus Pethia [Teleostei, Cyprinidae] in Sri Lanka�FHiranya Sudasinghe, Tharindu Ranasinghe, Jayampathi Herath, Kumudu Wijesooriya, Rohan Pethiyagoda, Lukas Rüber & Madhava Meegaskumbura�x�B
�@"ƒXƒ|ƒbƒgƒZ�[ƒ‹ƒoƒ‹ƒu"�B‚Q‚Q�`‚Q‚S�Ž�B

�Ÿ
�@Pethia poiensis Shangningam & Vishwanath 2018
�@^*2018�wPethia poiensis, a New Species of Cyprinid Fish from the Chindwin Basin of Manipur, India�FBungdon Shangningam & Waikhom Vishwanath�x�B
�@^*2021�wMolecular Phylogeny and Phylogeography of the Fresh-Water Fish Genus Pethia [Teleostei, Cyprinidae] in Sri Lanka�FHiranya Sudasinghe, Tharindu Ranasinghe, Jayampathi Herath, Kumudu Wijesooriya, Rohan Pethiyagoda, Lukas Rüber & Madhava Meegaskumbura�x�B

�Ÿ
�@Pethia pollux Conway, Pinion & Kottelat 2021
�@^*2021�wMolecular Phylogeny and Phylogeography of the Fresh-Water Fish Genus Pethia [Teleostei, Cyprinidae] in Sri Lanka�FHiranya Sudasinghe, Tharindu Ranasinghe, Jayampathi Herath, Kumudu Wijesooriya, Rohan Pethiyagoda, Lukas Rüber & Madhava Meegaskumbura�x�B
�@^*2021�wTwo New Species of Pethia [Teleostei, Cyprinidae], Representing a Sympatric Species Pair, from the Ayeyarwady Drainage, Myanmar�FKevin W.Conway, Amanda K.Pinion & Maurice Kottelat�x�B

�Ÿ
�@Pethia pookodensis (Mercy & Jacob 2007)
�@�@�@syn.Puntius pookodensis Mercy & Jacob 2007
�@^*2021�wMolecular Phylogeny and Phylogeography of the Fresh-Water Fish Genus Pethia [Teleostei, Cyprinidae] in Sri Lanka�FHiranya Sudasinghe, Tharindu Ranasinghe, Jayampathi Herath, Kumudu Wijesooriya, Rohan Pethiyagoda, Lukas Rüber & Madhava Meegaskumbura�x�B


�Ÿ
�@Pethia punctata (Day 1865)
�@�@�@syn.Barbus punctatus (Day 1865) Day 1878
�@�@�@�@�@Puntius punctatus Day 1865
�@�@�@�@�@Puntius ticto punctatus (Day 1865)
�@^*1878�wA Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters India, Burma and Ceylon 2, Atlas Constaining 198 Plates�FFrancis Day�x�B
�@^*1878�wThe Fishes of India, Being a Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters of India, Burma and Ceylon 1 Extra�FFrancis Day�x�B
�@^*2021�wMolecular Phylogeny and Phylogeography of the Fresh-Water Fish Genus Pethia [Teleostei, Cyprinidae] in Sri Lanka�FHiranya Sudasinghe, Tharindu Ranasinghe, Jayampathi Herath, Kumudu Wijesooriya, Rohan Pethiyagoda, Lukas Rüber & Madhava Meegaskumbura�x�B

�Ÿ
�@Pethia reval (Meegaskumbura, Silva, Maduwage & Pethiyagoda 2008)
�@�@�@syn.Puntius reval Meegaskumbura, Silva, Maduwage & Pethiyagoda 2008
�@^*2021�wMolecular Phylogeny and Phylogeography of the Fresh-Water Fish Genus Pethia [Teleostei, Cyprinidae] in Sri Lanka�FHiranya Sudasinghe, Tharindu Ranasinghe, Jayampathi Herath, Kumudu Wijesooriya, Rohan Pethiyagoda, Lukas Rüber & Madhava Meegaskumbura�x�B

�Ÿ
�@Pethia rutila Lalramliana, Knight & Laltlanhlua 2014
�@^*2014�wPethia rutila [Teleostei, Cyprinidae], a New Species from Mizoram, North-East India�FLalramliana Lalramliana, John Daniel Marcus Knight & Zathang Laltlanhlua�x�B
�@^*2021�wMolecular Phylogeny and Phylogeography of the Fresh-Water Fish Genus Pethia [Teleostei, Cyprinidae] in Sri Lanka�FHiranya Sudasinghe, Tharindu Ranasinghe, Jayampathi Herath, Kumudu Wijesooriya, Rohan Pethiyagoda, Lukas Rüber & Madhava Meegaskumbura�x�B

�Ÿ
�@Pethia sahit Katwate, Kumkar, Raghavan & Dahanukar 2018
�@^*2018�wA New Syntopic Species of Small Barb from the Western Ghats of India [Teleostei, Cyprinidae]�FUnmesh Katwate, Pradeep Kumkar, Rajeev Raghavan & Neelesh Dahanukar�x�B
�@^*2021�wMolecular Phylogeny and Phylogeography of the Fresh-Water Fish Genus Pethia [Teleostei, Cyprinidae] in Sri Lanka�FHiranya Sudasinghe, Tharindu Ranasinghe, Jayampathi Herath, Kumudu Wijesooriya, Rohan Pethiyagoda, Lukas Rüber & Madhava Meegaskumbura�x�B

�Ÿ
�@Pethia sanjaymoluri Katwate, Jadhav, Kumkar, Raghavan & Dahanukar 2016
�@^*2016�wPethia sanjaymoluri, a New Species of Barb [Teleostei, Cyprinidae] from the Northern Western Ghats, India�FUnmesh Katwate, Shrikant Jadhav, Pradeep Kumkar, Rajeev Raghavan & Neelesh Dahanukar�x�B
�@^*2021�wMolecular Phylogeny and Phylogeography of the Fresh-Water Fish Genus Pethia [Teleostei, Cyprinidae] in Sri Lanka�FHiranya Sudasinghe, Tharindu Ranasinghe, Jayampathi Herath, Kumudu Wijesooriya, Rohan Pethiyagoda, Lukas Rüber & Madhava Meegaskumbura�x�B

�Ÿ
�@Pethia setnai (Chhapgar & Sane 1992)
�@�@�@syn.Puntius setnai Chhapgar & Sane 1992
�@^*2021�wMolecular Phylogeny and Phylogeography of the Fresh-Water Fish Genus Pethia [Teleostei, Cyprinidae] in Sri Lanka�FHiranya Sudasinghe, Tharindu Ranasinghe, Jayampathi Herath, Kumudu Wijesooriya, Rohan Pethiyagoda, Lukas Rüber & Madhava Meegaskumbura�x�B


�Ÿ
�@Pethia shalynia (Yazdani & Talukdar 1975)
�@�@�@syn.Puntius shalynius Yazdani & Talukdar 1975
�@^*2021�wMolecular Phylogeny and Phylogeography of the Fresh-Water Fish Genus Pethia [Teleostei, Cyprinidae] in Sri Lanka�FHiranya Sudasinghe, Tharindu Ranasinghe, Jayampathi Herath, Kumudu Wijesooriya, Rohan Pethiyagoda, Lukas Rüber & Madhava Meegaskumbura�x�B

�Ÿ
�@Pethia sharmai (Menon & Devi 1993)
�@�@�@syn.Puntius sharmai Menon & Devi 1993
�@^*2012�wA Synopsis of the South Asian Fishes Referred to Puntius [Pisces, Cyprinidae]�FRohan Pethiyagoda, Madhava Meegaskumbura & Kalana Maduwage�x�B
�@^*2021�wMolecular Phylogeny and Phylogeography of the Fresh-Water Fish Genus Pethia [Teleostei, Cyprinidae] in Sri Lanka�FHiranya Sudasinghe, Tharindu Ranasinghe, Jayampathi Herath, Kumudu Wijesooriya, Rohan Pethiyagoda, Lukas Rüber & Madhava Meegaskumbura�x�B
�@"ƒVƒƒ�[ƒ€ƒoƒ‹ƒu"�B


�Ÿ
�@Pethia stoliczkana (Day 1871)
�@�@�@syn.Barbus stoliczkanus Day 1871
�@�@�@�@�@Barbus ticto stoliczkae�@�¦ƒXƒyƒ‹ƒ~ƒX�B
�@�@�@�@�@Puntius stoliczkanus (Day 1871)
�@�@�@�@�@Puntius ticto stoliczkanus (Day 1871)
�@^*1878�wA Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters India, Burma and Ceylon 2, Atlas Constaining 198 Plates�FFrancis Day�x�B
�@^*1878�wThe Fishes of India, Being a Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters of India, Burma and Ceylon 1 Extra�FFrancis Day�x�B
�@^*2021�wFishes of the Indo-Chinese Mekong�FNagao Natural Environment Foundation�x�B
�@^*2021�wMolecular Phylogeny and Phylogeography of the Fresh-Water Fish Genus Pethia [Teleostei, Cyprinidae] in Sri Lanka�FHiranya Sudasinghe, Tharindu Ranasinghe, Jayampathi Herath, Kumudu Wijesooriya, Rohan Pethiyagoda, Lukas Rüber & Madhava Meegaskumbura�x�B

�Ÿ
�@Pethia striata Atkore, Knight, Rema Devi & Krishnaswamy 2015
�@^*2021�wMolecular Phylogeny and Phylogeography of the Fresh-Water Fish Genus Pethia [Teleostei, Cyprinidae] in Sri Lanka�FHiranya Sudasinghe, Tharindu Ranasinghe, Jayampathi Herath, Kumudu Wijesooriya, Rohan Pethiyagoda, Lukas Rüber & Madhava Meegaskumbura�x�B

�Ÿ
�@Pethia thelys (Kullander 2008)
�@�@�@syn.Puntius thelys Kullander 2008
�@^*2021�wMolecular Phylogeny and Phylogeography of the Fresh-Water Fish Genus Pethia [Teleostei, Cyprinidae] in Sri Lanka�FHiranya Sudasinghe, Tharindu Ranasinghe, Jayampathi Herath, Kumudu Wijesooriya, Rohan Pethiyagoda, Lukas Rüber & Madhava Meegaskumbura�x�B

�Ÿ
�@Pethia tiantian (Kullander & Fang 2005)
�@�@�@syn.Puntius tiantian Kullander & Fang 2005
�@^*2021�wMolecular Phylogeny and Phylogeography of the Fresh-Water Fish Genus Pethia [Teleostei, Cyprinidae] in Sri Lanka�FHiranya Sudasinghe, Tharindu Ranasinghe, Jayampathi Herath, Kumudu Wijesooriya, Rohan Pethiyagoda, Lukas Rüber & Madhava Meegaskumbura�x�B


�Ÿ
�@Pethia ticto (Hamilton 1822)
�@�@�@syn.Barbus ticto (Hamilton 1822) Day 1876
�@�@�@�@�@Cyprinus ticto Hamilton 1822
�@�@�@�@�@Puntius ticto (Hamilton 1822)
�@�@�@�@�@Rothee ticto (Hamilton 1822)
�@�@�@�@�@Systomus ticto (Hamilton 1822)
�@�@�@�@�@Systomus tripunctatus Jerdon 1849
�@^*1876�wOn Some of the Fishes of the Deccan�FFrancis Day�x�B
�@^*1878�wA Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters India, Burma and Ceylon 2, Atlas Constaining 198 Plates�FFrancis Day�x�B
�@^*1878�wThe Fishes of India, Being a Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters of India, Burma and Ceylon 1 Extra�FFrancis Day�x�B
�@^*1928�wFauna of the Chilka Lake 1915-1924 4, Fish 1 1916�FB.L.Chaudhuri�x�B
�@^*2021�wMolecular Phylogeny and Phylogeography of the Fresh-Water Fish Genus Pethia [Teleostei, Cyprinidae] in Sri Lanka�FHiranya Sudasinghe, Tharindu Ranasinghe, Jayampathi Herath, Kumudu Wijesooriya, Rohan Pethiyagoda, Lukas Rüber & Madhava Meegaskumbura�x�B
�@"ƒ`ƒRƒoƒ‹ƒu"�B‚P‚S�`‚Q‚Q�Ž�E‚�‚g‚U�D‚T�`‚V�D‚O�B

�Ÿ
�@Pethia yuensis (Arunkumar & Singh 2002)
�@�@�@syn.Puntius yuensis Arunkumar & Singh 2002
�@^*2021�wMolecular Phylogeny and Phylogeography of the Fresh-Water Fish Genus Pethia [Teleostei, Cyprinidae] in Sri Lanka�FHiranya Sudasinghe, Tharindu Ranasinghe, Jayampathi Herath, Kumudu Wijesooriya, Rohan Pethiyagoda, Lukas Rüber & Madhava Meegaskumbura�x�B

---

Genus Plesiopuntius^♂ SUDASINGHE, RÜBER & MEEGASKUMBURA 2023�@ƒvƒŒƒVƒIƒvƒ“ƒeƒBƒEƒX‘®

�Ÿ
�@Plesiopuntius bimaculatus (Bleeker 1863) Sudasinghe, Rüber & Meegaskumbura 2023
�@�@�@syn.Barbus puckelli (Day 1868) Day 1878
�@�@�@�@�@Gnathopogon bimaculatus Bleeker 1863
�@�@�@�@�@Puntius bimaculatus (Bleeker 1863)
�@�@�@�@�@Puntius puckelli Day 1868�@�¦��Puntius (Capoeta) puckelli Day 1868�B
�@^*1878�wA Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters India, Burma and Ceylon 2, Atlas Constaining 198 Plates�FFrancis Day�x�B
�@^*1878�wThe Fishes of India, Being a Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters of India, Burma and Ceylon 1 Extra�FFrancis Day�x�B
�@^*2023�wMolecular Phylogeny and Systematics of the South Asian Fresh-Water Fish Genus Puntius [Teleostei, Cyprinidae]�FHiranya Sudasinghe, Lukas Rüber & Madhava Meegaskumbura�x�B
�@"ƒŒƒbƒhƒTƒCƒhƒoƒ‹ƒu"�B‚Q‚Q�`‚Q‚S�Ž�E‚�‚g‚U�D‚S�B

Barbus puckelli=Gnathopogon bimaculatus

---

Genus Prolabeops^♂ SCHULTZ 1941�@ƒvƒ�ƒ‰ƒxƒIƒvƒX‘®
�@•ª—ÞŠw“IˆÊ’u‚Í•sˆÀ’è�A‚µ‚΂µ‚ÎBarbinae�EOreininae�B

�Ÿ
�@Prolabeops melanhypopterus (Pellégrin 1928)
�@�@�@syn.Barbus melanhypopterus Pellégrin 1928
�@�@�@�@�@Prolabeops cameroonensis Schultz 1941
�@�@�@�@�@�@�@�¦–ÍŽ®Ží�A’A‚µ�æ‚É‹L�Ú‚³‚ꂽ Barbus melanhypopterus Pellégrin 1928 ‚É“��‡�B
�@^*2022�wTowards the Phylogenetic Placement of the Enigmatic African Genus Prolabeops Schultz 1941�FFrederic Dieter Benedikt Schedel, Zuzana Musilova, Adrian Indermaur, Arnold Roger Bitja-Nyom, Walter Salzburger & Ulrich Kurt Schliewen�x�B
�@IUCN Least Concern ver.3.1�B

�Ÿ
�@Prolabeops nyongensis�@IUCN Data Deficient ver.3.1�B

---

Genus Pseudobarbus^♂ (SMITH 1841)�@ƒvƒZƒEƒhƒoƒ‹ƒuƒX‘®
�@�@�@�¦��Genus Barbus^♂ CUVIER & CLOQUET 1816 Sub-Genus Pseudobarbus^♂ SMITH 1841�B
�@�@�@�¦�{Genus Gnathendalia^♀ DE CASTELNAU 1861 (Type species : Gnatendalia vulnerata de Castelnau 1861)�B
�@�@�@�¦�{Genus Gnatendalia^♀ LÉVÊQUE & DAGET 1984 (Type species : Gnatendalia vulnerata de Castelnau 1861)�BƒXƒyƒ‹ƒ~ƒX�B
�@�@�@�¦�{Genus Oreodaimon^⚲ GREENWOOD & JUBB 1967 (Type species : Labeo quathlambae Barnard 1938)�B
�@�@�@�¦�‚Genus Pseudobarbus^♂ BIELZ 1853 [Genus Barbus]�B

�Ÿ
�@Pseudobarbus afer (Peters 1865)
�@�@�@syn.Barbus afer Peters 1865�@�¦��Barbus (Capoëta) afer Peters 1865�B
�@�@�@�@�@Barbus anoplus Boulenger 1911 non Weber 1897
�@�@�@�@�@Barbus asper Barnard 1943 non Boulenger 1911
�@^*1865�wHr. W.Peters Berichtete über Einige Neue Saugethiere [Mormops, Macrotus, Vesperus, Molossus, Capromys], Amphibien [Platydactylus, Otocryptis, Euprepes, Unglia, Dromicus, Tropidonotus, Xenodon, Hylodes] und Fische [Sillago, Sebastes, Channa, Myctophum, Carassius, Barbus, Capoeta, Poecilia, Saurenchelys, Leptocephalus] 1864�FWilhelm Karl Hartwich Peters�x�B
�@"ƒC�[ƒXƒgƒP�[ƒvƒŒƒbƒhƒtƒBƒ“"�BIUCN Endangered�@ B2ab(ii,iii,v)c(ii,iv)�@ ver.3.1�B

�Ÿ
�@Pseudobarbus asper
�@�@�@syn.Barbus asper
�@"ƒXƒ‚�[ƒ‹ƒXƒP�[ƒ‹ƒŒƒbƒhƒtƒBƒ“"�BIUCN Endangered�@ B2ab(ii,iii,v)�@ ver.3.1�B


�Ÿ
�@Pseudobarbus burchelli
�@�@�@syn.Barbus burchelli Smith 1841�@�¦��Barbus (Pseudobarbus) burchelli Smith 1841�B
�@�@�@�@�@? Barbus gobionides Valenciennes 1842
�@�@�@�@�@Barbus multimaculatus Steindachner 1870
�@�@�@�@�@Barbus vulneratus (de Castelnau 1861)
�@�@�@�@�@Gnathendalia vulnerata de Castelnau 1861
�@�@�@�@�@Gnatendalia vulnerata (de Castelnau 1861)
�@"ƒo�[ƒ`ƒFƒ‹ƒŒƒbƒhƒtƒBƒ“"�BIUCN Critically Endangered�@ B1ab(iii,v)+2ab(iii,v)�@ ver.3.1�B

�Ÿ
�@Pseudobarbus burgi
�@�@�@syn.Barbus burgi
�@"ƒo�[ƒOƒŒƒbƒhƒtƒBƒ“"�BIUCN Endangered�@ B2ab(iii,v)�@ ver.3.1�B

�Ÿ
�@Pseudobarbus erubescens (Skelton 1974)
�@�@�@syn.Barbus erubescens Skelton 1974
�@"ƒgƒEƒB�[ƒŒƒbƒhƒtƒBƒ“"�BIUCN Critically Endangered�@ B2ab(ii,iii,v)�@ ver.3.1�B

�Ÿ
�@Pseudobarbus phlegethon
�@�@�@syn.Barbus phlegethon
�@"ƒtƒBƒGƒŠƒA�[ƒŒƒbƒhƒtƒBƒ“"�BIUCN Endangered�@ B1ab(ii,iii,v)+2ab(ii,iii,v)�@ ver.3.1�B


�Ÿ
�@Pseudobarbus quathlambae (Barnard 1938)
�@�@�@syn.Labeo quathlambae Barnard 1938
�@�@�@�@�@Oreodaimon quathlambae (Barnard 1938)
�@"ƒ}ƒ‹ƒ`ƒ~ƒm�["�BIUCN Endangered�@ B2ab(ii,iii,v)�@ ver.3.1�B

�Ÿ
�@Pseudobarbus senticeps (Smith 1936)
�@�@�@syn.Barbus senticeps Smith 1936

�Ÿ
�@Pseudobarbus serrus (Peters 1865)
�@�@�@syn.Barbus serrus Peters 1865
�@^*1865�wHr. W.Peters Berichtete über Einige Neue Saugethiere [Mormops, Macrotus, Vesperus, Molossus, Capromys], Amphibien [Platydactylus, Otocryptis, Euprepes, Unglia, Dromicus, Tropidonotus, Xenodon, Hylodes] und Fische [Sillago, Sebastes, Channa, Myctophum, Carassius, Barbus, Capoeta, Poecilia, Saurenchelys, Leptocephalus] 1864�FWilhelm Karl Hartwich Peters�x�B
�@"ƒ\�[ƒtƒBƒ“"�BIUCN Endangered�@ B2ab(iii,v)�@ ver.3.1�B

�Ÿ
�@Pseudobarbus skeltoni Chakona & Swartz 2013

�Ÿ
�@Pseudobarbus swartzi Chakona & Skelton 2017

�Ÿ
�@Pseudobarbus tenuis
�@�@�@syn.Barbus tenuis
�@"ƒXƒŒƒ“ƒ_�[ƒŒƒbƒhƒtƒBƒ“"�BIUCN Near Threatened ver.3.1�B

�Ÿ
�@Pseudobarbus verloreni Chakona, Swartz & Skelton 2014

---

Genus Puntigrus^♂ KOTTELAT 2013�@ƒvƒ“ƒeƒBƒOƒ‹ƒX‘®

�Ÿ
�@Puntigrus anchisporus (Vaillant 1902)
�@�@�@syn.Barbus anchisporus Vaillant 1902
�@�@�@�@�@Puntius anchisporus (Vaillant 1902)
�@�@�@�@�@Systomus anchisporus (Vaillant 1902)

�Ÿ
�@Puntigrus navjotsodhii (Tan 2012)
�@�@�@syn.Systomus navjotsodhii Tan 2012

�Ÿ
�@Puntigrus partipentazonus (Fowler 1934)
�@�@�@syn.Barbus partipentazonus Fowler 1934
�@�@�@�@�@Barbus tetrazonus partipentazonus (Fowler 1934)
�@�@�@�@�@Puntius partipentazonus (Fowler 1934)
�@�@�@�@�@Systomus partipentazonus (Fowler 1934)
�@^*2021�wFishes of the Indo-Chinese Mekong�FNagao Natural Environment Foundation�x�B
�@‚Q‚Q�`‚Q‚T�Ž�E‚V�D‚O�B

�Ÿ
�@Puntigrus pulcher (Rendahl 1922)
�@�@�@syn.Puntius pulcher Rendahl 1922
�@�@�@�@�@Systomus pulcher (Rendahl 1922)


�Ÿ
�@Puntigrus tetrazonus (Bleeker 1855)
�@�@�@syn.Barbus tetrazonus (Bleeker 1855)
�@�@�@�@�@Capoeta tetrazona Bleeker 1855 non Bleeker 1856
�@�@�@�@�@Puntius tetrazonus (Bleeker 1855)
�@�@�@�@�@Systomus tetrazonus (Bleeker 1855)
�@"ƒXƒ}ƒgƒ‰(ƒAƒ‹ƒrƒm�EƒOƒŠ�[ƒ“)"�B‚Q‚O�`‚Q‚U�Ž�E‚�‚g‚U�D‚O�`‚W�D‚O�B

---

Genus Puntius^♂ (HAMILTON 1822)�@ƒvƒ“ƒeƒBƒEƒX‘®
�@�@�@�¦��Genus Cyprinus^♂ VON LINNÉ 1758 Sub-Genus Puntius^♂ HAMILTON 1822�B
�@�@�@�¦�© [Puntiini KARAMAN 1971]�B
�@�«�·–¾—Ä‚ÌŒXŒü�B(‚P‚T�`�j‚Q‚T�`‚Q‚U(�`‚R‚O�j�Ž�E‚�‚g(‚U�D‚O�`�j‚U�D‚W�`‚V�D‚Q(�`‚V�D‚T�j�B


�Ÿ
�@Puntius ambassis (Day 1869)
�@�@�@syn.Barbus ambassis Day 1869
�@^*1878�wA Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters India, Burma and Ceylon 2, Atlas Constaining 198 Plates�FFrancis Day�x�B
�@^*1878�wThe Fishes of India, Being a Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters of India, Burma and Ceylon 1 Extra�FFrancis Day�x�B


�Ÿ
�@Puntius amphibius (Valenciennes 1842)
�@�@�@syn.Barbus amphibius (Valenciennes 1842) Day 1878
�@�@�@�@�@Capoeta amphibia Valenciennes 1842
�@�@�@�@�@Systomus amphibius (Valenciennes 1842)
�@^*1878�wA Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters India, Burma and Ceylon 2, Atlas Constaining 198 Plates�FFrancis Day�x�B
�@^*1878�wThe Fishes of India, Being a Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters of India, Burma and Ceylon 1 Extra�FFrancis Day�x�B
�@"ƒXƒJ�[ƒŒƒbƒgƒoƒ“ƒhƒoƒ‹ƒu"�B‹D�…ˆæ�i“ü�«�B

�Ÿ
�@Puntius aphyus
�@�@�@syn.Barbus aphyus


�Ÿ
�@Puntius arenatus (Day 1878)
�@�@�@syn.Barbus arenatus Day 1878
�@^*1878�wA Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters India, Burma and Ceylon 2, Atlas Constaining 198 Plates�FFrancis Day�x�B
�@^*1878�wThe Fishes of India, Being a Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters of India, Burma and Ceylon 1 Extra�FFrancis Day�x�B

�Ÿ
�@Puntius bramoides
�@�@�@syn.Barbus bramoides


�Ÿ
�@Puntius brevis (Bleeker 1849)
�@�@�@syn.Barbus brevis (Bleeker 1849)
�@�@�@�@�@Capoeta brevis Bleeker 1849
�@�@�@�@�@Puntius leiacanthus (Bleeker 1860)
�@�@�@�@�@Puntius ocellatus Mai 1978
�@�@�@�@�@Systomus brevis (Bleeker 1849)
�@�@�@�@�@Systomus leiacanthus Bleeker 1860
�@^*2021�wFishes of the Indo-Chinese Mekong�FNagao Natural Environment Foundation�x�B


�Ÿ
�@Puntius burmanicus (Day 1878)
�@�@�@syn.Barbus burmanicus Day 1878
�@^*1878�wA Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters India, Burma and Ceylon 2, Atlas Constaining 198 Plates�FFrancis Day�x�B
�@^*1878�wThe Fishes of India, Being a Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters of India, Burma and Ceylon 1 Extra�FFrancis Day�x�B
�@"ƒrƒ‹ƒ}ƒoƒ‹ƒu"�B‚Q‚S�`‚Q‚U�Ž�B

�Ÿ
�@Puntius cauveriensis (Hora 1937)
�@�@�@syn.Barbus cauveriensis Hora 1937


�Ÿ
�@Puntius cholus (Hamilton 1822)
�@�@�@syn.Barbus cholus (Hamilton 1822) Day 1878
�@�@�@�@�@Barbus tetrarupagus (McClelland 1839) Day 1878
�@�@�@�@�@Barbus titius (Hamilton 1822)
�@�@�@�@�@Barbus unimaculatus (Blyth 1860) Day 1878
�@�@�@�@�@Capoeta chola (Hamilton 1822)
�@�@�@�@�@Cyprinus cholus Hamilton 1822
�@�@�@�@�@Cyprinus titius Hamilton 1822
�@�@�@�@�@Puntius tetrarupagus (McClelland 1839)
�@�@�@�@�@Puntius titius (Hamilton 1822)
�@�@�@�@�@Systomus tetrarupagus McClelland 1839
�@�@�@�@�@Systomus unimaculatus Blyth 1860
�@^*1878�wA Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters India, Burma and Ceylon 2, Atlas Constaining 198 Plates�FFrancis Day�x�B
�@^*1878�wThe Fishes of India, Being a Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters of India, Burma and Ceylon 1 Extra�FFrancis Day�x�B
�@"ƒXƒ�ƒ“ƒvƒoƒ‹ƒu"�B‚Q‚O�`‚Q‚T�Ž�E‚�‚g‚U�D‚O�`‚U�D‚T�B

Barbus unimaculatus=Cyprinus cholus

Barbus tetrarupagus=Cyprinus cholus

�Ÿ
�@Puntius crescentus

�Ÿ
�@Puntius deccanensis Yazdani & Rao 1976
�@^*2012�wA Synopsis of the South Asian Fishes Referred to Puntius [Pisces, Cyprinidae]�FRohan Pethiyagoda, Madhava Meegaskumbura & Kalana Maduwage�x�B
�@"ƒfƒbƒJƒoƒ‹ƒu"�B

�Ÿ
�@Puntius dolichopterus Plamoottil 2015


�Ÿ
�@Puntius dorsalis (Jerdon 1849)
�@�@�@syn.Barbus dorsalis (Jerdon 1849) Day 1878
�@�@�@�@�@Labeo dorsalis (Jerdon 1849) Jordan & Starks 1917
�@�@�@�@�@Systomus dorsalis Jerdon 1849
�@�@�@�@�@Systomus tristis Jerdon 1849
�@^*1878�wA Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters India, Burma and Ceylon 2, Atlas Constaining 198 Plates�FFrancis Day�x�B
�@^*1878�wThe Fishes of India, Being a Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters of India, Burma and Ceylon 1 Extra�FFrancis Day�x�B
�@^*1917�wNotes on a Collection of Fishes from Ceylon with Descriptions of New Species�FDavid Starr Jordan & Edwin Chapin Starks�x�B
�@"ƒ�ƒ“ƒOƒXƒiƒEƒgƒoƒ‹ƒu"�B

�Ÿ
�@Puntius euspilurus Plamoottil 2016
�@^*2016�wPuntius euspilurus, a New Fish Species [Cypriniformes, Cyprinidae] from Kerala, India�FMathews Plamoottil�x�B

�Ÿ
�@Puntius fraseri (Hora & Misra 1938)
�@�@�@syn.Barbus fraseri Hora & Misra 1938
�@^*2012�wA Synopsis of the South Asian Fishes Referred to Puntius [Pisces, Cyprinidae]�FRohan Pethiyagoda, Madhava Meegaskumbura & Kalana Maduwage�x�B
�@"ƒ_�[ƒ‹ƒiƒoƒ‹ƒu"�B


�Ÿ
�@Puntius kamalikus

�Ÿ
�@Puntius kelumi

�Ÿ
�@Puntius khohi

�Ÿ
�@Puntius kyphus Plamoottil 2019
�@^*2016�wPuntius kyphus [Cypriniformes, Cyprinidae], a New Fish Species from Kerala, India�FMathews Plamoottil�x�B

�Ÿ
�@Puntius layardi
�@�@�@syn.Barbus layardi

�Ÿ
�@Puntius madhusoodani Krishnakumar, Benno Pereira & Radhakrishnan 2012


�Ÿ
�@Puntius mahecolus (Valenciennes 1844)
�@�@�@syn.Barbus mahicolus Day 1878�@�¦ƒXƒyƒ‹ƒ~ƒX�B
�@�@�@�@�@Leuciscus mahecolus Valenciennes 1844
�@^*1878�wA Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters India, Burma and Ceylon 2, Atlas Constaining 198 Plates�FFrancis Day�x�B
�@^*1878�wThe Fishes of India, Being a Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters of India, Burma and Ceylon 1 Extra�FFrancis Day�x�B

�Ÿ
�@Puntius masyai Smith 1945
�@^*2021�wFishes of the Indo-Chinese Mekong�FNagao Natural Environment Foundation�x�B


�Ÿ
�@Puntius melanostigmus (Day 1878)
�@�@�@syn.Barbus melanostigmus Day 1878
�@^*1878�wA Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters India, Burma and Ceylon 2, Atlas Constaining 198 Plates�FFrancis Day�x�B
�@^*1878�wThe Fishes of India, Being a Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters of India, Burma and Ceylon 1 Extra�FFrancis Day�x�B
�@"ƒ�ƒCƒi�[ƒhƒoƒ‹ƒu"�B

�Ÿ
�@Puntius morehensis

�Ÿ
�@Puntius mudumalaiensis

�Ÿ
�@Puntius muzaffarpurensis Srivastava, Verma & Sharma 1977

�Ÿ
�@Puntius nangalensis Jayaram 1990
�@^*2012�wA Synopsis of the South Asian Fishes Referred to Puntius [Pisces, Cyprinidae]�FRohan Pethiyagoda, Madhava Meegaskumbura & Kalana Maduwage�x�B

�Ÿ
�@Puntius nelsoni Plamoottil 2014

�Ÿ
�@Puntius nigronotus Plamoottil 2015

�Ÿ
�@Puntius ocellus Plamoottil & Vineeth 2020
�@^*2020�wDiscovery of a New Teleost Fish, Puntius ocellus from Kerala, India�FMathews Plamoottil & Kumar Vineeth�x�B


�Ÿ
�@Puntius parrah Day 1865
�@�@�@syn.Barbus parrah (Day 1865) Day 1878
�@�@�@�@�@Puntius perlee Day 1865
�@^*1878�wA Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters India, Burma and Ceylon 2, Atlas Constaining 198 Plates�FFrancis Day�x�B
�@^*1878�wThe Fishes of India, Being a Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters of India, Burma and Ceylon 1 Extra�FFrancis Day�x�B

�Ÿ
�@Puntius paucimaculatus


�Ÿ
�@Puntius pinnauratus (Day 1865)
�@�@�@syn.Barbus pinnauratus (Day 1865) Day 1878
�@�@�@�@�@Cyclocheilichthys pinnauratus Day 1865
�@^*1878�wA Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters India, Burma and Ceylon 2, Atlas Constaining 198 Plates�FFrancis Day�x�B
�@^*1878�wThe Fishes of India, Being a Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters of India, Burma and Ceylon 1 Extra�FFrancis Day�x�B
�@^*1917�wNotes on a Collection of Fishes from Ceylon with Descriptions of New Species�FDavid Starr Jordan & Edwin Chapin Starks�x�B

�Ÿ
�@Puntius pugio 2008


�Ÿ
�@Puntius punjaubensis (Day 1871)
�@�@�@syn.Barbus punjabensis�@�¦ƒXƒyƒ‹ƒ~ƒX�B
�@�@�@�@�@Barbus punjaubensis Day 1871
�@^*1878�wA Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters India, Burma and Ceylon 2, Atlas Constaining 198 Plates�FFrancis Day�x�B
�@^*1878�wThe Fishes of India, Being a Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters of India, Burma and Ceylon 1 Extra�FFrancis Day�x�B
�@^*2012�wA Synopsis of the South Asian Fishes Referred to Puntius [Pisces, Cyprinidae]�FRohan Pethiyagoda, Madhava Meegaskumbura & Kalana Maduwage�x�B


�Ÿ
�@Puntius puntio (Hamilton 1822)
�@�@�@syn.Barbus puntio (Hamilton 1822) Day 1878
�@�@�@�@�@Cyprinus puntio Hamilton 1822 non Hamilton 1822 (�‚phutunio)
�@^*1878�wA Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters India, Burma and Ceylon 2, Atlas Constaining 198 Plates�FFrancis Day�x�B
�@^*1878�wThe Fishes of India, Being a Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters of India, Burma and Ceylon 1 Extra�FFrancis Day�x�B
�@"ƒvƒ“ƒeƒBƒIƒoƒ‹ƒu"�B

�Ÿ
�@Puntius sachsii
�@�@�@syn.Barbus sachsii
�@"ƒS�[ƒ‹ƒfƒ“ƒoƒ‹ƒu"�E"ƒS�[ƒ‹ƒhƒtƒBƒ“ƒoƒ‹ƒu"�B

�Ÿ
�@Puntius sanctus Plamoottil 2020
�@^*2020�wPuntius sanctus, a New Fish [Cypriniformes, Cyprinidae] Species from Tamil Nadu, India�FMathews Plamoottil�x�B

�Ÿ
�@Puntius sealei
�@�@�@syn.Barbodes sealei
�@�@�@�@�@? Barbus elongatus
�@�@�@�@�@Barbus sealei
�@�@�@�@�@Puntius elongatus


�Ÿ
�@Puntius sophore (Hamilton 1822)
�@�@�@syn.Barbus annandalei Fowler 1924
�@�@�@�@�@Barbus chrysopterus (McClelland 1839) Day 1878
�@�@�@�@�@Barbus duvaucelii (Valenciennes 1844)
�@�@�@�@�@Barbus sophore (Hamilton 1822) Day 1878
�@�@�@�@�@Cyprinus sophore Hamilton 1822
�@�@�@�@�@Leuciscus duvaucelii Valenciennes 1844
�@�@�@�@�@Puntius chrysopterus (McClelland 1839)
�@�@�@�@�@Puntius modestus Kner 1867
�@�@�@�@�@Systomus chrysopterus McClelland 1839
�@�@�@�@�@Systomus duvaucelii (Valenciennes 1844)
�@�@�@�@�@Systomus sophore (Hamilton 1822)
�@^*1878�wA Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters India, Burma and Ceylon 2, Atlas Constaining 198 Plates�FFrancis Day�x�B
�@^*1878�wThe Fishes of India, Being a Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters of India, Burma and Ceylon 1 Extra�FFrancis Day�x�B
�@^*1928�wFauna of the Chilka Lake 1915-1924 4, Fish 1 1916�FB.L.Chaudhuri�x�B
�@^*2012�wA Synopsis of the South Asian Fishes Referred to Puntius [Pisces, Cyprinidae]�FRohan Pethiyagoda, Madhava Meegaskumbura & Kalana Maduwage�x�B
�@"ƒv�[ƒ‹ƒoƒ‹ƒu"�B‚Q‚Q�Ž�E‚�‚g‚W�D‚O�B

Barbus chrysopterus=Cyprinus sophore

�Ÿ
�@Puntius sophoroides
�@�@�@syn.Barbus sophoroides

�Ÿ
�@Puntius spilopterus
�@�@�@syn.Barbus spilopterus


�Ÿ
�@Puntius stigmus (Valenciennes 1844)
�@�@�@syn.Barbus stigmus (Valenciennes 1844) Day 1876
�@�@�@�@�@Leuciscus stigmus Valenciennes 1844
�@^*1876�wOn Some of the Fishes of the Deccan�FFrancis Day�x�B
�@^*1878�wA Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters India, Burma and Ceylon 2, Atlas Constaining 198 Plates�FFrancis Day�x�B
�@^*1878�wThe Fishes of India, Being a Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters of India, Burma and Ceylon 1 Extra�FFrancis Day�x�B

�Ÿ
�@Puntius takhoaensis


�Ÿ
�@Puntius terio (Hamilton 1822)
�@�@�@syn.Barbus terio (Hamilton 1822) Day 1878
�@�@�@�@�@Cyprinus terio Hamilton 1822
�@�@�@�@�@Systomus gibbosus McClelland 1839
�@�@�@�@�@Systomus terio (Hamilton 1822)
�@^*1878�wA Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters India, Burma and Ceylon 2, Atlas Constaining 198 Plates�FFrancis Day�x�B
�@^*1878�wThe Fishes of India, Being a Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters of India, Burma and Ceylon 1 Extra�FFrancis Day�x�B
�@"ƒ�ƒ“ƒXƒ|ƒbƒgƒoƒ‹ƒu"�B‚P‚W�`‚Q‚W�Ž�E‚�‚g‚V�D‚O�`‚W�D‚O�B

�Ÿ
�@Puntius tetraspilus (Günther 1868)
�@�@�@syn.Barbus innominatus Day 1870
�@�@�@�@�@Barbus tetraspilus Günther 1868
�@�@�@�@�@Leuciscus binotatus Blyth 1858
�@�@�@�@�@Neolissochilus innominatus (Day 1870)
�@^*1878�wThe Fishes of India, Being a Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters of India, Burma and Ceylon 1 Extra�FFrancis Day�x�B

�Ÿ
�@Puntius thermalis (Valenciennes 1844)
�@�@�@syn.Barbus thermalis (Valenciennes 1844) Day 1878
�@�@�@�@�@Leuciscus thermalis Valenciennes 1844
�@^*1878�wThe Fishes of India, Being a Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters of India, Burma and Ceylon 1 Extra�FFrancis Day�x�B

�Ÿ
�@Puntius viridis Plamoottil & Abraham 2014


�Ÿ
�@Puntius waageni (Day 1872)
�@�@�@syn.Barbus waageni Day 1872
�@^*1878�wA Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters India, Burma and Ceylon 2, Atlas Constaining 198 Plates�FFrancis Day�x�B
�@^*1878�wThe Fishes of India, Being a Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters of India, Burma and Ceylon 1 Extra�FFrancis Day�x�B

---

Genus Rohanella^♀ SUDASINGHE, RÜBER & MEEGASKUMBURA 2023�@ƒ�ƒnƒlƒ‹ƒ‰‘®


�Ÿ
�@Rohanella titteya (Deraniyagala 1929) Sudasinghe, Rüber & Meegaskumbura 2023
�@�@�@syn.Barbus titteyus (Deraniyagala 1929)
�@�@�@�@�@Capoeta titteya (Deraniyagala 1929)
�@�@�@�@�@Puntius titteyus Deraniyagala 1929
�@�@�@�@�@Puntius titteyus var.rubripinnis Deraniyagala 1958
�@^*2023�wMolecular Phylogeny and Systematics of the South Asian Fresh-Water Fish Genus Puntius [Teleostei, Cyprinidae]�FHiranya Sudasinghe, Lukas Rüber & Madhava Meegaskumbura�x�B
�@"ƒ`ƒFƒŠ�[ƒoƒ‹ƒu"�B‚Q‚R�`‚Q‚V�Ž�E‚�‚g‚U�D‚O�`‚W�D‚O�BIUCN Lower Risk/conservation dependent ver.2.3�B

---

Genus Rohtee^♂ SYKES 1839�@ƒ��[ƒeƒG‘®
�@‚Q‚U�Ž�E‚�‚g‚U�D‚W�B


�Ÿ
�@Rohtee ogilbii Sykes 1839
�@�@�@syn.Abramis orgibii (Sykes 1839)
�@�@�@�@�@Mystacoleucus ogilbii (Sykes 1839)
�@�@�@�@�@Osteobrama ogilbii (Sykes 1839)
�@�@�@�@�@Systomus ogilbii (Sykes 1839)
�@^*1876�wOn Some of the Fishes of the Deccan�FFrancis Day�x�B
�@^*1878�wA Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters India, Burma and Ceylon 2, Atlas Constaining 198 Plates�FFrancis Day�x�B
�@^*1878�wThe Fishes of India, Being a Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters of India, Burma and Ceylon 1 Extra�FFrancis Day�x�B
�@"ƒoƒ^ƒjƒ��[ƒe�["�B

---

Genus Rohteichthys^♂ BLEEKER 1860�@ƒ��[ƒeƒCƒNƒeƒBƒX‘®
�@�@�@�¦�© [Rohteichthyina GÜNTHER 1868]�B
�@•ª—ÞŠw“IˆÊ’u‚Í•sˆÀ’è�B

�Ÿ
�@Rohteichthys microlepis (Bleeker 1851)
�@�@�@syn.Barbus microlepis Bleeker 1851 non Keyserling 1861
�@�@�@�@�@Rohtee microlepis (Bleeker 1851)
�@�@�@�@�@Systomus microlepis (Bleeker 1851)

---

Genus Sahyadria^♀ RAGHAVAN, PHILIP, ALI & DAHANUKAR 2013�@ƒTƒqƒAƒhƒŠƒA‘®

�Ÿ
�@Sahyadria chalakkudiensis (Menon, Rema Devi & Thobias 1999)
�@�@�@syn.Puntius chalakkudiensis Menon, Rema Devi & Thobias 1999


�Ÿ
�@Sahyadria denisonii (Day 1865)
�@�@�@syn.Barbus denisonii (Day 1865) Day 1878
�@�@�@�@�@Crossocheilus denisonii (Day 1865)
�@�@�@�@�@Labeo denisonii Day 1865
�@�@�@�@�@Puntius denisonii (Day 1865)
�@^*1878�wA Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters India, Burma and Ceylon 2, Atlas Constaining 198 Plates�FFrancis Day�x�B
�@^*1878�wThe Fishes of India, Being a Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters of India, Burma and Ceylon 1 Extra�FFrancis Day�x�B
�@"ƒfƒjƒ\ƒ“ƒoƒ‹ƒu"�E"ƒg�[ƒs�[ƒhƒoƒ‹ƒu(ƒŒƒbƒhƒ‰ƒCƒ“�j"�B‚P‚T�`‚Q‚T�Ž�E‚�‚g‚U�D‚W�`‚V�D‚W�BIUCN Vulnerable�@ B1ab(iii,v)�@ ver.3.1�B

---

Genus Sedercypris^♀ SKELTON, SWARTZ & VREVEN 2018�@ƒZƒfƒ‹ƒLƒvƒŠƒX‘®

�Ÿ
�@Sedercypris calida (Barnard 1938)
�@�@�@syn.Barbus calidus Barnard 1938
�@�@�@�@�@Pseudobarbus calidus (Barnard 1938)�@�¦ƒtƒBƒbƒVƒ…ƒx�[ƒX‚Ì—LŒø–¼�Ì�B
�@^*2018�wThe Identity of Barbus capensis Smith 1841 and the Generic Status of Southern African Tetraploid Cyprinids [Teleostei, Cyprinidae]�FPaul H.Skelton, Ernst R.Swartz & Emmanuel J.Vreven�x�B
�@"ƒNƒ‰�[ƒ“ƒEƒBƒŠƒAƒ€ƒŒƒbƒhƒtƒBƒ“"�BIUCN Vulnerable�@ B2ab(ii,iii,v)�@ ver.3.1�B

---

Genus Striuntius^♂ KOTTELAT 2013�@ƒXƒgƒŠƒEƒ“ƒeƒBƒEƒX‘®

�Ÿ
�@Striuntius lineatus (Duncker 1904)
�@�@�@syn.Barbus lineatus Duncker 1904 non Popta 1905
�@�@�@�@�@Puntius lineatus (Duncker 1904)
�@"ƒ‰ƒCƒ“ƒoƒ‹ƒu"�B

---

Genus Systomus^♂ MCCLELLAND 1839�@ƒVƒXƒgƒ€ƒX‘®
�@�@�@�¦�© [Sub-Family Sarcoborinae MCCLELLAND 1838]�B
�@�@�@�¦�© [Systomi BLEEKER 1863-1864]�B
�@�@�@�¦�© [Systomini MENON 1999]�B
�@�@�@�¦�© [Systomini RAINBOTH 1991]�B

�Ÿ
�@Systomus asokus
�@�@�@syn.Barbichthys asokae
�@�@�@�@�@Barbus asokus
�@�@�@�@�@Puntius asokus
�@"ƒAƒ\ƒJƒoƒ‹ƒu"�B‚Q‚T�`‚R‚O�Ž�E‚�‚g‚U�D‚T�`‚V�D‚T�BIUCN Endangered�@ A1c, B1+2c�@ ver.2.3�B

�Ÿ
�@Systomus chryseus Plamoottil 2014

�Ÿ
�@Systomus clavatus (McClelland 1845)
�@�@�@syn.Barbus clavatus McClelland 1845
�@�@�@�@�@Poropuntius clavatus (McClelland 1845)
�@�@�@�@�@Puntius clavatus (McClelland 1845)
�@^*1878�wThe Fishes of India, Being a Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters of India, Burma and Ceylon 1 Extra�FFrancis Day�x�B
�@"ƒXƒeƒbƒhƒ}ƒ“ƒoƒ‹ƒu"�B

�Ÿ
�@Systomus compressiformis
�@�@�@syn.Barbus compressiformis
�@�@�@�@�@Puntius compressiformis

�Ÿ
�@Systomus endecanalis (Roberts 1989)
�@�@�@syn.Desmopuntius endecanalis (Roberts 1989)
�@�@�@�@�@Puntius endecanalis Roberts 1989

�Ÿ
�@Systomus gracilus Plamoottil & Maji 2020
�@^*2020�wSystomus gracilus, A New Fish [Cypriniformes, Cyprinidae] Species from West Bengal�FMathews Plamoottil & Debarghya Maji�x�B

�Ÿ
�@Systomus immaculatus McClelland 1839
�@^*2012�wA Synopsis of the South Asian Fishes Referred to Puntius [Pisces, Cyprinidae]�FRohan Pethiyagoda, Madhava Meegaskumbura & Kalana Maduwage�x�B
�@"ƒIƒŠ�[ƒuƒoƒ‹ƒu"�B‹D�…ˆæ�i“ü�«�B

�Ÿ
�@Systomus jayarami
�@�@�@syn.Puntius jayarami

�Ÿ
�@Systomus laticeps Plamoottil 2016
�@^*2016�wSystomus laticeps, a New Species of Cyprinid Fish from Kerala, India�FMathews Plamoottil�x�B

�Ÿ
�@Systomus martenstyni
�@�@�@syn.Puntius martenstyni
�@"ƒ}ƒ‹ƒeƒ“ƒXƒeƒBƒoƒ‹ƒu"�BIUCN Endangered�@ B1+2c�@ ver.2.3�B

�Ÿ
�@Systomus orphoides (Valenciennes 1842)
�@�@�@syn.Barbodes saranus orphoides (Valenciennes 1842)
�@�@�@�@�@Barbus orphoides Valenciennes 1842
�@�@�@�@�@Puntius orphoides (Valenciennes 1842)
�@�@�@�@�@Puntius saranus orphoides (Valenciennes 1842)

�Ÿ
�@Systomus pleurotaenius (Bleeker 1863)
�@�@�@syn.Barbus pleurotaenius (Bleeker 1863) Day 1878
�@�@�@�@�@Puntius pleurotaenius Bleeker 1863
�@^*1878�wThe Fishes of India, Being a Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters of India, Burma and Ceylon 1 Extra�FFrancis Day�x�B
�@"ƒuƒ‰ƒbƒNƒ‰ƒCƒ“ƒoƒ‹ƒu"�B‚Q‚Q�`‚Q‚U�Ž�E‚�‚g‚U�D‚T�`‚V�D‚T�BIUCN Lower Risk/conservation dependent ver.2.3�B

�Ÿ
�@Systomus rubripinnis (Valenciennes 1842)
�@�@�@syn.Barbodes rubripinnis (Valenciennes 1842)
�@�@�@�@�@Barbus caudimarginatus Blyth 1860
�@�@�@�@�@Barbus rubripinnis Valenciennes 1842
�@�@�@�@�@Barbus saranus caudimarginatus (Blyth 1860)
�@�@�@�@�@Barbus sarananellus Bleeker 1849
�@�@�@�@�@Barbus sewelli Prashad & Mukerji 1929
�@�@�@�@�@Puntius rubripinnis (Valenciennes 1842)
�@^*2021�wFishes of the Indo-Chinese Mekong�FNagao Natural Environment Foundation�x�B
�@"ƒWƒƒƒ�ƒoƒ‹ƒu"�B‚Q‚Q�`‚Q‚T�Ž�E‚�‚g‚U�D‚O�`‚U�D‚T�B

�Ÿ
�@Systomus rufus Plamoottil 2014


�Ÿ
�@Systomus saranus (Hamilton 1822)
�@�@�@syn.Barbodes saranus (Hamilton 1822)
�@�@�@�@�@Barbus binduchitrus Hora 1937
�@�@�@�@�@Barbus chrysopomus Valenciennes 1842
�@�@�@�@�@Barbus diliciosus McClelland 1839
�@�@�@�@�@Barbus duvaucelii Valenciennes 1842
�@�@�@�@�@Barbus gardonides Valenciennes 1842
�@�@�@�@�@Barbus gibbosus Valenciennes 1842 non Peters 1852
�@�@�@�@�@Barbus kakus Valenciennes 1842
�@�@�@�@�@Barbus polydori Valenciennes 1842
�@�@�@�@�@Barbus roseipinnis Valenciennes 1842
�@�@�@�@�@Barbus saranus (Hamilton 1822) Day 1876
�@�@�@�@�@Cyprinus saranus Hamilton 1822
�@�@�@�@�@Puntius chrysopomus (Valenciennes 1842)
�@�@�@�@�@Puntius roseipinnis (Valenciennes 1842)
�@�@�@�@�@Puntius saberi Datta & Karmakar 1981
�@�@�@�@�@Puntius saranus (Hamilton 1822)
�@�@�@�@�@Systomus chrysopomus (Valenciennes 1842)
�@^*1876�wOn Some of the Fishes of the Deccan�FFrancis Day�x�B
�@^*1878�wA Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters India, Burma and Ceylon 2, Atlas Constaining 198 Plates�FFrancis Day�x�B
�@^*1878�wThe Fishes of India, Being a Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters of India, Burma and Ceylon 1 Extra�FFrancis Day�x�B
�@"ƒ|ƒ“ƒfƒBƒ`ƒFƒŠ�[ƒoƒ‹ƒu"�B

�Ÿ
�@Systomus simus (Smith 1945)
�@�@�@syn.Barbus jacobus-boehlkei Fowler 1958
�@�@�@�@�@Barbus jacobusboehlkei (Fowler 1958)
�@�@�@�@�@Puntius simus Smith 1945
�@�@�@�@�@Puntius jacobusboehlkei (Fowler 1958)
�@�@�@�@�@Systomus jacobusboehlkei (Fowler 1958)�@�¦ƒtƒBƒbƒVƒ…ƒx�[ƒX‚Ì—LŒø–¼�Ì�B
�@^*1958�wSome New Taxonomic Names of Fishlike Vertebrates�FHenry Weed Fowler�x�B
�@^*2021�wFishes of the Indo-Chinese Mekong�FNagao Natural Environment Foundation�x�B
�@Barbus jacobus-boehlkei Fowler 1958 ‚ÍPuntius simus Smith 1945‚Ì•Ï–¼�AˆÈ‘O“¯‘®‚¾‚Á‚½Barbus simus Sauvage & Dabry de Thiersant 1874��Œ»Onychostoma simum‚Æ‚©‚Ԃ邽‚ß�B�®�AƒtƒBƒbƒVƒ…ƒx�[ƒX‚Å‚Í‚±‚ê‚ð“Æ—§‚³‚¹‚Ä‚¢‚é�B

�Ÿ
�@Systomus spilurus (Günther 1868)
�@�@�@syn.Barbus spilurus Günther 1868

�Ÿ
�@Systomus subnasutus (Valenciennes 1842)
�@�@�@syn.Barbodes saranus subnasutus (Valenciennes 1842)
�@�@�@�@�@Barbus subnasutus Valenciennes 1842
�@�@�@�@�@Puntius saranus subnasutus (Valenciennes 1842)
�@�@�@�@�@Puntius subnasutus (Valenciennes 1842)
�@�@�@�@�@Systomus saranus subnasutus (Valenciennes 1842)

�Ÿ
�@Systomus timbiri (Deraniyagala 1963)
�@�@�@syn.Puntius timbiri Deraniyagala 1963

---

Genus Waikhomia^♀ KATWATE, KUMKAR, RAGHAVAN & DAHANUKAR 2020�@ƒ�ƒCƒRƒ~ƒA‘®

�Ÿ
�@Waikhomia hira Katwate, Kumkar, Raghavan & Dahanukar 2020
�@^*2020�wTaxonomy and systematics of the �eMaharaja Barbs�f (Teleostei: Cyprinidae), with the description of a new genus and species from the Western Ghats, India�FUnmesh Katwate, Pradeep Kumkar, Rajeev Raghavan & Neelesh Dahanukar�x�B

�Ÿ
�@Waikhomia sahyadriensis (Silas 1953) Katwate, Kumkar, Raghavan & Dahanukar 2020
�@�@�@syn.Puntius sahyadriensis Silas 1953
�@^*2020�wTaxonomy and systematics of the �eMaharaja Barbs�f (Teleostei: Cyprinidae), with the description of a new genus and species from the Western Ghats, India�FUnmesh Katwate, Pradeep Kumkar, Rajeev Raghavan & Neelesh Dahanukar�x�B
�@"ƒJƒuƒŠƒoƒ‹ƒu"�B

---